10-Q

ALEXANDRIA REAL ESTATE EQUITIES, INC. (ARE)

10-Q 2025-04-28 For: 2025-03-31
View Original
Added on April 01, 2026

UNITED STATES

SECURITIES AND EXCHANGE COMMISSION

Washington, D.C. 20549

FORM 10-Q

(Mark One)

☒QUARTERLY REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES

EXCHANGE ACT OF 1934

For the quarterly period ended March 31, 2025

OR

☐TRANSITION REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES

EXCHANGE ACT OF 1934

For the transition period from ____________ to ____________

Commission file number 1-12993

ALEXANDRIA REAL ESTATE EQUITIES, INC.

(Exact name of registrant as specified in its charter)

Maryland 95-4502084
(State or other jurisdiction of<br><br>incorporation or organization) (I.R.S. Employer Identification Number)

26 North Euclid Avenue, Pasadena, California 91101

(Address of principal executive offices) (Zip code)

(626) 578-0777

(Registrant’s telephone number, including area code)

N/A

(Former name, former address and former fiscal year, if changed since last report)

Securities registered pursuant to Section 12(b) of the Exchange Act:

Title of each class Trading Symbol(s) Name of each exchange on which registered
Common Stock, $0.01 par value per share ARE New York Stock Exchange

Indicate by check mark whether the registrant (1) has filed all reports required to be filed by Section 13 or 15(d) of the Securities

Exchange Act of 1934 during the preceding 12 months (or for such shorter period that the registrant was required to file such reports),

and (2) has been subject to such filing requirements for the past 90 days. Yes ☒  No ☐

Indicate by check mark whether the registrant has submitted electronically every Interactive Data File required to be submitted and

posted pursuant to Rule 405 of Regulation S-T (§ 232.405 of this chapter) during the preceding 12 months (or for such shorter period

that the registrant was required to submit and post such files). Yes ☒  No  ☐

Indicate by check mark whether the registrant is a large accelerated filer, an accelerated filer, a non-accelerated filer, a smaller reporting

company, or an emerging growth company. See the definitions of “large accelerated filer,” “accelerated filer,” “smaller reporting

company,” and “emerging growth company” in Rule 12b-2 of the Exchange Act.

Large accelerated filer Smaller reporting company
Accelerated filer Emerging growth company
Non-accelerated filer

If an emerging growth company, indicate by check mark if the registrant has elected not to use the extended transition period for

complying with any new or revised financial accounting standards provided pursuant to Section 13(a) of the Exchange Act. ☐

Indicate by check mark whether the registrant is a shell company (as defined in Rule 12b-2 of the Exchange Act). Yes ☐  No ☒

As of April 15, 2025, 172,988,874 shares of common stock, par value $0.01 per share, were outstanding.

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TABLE OF CONTENTS

Page
PART I – FINANCIAL INFORMATION
Item 1. FINANCIAL STATEMENTS (UNAUDITED)
Consolidated Balance Sheets as of March 31, 2025 and December 31, 2024 .......................................................... 1
Consolidated Financial Statements for the Three Months Ended March 31, 2025 and 2024:
Consolidated Statements of Operations ................................................................................................................... 2
Consolidated Statements of Comprehensive Income ............................................................................................ 3
Consolidated Statements of Changes in Stockholders’ Equity and Noncontrolling Interests .......................... 4
Consolidated Statements of Cash Flows ................................................................................................................ 6
Notes to Consolidated Financial Statements .................................................................................................................... 8
Item 2. MANAGEMENT’S DISCUSSION AND ANALYSIS OF FINANCIAL CONDITION AND RESULTS OF<br><br>OPERATIONS ........................................................................................................................................................................ 44
Item 3. QUANTITATIVE AND QUALITATIVE DISCLOSURES ABOUT MARKET RISK ......................................................... 104
Item 4. CONTROLS AND PROCEDURES ..................................................................................................................................... 105
PART II – OTHER INFORMATION
Item 1. LEGAL PROCEEDINGS ...................................................................................................................................................... 106
Item 1A. RISK FACTORS .................................................................................................................................................................... 107
Item 2. UNREGISTERED SALES OF EQUITY SECURITIES AND USE OF PROCEEDS ................................................... 109
Item 5. OTHER INFORMATION ....................................................................................................................................................... 109
Item 6. EXHIBITS ............................................................................................................................................................................... 110
SIGNATURES ................................................................................................................................................................................................. 111

ii

GLOSSARY

The following abbreviations or acronyms that may be used in this document

shall have the adjacent meanings set forth below:

ASU Accounting Standards Update
ATM At the Market
CIP Construction in Progress
EPS Earnings per Share
FASB Financial Accounting Standards Board
FFO Funds From Operations
GAAP U.S. Generally Accepted Accounting Principles
IRS Internal Revenue Service
JV Joint Venture
Nareit National Association of Real Estate Investment Trusts
NAV Net Asset Value
NYSE New York Stock Exchange
REIT Real Estate Investment Trust
RSF Rentable Square Feet/Foot
SEC Securities and Exchange Commission
SF Square Feet/Foot
SoDo South of Downtown submarket of Seattle
SOFR Secured Overnight Financing Rate
SoMa South of Market submarket of the San Francisco Bay Area
U.S. United States
VIE Variable Interest Entity

1

PART I – FINANCIAL INFORMATION

ITEM 1. FINANCIAL STATEMENTS (UNAUDITED)

Alexandria Real Estate Equities, Inc.

Consolidated Balance Sheets

(In thousands)

March 31, 2025 December 31, 2024
(Unaudited)
Assets
Investments in real estate $32,121,712 $32,110,039
Investments in unconsolidated real estate joint ventures 50,086 39,873
Cash and cash equivalents 476,430 552,146
Restricted cash 7,324 7,701
Tenant receivables 6,875 6,409
Deferred rent 1,210,584 1,187,031
Deferred leasing costs 489,287 485,959
Investments 1,479,688 1,476,985
Other assets 1,758,442 1,661,306
Total assets $37,600,428 $37,527,449
Liabilities, Noncontrolling Interests, and Equity
Secured notes payable $150,807 $149,909
Unsecured senior notes payable 12,640,144 12,094,465
Unsecured senior line of credit and commercial paper 299,883
Accounts payable, accrued expenses, and other liabilities 2,281,414 2,654,351
Dividends payable 228,622 230,263
Total liabilities 15,600,870 15,128,988
Commitments and contingencies
Redeemable noncontrolling interests 9,612 19,972
Alexandria Real Estate Equities, Inc.’s stockholders’ equity:
Common stock 1,701 1,722
Additional paid-in capital 17,509,148 17,933,572
Accumulated other comprehensive loss (46,202) (46,252)
Alexandria Real Estate Equities, Inc.’s stockholders’ equity 17,464,647 17,889,042
Noncontrolling interests 4,525,299 4,489,447
Total equity 21,989,946 22,378,489
Total liabilities, noncontrolling interests, and equity $37,600,428 $37,527,449

The accompanying notes are an integral part of these consolidated financial statements.

2

Alexandria Real Estate Equities, Inc.

Consolidated Statements of Operations

(In thousands, except per share amounts)

(Unaudited)

Three Months Ended March 31,
2025 2024
Revenues:
Income from rentals $743,175 $755,551
Other income 14,983 13,557
Total revenues 758,158 769,108
Expenses:
Rental operations 226,395 218,314
General and administrative 30,675 47,055
Interest 50,876 40,840
Depreciation and amortization 342,062 287,554
Impairment of real estate 32,154
Total expenses 682,162 593,763
Equity in (losses) earnings of unconsolidated real estate joint ventures (507) 155
Investment (loss) income (49,992) 43,284
Gain on sales of real estate 13,165 392
Net income 38,662 219,176
Net income attributable to noncontrolling interests (47,601) (48,631)
Net (loss) income attributable to Alexandria Real Estate Equities, Inc.’s stockholders (8,939) 170,545
Net income attributable to unvested restricted stock awards (2,660) (3,659)
Net (loss) income attributable to Alexandria Real Estate Equities, Inc.’s common stockholders $(11,599) $166,886
Net (loss) income per share attributable to Alexandria Real Estate Equities, Inc.’s common<br><br>stockholders:
Basic $(0.07) $0.97
Diluted $(0.07) $0.97

The accompanying notes are an integral part of these consolidated financial statements.

3

Alexandria Real Estate Equities, Inc.

Consolidated Statements of Comprehensive Income

(In thousands)

(Unaudited)

Three Months Ended March 31,
2025 2024
Net income $38,662 $219,176
Other comprehensive income (loss)
Unrealized gains (losses) on foreign currency translation:
Unrealized foreign currency translation gains (losses) arising during the period 50 (7,919)
Unrealized gains (losses) on foreign currency translation, net 50 (7,919)
Total other comprehensive income (loss) 50 (7,919)
Comprehensive income 38,712 211,257
Less: comprehensive income attributable to noncontrolling interests (47,601) (48,631)
Comprehensive (loss) income attributable to Alexandria Real Estate Equities, Inc.’s stockholders $(8,889) $162,626

The accompanying notes are an integral part of these consolidated financial statements.

4

Alexandria Real Estate Equities, Inc.

Consolidated Statement of Changes in Stockholders’ Equity and Noncontrolling Interests

(Dollars in thousands)

(Unaudited)

Alexandria Real Estate Equities, Inc.’s Stockholders’ Equity
Number of<br><br>Common<br><br>Shares Common<br><br>Stock Additional<br><br>Paid-In<br><br>Capital Retained<br><br>Earnings Accumulated<br><br>Other<br><br>Comprehensive<br><br>Loss Noncontrolling<br><br>Interests Total<br><br>Equity Redeemable<br><br>Noncontrolling<br><br>Interests
Balance as of December 31, 2024 172,203,443 $1,722 $17,933,572 $— $(46,252) $4,489,447 $22,378,489 $19,972
Net (loss) income (8,939) 47,331 38,392 270
Total other comprehensive income 50 50
Contributions from and sales of noncontrolling interests 54 54,354 54,408
Distributions to and redemption of noncontrolling interests (7,048) (65,833) (72,881) (10,630)
Issuance pursuant to stock plan 125,280 1 32,755 32,756
Taxes related to net settlement of equity awards (46,547) (4,735) (4,735)
Repurchase of common stock (2,152,293) (22) (208,165) (208,187)
Dividends declared on common stock ($1.32 per share) (228,346) (228,346)
Reclassification of distributions and net loss (237,285) 237,285
Balance as of March 31, 2025 170,129,883 $1,701 $17,509,148 $— $(46,202) $4,525,299 $21,989,946 $9,612

The accompanying notes are an integral part of these consolidated financial statements.

5

Alexandria Real Estate Equities, Inc.

Consolidated Statement of Changes in Stockholders’ Equity and Noncontrolling Interests

(Dollars in thousands)

(Unaudited)

Alexandria Real Estate Equities, Inc.’s Stockholders’ Equity
Number of<br><br>Common<br><br>Shares Common<br><br>Stock Additional<br><br>Paid-In<br><br>Capital Retained<br><br>Earnings Accumulated<br><br>Other<br><br>Comprehensive<br><br>Loss Noncontrolling<br><br>Interests Total<br><br>Equity Redeemable<br><br>Noncontrolling<br><br>Interests
Balance as of December 31, 2023 171,910,599 $1,719 $18,485,352 $— $(15,896) $4,135,338 $22,606,513 $16,480
Net income 170,545 48,359 218,904 272
Total other comprehensive loss (7,919) (7,919)
Contributions from and sales of noncontrolling interests 7,201 180,978 188,179
Distributions to and redemption of noncontrolling interests (8,070) (67,907) (75,977) (382)
Transfer of noncontrolling interests (250) (250) 250
Reallocation of capital to joint venture partner (30,185) 30,185
Issuance pursuant to stock plan 164,784 2 39,376 39,378
Taxes related to net settlement of equity awards (67,416) (1) (7,395) (7,396)
Dividends declared on common stock ($1.27 per share) (222,134) (222,134)
Reclassification of distributions in excess of earnings (51,589) 51,589
Balance as of March 31, 2024 172,007,967 $1,720 $18,434,690 $— $(23,815) $4,326,703 $22,739,298 $16,620

The accompanying notes are an integral part of these consolidated financial statements.

6

Alexandria Real Estate Equities, Inc.<br><br>Consolidated Statements of Cash Flows<br><br>(In thousands)<br><br>(Unaudited)
Three Months Ended March 31,
2025 2024
Operating Activities:
Net income $38,662 $219,176
Adjustments to reconcile net income to net cash provided by operating activities:
Depreciation and amortization 342,062 287,554
Impairment of real estate 32,154
Gain on sales of real estate (13,165) (392)
Equity in losses (earnings) of unconsolidated real estate joint ventures 507 (155)
Distributions of earnings from unconsolidated real estate joint ventures 172 884
Amortization of loan fees 4,691 4,142
Amortization of debt discounts 349 318
Amortization of acquired above- and below-market leases (15,222) (30,340)
Deferred rent (22,023) (48,251)
Stock compensation expense 10,064 17,125
Investment loss (income) 49,992 (43,284)
Changes in operating assets and liabilities:
Tenant receivables (467) 731
Deferred leasing costs (26,645) (23,970)
Other assets (37,034) (40,728)
Accounts payable, accrued expenses, and other liabilities (156,148) (1,653)
Net cash provided by operating activities 207,949 341,157
Investing Activities:
Proceeds from sales of real estate 68,182 16,670
Additions to real estate (645,841) (693,268)
Purchases of real estate (194,002)
Change in escrow deposits (9,506) (1,008)
Investments in unconsolidated real estate joint ventures (10,994) (3,224)
Additions to non-real estate investments (69,311) (60,572)
Sales of and distributions from non-real estate investments 12,691 40,550
Net cash used in investing activities $(654,779) $(894,854)

7

Alexandria Real Estate Equities, Inc.<br><br>Consolidated Statements of Cash Flows<br><br>(In thousands)<br><br>(Unaudited)
Three Months Ended March 31,
2025 2024
Financing Activities:
Borrowings under secured notes payable $824 $10,216
Proceeds from issuance of unsecured senior notes payable 548,532 998,806
Proceeds from issuances under commercial paper program 2,700,000 3,170,000
Repayments of borrowings under commercial paper program (2,400,000) (3,270,000)
Payments of loan fees (5,406) (10,118)
Taxes paid related to net settlement of equity awards (5,558) (23,964)
Repurchase of common stock (208,187)
Dividends on common stock (229,987) (221,824)
Contributions from and sales of noncontrolling interests 54,409 82,853
Distributions to and purchases of noncontrolling interests (83,852) (111,540)
Net cash provided by financing activities 370,775 624,429
Effect of foreign exchange rate changes on cash and cash equivalents (38) 192
Net (decrease) increase in cash, cash equivalents, and restricted cash (76,093) 70,924
Cash, cash equivalents, and restricted cash as of the beginning of period 559,847 660,771
Cash, cash equivalents, and restricted cash as of the end of period $483,754 $731,695
Supplemental Disclosure and Non-Cash Investing and Financing Activities:
Cash paid during the period for interest, net of interest capitalized $33,776 $34,901
Accrued construction for current-period additions to real estate $147,045 $318,041
Transfer of real estate assets and/or equipment from tenants $39,950 $11,965
Notes receivable issued in connection with sales of real estate $91,000 $—
Derecognition of net investment in real estate from sales-type lease $4,677 $—
Contribution of assets from and issuance of noncontrolling interest to real estate joint<br><br>venture partner $— $103,547
Reallocation of additional paid-in-capital to consolidated joint venture partner’s non-<br><br>controlling interest $— $30,185

The accompanying notes are an integral part of these consolidated financial statements.

8

Alexandria Real Estate Equities, Inc.

Notes to Consolidated Financial Statements

(Unaudited)

1.ORGANIZATION AND BASIS OF PRESENTATION

Alexandria Real Estate Equities, Inc. (NYSE: ARE), an S&P 500® life science REIT, is the pioneer of the life science real estate

niche since its founding in 1994. Alexandria is the preeminent and longest-tenured owner, operator, and developer of collaborative

Megacampus™ ecosystems in AAA life science innovation cluster locations, including Greater Boston, the San Francisco Bay Area,

San Diego, Seattle, Maryland, Research Triangle, and New York City. As of March 31, 2025, Alexandria has a total market capitalization

of $28.8 billion and an asset base in North America that includes 39.6 million RSF of operating properties and 4.0 million RSF of Class

A/A+ properties undergoing construction. As used in this quarterly report on Form 10-Q, references to the “Company,” “Alexandria,”

“ARE,” “we,” “us,” and “our” refer to Alexandria Real Estate Equities, Inc. and its consolidated subsidiaries. The accompanying

unaudited consolidated financial statements include the accounts of Alexandria Real Estate Equities, Inc. and its consolidated

subsidiaries. All significant intercompany balances and transactions have been eliminated.

We have prepared the accompanying interim consolidated financial statements in accordance with GAAP and in conformity

with the rules and regulations of the SEC. In our opinion, these interim consolidated financial statements presented herein reflect all

adjustments, of a normal recurring nature, that are necessary to fairly present the interim consolidated financial statements. The results

of operations for the interim period are not necessarily indicative of the results that may be expected for the year ending December 31,

  1. These unaudited consolidated financial statements should be read in conjunction with the audited consolidated financial

statements and the notes thereto included in our annual report on Form 10-K for the year ended December 31, 2024. Any references to

our total market capitalization, number or quality of buildings or tenants, quality of location, square footage, number of leases, or

occupancy percentage, and any amounts derived from these values in these notes to consolidated financial statements are outside the

scope of our independent registered public accounting firm’s procedures.

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES

Consolidation

On an ongoing basis, as circumstances indicate the need for reconsideration, we evaluate each legal entity that is not wholly

owned by us in accordance with the consolidation accounting guidance. Our evaluation considers all of our variable interests, including

equity ownership, as well as fees paid to us for our involvement in the management of each partially owned entity. To fall within the

scope of the consolidation guidance, an entity must meet both of the following criteria:

•The entity has a legal structure that has been established to conduct business activities and to hold assets; such entity

can be in the form of a partnership, limited liability company, or corporation, among others; and

•We have a variable interest in the legal entity — i.e., variable interests that are contractual, such as equity ownership, or

other financial interests that change with changes in the fair value of the entity’s net assets.

If an entity does not meet both criteria above, we apply other accounting literature, such as the equity method of accounting. If

an entity does meet both criteria above, we evaluate such entity for consolidation under either the variable interest model if the legal

entity meets any of the characteristics below to qualify as a VIE, or under the voting model for all other legal entities that are not VIEs.

A legal entity is determined to be a VIE if it has any of the following three characteristics:

1)The entity does not have sufficient equity to finance its activities without additional subordinated financial support;

2)The entity is established with non-substantive voting rights (i.e., the entity deprives the majority economic interest

holder(s) of voting rights); or

3)The equity holders, as a group, lack the characteristics of a controlling financial interest. Equity holders meet this criterion

if they lack any of the following:

•The power, through voting rights or similar rights, to direct the activities of the entity that most significantly influence

the entity’s economic performance, as evidenced by:

•Substantive participating rights in day-to-day management of the entity’s activities; or

•Substantive kick-out rights over the party responsible for significant decisions;

•The obligation to absorb the entity’s expected losses; or

•The right to receive the entity’s expected residual returns.

9

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

For an entity, including our real estate joint ventures, structured as a limited partnership or a limited liability company, our

evaluation of whether the equity holders (equity partners other than the general partner or the managing member of a joint venture) lack

the characteristics of a controlling financial interest includes the evaluation of whether the limited partners or non-managing members

(the noncontrolling equity holders) lack both substantive participating rights and substantive kick-out rights, defined as follows:

•Participating rights provide the noncontrolling equity holders the ability to direct significant financial and operating

decisions made in the ordinary course of business that most significantly influence the entity’s economic performance.

•Kick-out rights allow the noncontrolling equity holders to remove the general partner or managing member without cause.

If we conclude that any of the three characteristics of a VIE are met, including that the equity holders lack the characteristics of

a controlling financial interest because they lack both substantive participating rights and substantive kick-out rights, we conclude that

the entity is a VIE and evaluate it for consolidation under the variable interest model.

Variable interest model

If an entity is determined to be a VIE, we evaluate whether we are the primary beneficiary. The primary beneficiary analysis is

a qualitative analysis based on power and benefits. We consolidate a VIE if we have both power and benefits — that is, (i) we have the

power to direct the activities of a VIE that most significantly influence the VIE’s economic performance (power) and (ii) we have the

obligation to absorb losses of or the right to receive benefits from the VIE that could potentially be significant to the VIE (benefits). We

consolidate VIEs whenever we determine that we are the primary beneficiary. Refer to Note 4 – “Consolidated and unconsolidated real

estate joint ventures” and Note 7 – “Investments” to our unaudited consolidated financial statements for information on specific entities

that qualify as VIEs. If we have a variable interest in a VIE but are not the primary beneficiary, we account for our investment using the

equity method.

Voting model

If a legal entity fails to meet any of the three characteristics of a VIE (i.e., insufficiency of equity, existence of non-substantive

voting rights, or lack of a controlling financial interest), we then evaluate such entity under the voting model. Under the voting model, we

consolidate the entity if we determine that we, directly or indirectly, have greater than 50% of the voting shares (or own a majority of the

limited partnership’s kick-out rights through voting interests), and that other equity holders do not have substantive participating rights.

Refer to Note 4 – “Consolidated and unconsolidated real estate joint ventures” to our unaudited consolidated financial statements for

information on specific joint ventures that qualify for evaluation under the voting model.

Use of estimates

The preparation of consolidated financial statements in conformity with GAAP requires us to make estimates and assumptions

that affect the reported amounts of assets, liabilities, and equity; the disclosure of contingent assets and liabilities as of the date of the

consolidated financial statements; and the amounts of revenues and expenses during the reporting period. Actual results could

materially differ from those estimates.

Investments in real estate

Evaluation of business combination or asset acquisition

We evaluate each acquisition of real estate or in-substance real estate (including equity interests in entities that predominantly

hold real estate assets) to determine whether the integrated set of assets and activities acquired meets the definition of a business and

needs to be accounted for as a business combination. An acquisition of an integrated set of assets and activities that does not meet the

definition of a business is accounted for as an asset acquisition. If either of the following criteria is met, the integrated set of assets and

activities acquired would not qualify as a business:

•Substantially all of the fair value of the gross assets acquired is concentrated in either a single identifiable asset or a group

of similar identifiable assets; or

•The integrated set of assets and activities is lacking, at a minimum, an input and a substantive process that together

significantly contribute to the ability to create outputs (i.e., revenue generated before and after the transaction).

An acquired process is considered substantive if:

•The process includes an organized workforce (or includes an acquired contract that provides access to an organized

workforce) that is skilled, knowledgeable, and experienced in performing the process;

•The process cannot be replaced without significant cost, effort, or delay; or

•The process is considered unique or scarce.

10

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

Generally, our acquisitions of real estate or in-substance real estate do not meet the definition of a business because

substantially all of the fair value is concentrated in a single identifiable asset or group of similar identifiable assets (i.e., land, buildings,

and related intangible assets) or because the acquisition does not include a substantive process in the form of an acquired workforce or

an acquired contract that cannot be replaced without significant cost, effort, or delay. When evaluating acquired service or management

contracts, we consider the nature of the services performed, the terms of the contract relative to similar arm’s-length contracts, and the

availability of comparable vendors in evaluating whether the acquired contract constitutes a substantive process.

Recognition of real estate acquired

We evaluate each acquisition of real estate or in-substance real estate (including equity interests in entities that predominantly

hold real estate assets) to determine whether the integrated set of assets and activities acquired meets the definition of a business and

needs to be accounted for as a business combination. An acquisition of an integrated set of assets and activities that does not meet the

definition of a business is accounted for as an asset acquisition.

For acquisitions of real estate or in-substance real estate that are accounted for as business combinations, we allocate the

acquisition consideration (excluding acquisition costs) to the assets acquired, liabilities assumed, noncontrolling interests, and

previously existing ownership interests at fair value as of the acquisition date. Assets include intangible assets such as tenant

relationships, acquired in-place leases, and favorable intangibles associated with in-place leases in which we are the lessor. Liabilities

include unfavorable intangibles associated with in-place leases in which we are the lessor. In addition, for acquired in-place finance or

operating leases in which we are the lessee, acquisition consideration is allocated to lease liabilities and related right-of-use assets,

adjusted to reflect favorable or unfavorable terms of the lease when compared with market terms. Any excess (deficit) of the

consideration transferred relative to the fair value of the net assets acquired is accounted for as goodwill (bargain purchase gain).

Acquisition costs related to business combinations are expensed as incurred.

Generally, we expect that acquisitions of real estate or in-substance real estate will not meet the definition of a business

because substantially all of the fair value is concentrated in a single identifiable asset or group of similar identifiable assets (i.e., land,

buildings, and related intangible assets). The accounting model for asset acquisitions is similar to the accounting model for business

combinations, except that the acquisition consideration (including acquisition costs) is allocated to the individual assets acquired and

liabilities assumed on a relative fair value basis. Any excess (deficit) of the consideration transferred relative to the sum of the fair value

of the assets acquired and liabilities assumed is allocated to the individual assets and liabilities based on their relative fair values. As a

result, asset acquisitions do not result in the recognition of goodwill or a bargain purchase gain. Incremental and external direct

acquisition costs related to acquisitions of real estate or in-substance real estate (such as legal and other third-party services) are

capitalized.

We exercise judgment to determine the key assumptions used to allocate the purchase price of real estate acquired among its

components. The allocation of the consideration to the various components of properties acquired during the year can have an effect on

our net income due to the useful depreciable and amortizable lives applicable to each component and the recognition of the related

depreciation and amortization expense in our consolidated statements of operations. We apply judgment in utilizing available

comparable market information to assess relative fair value. We assess the relative fair values of tangible and intangible assets and

liabilities based on available comparable market information, including estimated replacement costs, rental rates, and recent market

transactions. In addition, we may use estimated cash flow projections that utilize appropriate discount and capitalization rates.

Estimates of future cash flows are based on a number of factors, including the historical operating results, known and anticipated

trends, and market/economic conditions that may affect the property.

The value of tangible assets acquired is based upon our estimation of fair value on an “as if vacant” basis. The value of

acquired in-place leases includes the estimated costs during the hypothetical lease-up period and other costs that would have been

incurred in the execution of similar leases under the market conditions at the acquisition date of the acquired in-place lease. If there is a

bargain fixed-rate renewal option for the period beyond the noncancelable lease term of an in-place lease, we evaluate intangible

factors, such as the business conditions in the industry in which the lessee operates, the economic conditions in the area in which the

property is located, and the ability of the lessee to sublease the property during the renewal term, in order to determine the likelihood

that the lessee will renew. When we determine that there is reasonable assurance that such bargain purchase option will be exercised,

we consider the option in determining the intangible value of such lease and its related amortization period. We also recognize the

relative fair values of assets acquired, the liabilities assumed, and any noncontrolling interest in acquisitions of less than a 100%

interest when the acquisition constitutes a change in control of the acquired entity.

11

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

Depreciation and amortization

The values allocated to buildings and building improvements, land improvements, tenant improvements, and equipment are

depreciated on a straight-line basis. For buildings and building improvements, we depreciate using the shorter of the respective ground

lease terms or their estimated useful lives, not to exceed 40 years. Land improvements are depreciated over their estimated useful

lives, not to exceed 20 years. Tenant improvements are depreciated over their respective lease terms or estimated useful lives, and

equipment is depreciated over the shorter of the lease term or its estimated useful life. The values of the right-of-use assets are

amortized on a straight-line basis over the remaining terms of each related lease. The values of acquired in-place leases and

associated favorable intangibles (i.e., acquired above-market leases) are classified in other assets in our consolidated balance sheets

and are amortized over the remaining terms of the related leases as a reduction of income from rentals in our consolidated statements

of operations. The values of unfavorable intangibles (i.e., acquired below-market leases) associated with acquired in-place leases are

classified in accounts payable, accrued expenses, and other liabilities in our consolidated balance sheets and are amortized over the

remaining terms of the related leases as an increase in income from rentals in our consolidated statements of operations.

Capitalized project costs

We capitalize project costs, including pre-construction costs, interest, property taxes, insurance, and other costs directly

related and essential to the development, redevelopment, pre-construction, or construction of a project. Capitalization of development,

redevelopment, pre-construction, and construction costs is required while activities are ongoing to prepare an asset for its intended use.

Fluctuations in our development, redevelopment, pre-construction, and construction activities could result in significant changes to total

expenses and net income. Costs incurred after a project is substantially complete and ready for its intended use are expensed as

incurred. Should development, redevelopment, pre-construction, or construction activity cease, interest, property taxes, insurance, and

certain other costs would no longer be eligible for capitalization and would be expensed as incurred. Expenditures for repairs and

maintenance are expensed as incurred.

Real estate sales

A property is classified as held for sale when all of the following criteria for a plan of sale have been met: (i) management,

having the authority to approve the action, commits to a plan to sell the property; (ii) the property is available for immediate sale in its

present condition, subject only to terms that are usual and customary; (iii) an active program to locate a buyer and other actions

required to complete the plan to sell have been initiated; (iv) the sale of the property is probable and is expected to be completed within

one year; (v) the property is being actively marketed for sale at a price that is reasonable in relation to its current fair value; and

(vi) actions necessary to complete the plan of sale indicate that it is unlikely that significant changes to the plan will be made or that the

plan will be withdrawn. Depreciation of assets ceases upon designation of a property as held for sale.

If the disposal of a property represents a strategic shift that has (or will have) a major effect on our operations or financial

results, such as (i) a major line of business, (ii) a major geographic area, (iii) a major equity method investment, or (iv) other major parts

of an entity, then the operations of the property, including any interest expense directly attributable to it, are classified as discontinued

operations in our consolidated statements of operations, and amounts for all prior periods presented are reclassified from continuing

operations to discontinued operations. The disposal of an individual property generally will not represent a strategic shift and therefore

will typically not meet the criteria for classification as a discontinued operation.

We recognize gains or losses on real estate sales in accordance with the accounting standard on the derecognition of

nonfinancial assets arising from contracts with noncustomers. Our ordinary output activities consist of the leasing of space to our

tenants in our operating properties, not the sales of real estate. Therefore, sales of real estate (in which we are the seller) qualify as

contracts with noncustomers. In our transactions with noncustomers, we apply certain recognition and measurement principles

consistent with our method of recognizing revenue arising from contracts with customers. Derecognition of the asset is based on the

transfer of control. If a real estate sales contract includes our ongoing involvement with the property, then we evaluate each promised

good or service under the contract to determine whether it represents a separate performance obligation, constitutes a guarantee, or

prevents the transfer of control. If a good or service is considered a separate performance obligation, an allocated portion of the

transaction price is recognized as revenue as we transfer the related good or service to the buyer.

The recognition of gain or loss on the sale of a partial interest also depends on whether we retain a controlling or

noncontrolling interest in the property. If we retain a controlling interest in the property upon completion of the sale, we continue to

reflect the asset at its book value, record a noncontrolling interest for the book value of the partial interest sold, and recognize additional

paid-in capital for the difference between the consideration received and the partial interest at book value. Conversely, if we retain a

noncontrolling interest upon completion of the sale of a partial interest of real estate, we recognize a gain or loss as if 100% of the asset

were sold.

12

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

Impairment of long-lived assets

Prior to and subsequent to the end of each quarter, we review current activities and changes in the business conditions of all of

our long-lived assets to determine the existence of any triggering events or impairment indicators requiring an impairment analysis. If

triggering events or impairment indicators are identified, we review an estimate of the future undiscounted cash flows, including, if

necessary, a probability-weighted approach if multiple outcomes are under consideration.

Long-lived assets to be held and used, including our rental properties, CIP, land held for development, right-of-use assets

related to operating leases in which we are the lessee, and intangibles, are individually evaluated for impairment when conditions exist

that may indicate that the carrying amount of a long-lived asset may not be recoverable. The carrying amount of a long-lived asset to be

held and used is not recoverable if it exceeds the sum of the undiscounted cash flows expected to result from the use and eventual

disposition of the asset. Triggering events or impairment indicators for long-lived assets to be held and used are assessed by project

and include significant fluctuations in estimated net operating income, occupancy changes, significant near-term lease expirations,

current and historical operating and/or cash flow losses, construction costs, estimated completion dates, rental rates, and other market

factors. We assess the expected undiscounted cash flows based upon numerous factors, including, but not limited to, projected rental

rates, estimated exit capitalization rates, and anticipated construction costs for projects under construction, which are based on

available market information, current and historical operating results, known trends, current market/economic conditions that may affect

the asset, and our assumptions about the use of the asset, including, if necessary, a probability-weighted approach if multiple outcomes

are under consideration.

Upon determination that an impairment has occurred, a write-down is recognized to reduce the carrying amount of the asset to

its estimated fair value. If an impairment charge is not required to be recognized, the recognition of depreciation or amortization is

adjusted prospectively, as necessary, to reduce the carrying amount of the asset to its estimated disposition value over the remaining

period that the asset is expected to be held and used. We may adjust depreciation of properties that are expected to be disposed of or

redeveloped prior to the end of their useful lives.

We use the held for sale impairment model for our properties classified as held for sale, which is different from the held and

used impairment model. Under the held for sale impairment model, an impairment charge is recognized if the carrying amount of the

long-lived asset classified as held for sale exceeds its fair value less cost to sell. Because of these two different models, it is possible for

a long-lived asset previously classified as held and used to require the recognition of an impairment charge upon classification as held

for sale.

International operations

In addition to operating properties in the U.S., we have 11 properties in Canada. The functional currency for our subsidiaries

operating in the U.S. is the U.S. dollar. The local currency of a foreign subsidiary serves as its functional currency. The assets and

liabilities of our foreign subsidiaries are translated into U.S. dollars at the exchange rate in effect as of the financial statement date.

Revenue and expense accounts of our foreign subsidiaries are translated using the weighted-average exchange rate for the periods

presented. Gains or losses resulting from the translation are classified in accumulated other comprehensive income (loss) as a

separate component of total equity and are excluded from net income (loss).

Whenever a foreign investment meets the criteria for classification as held for sale, we evaluate the recoverability of the

investment under the held for sale impairment model. We may recognize an impairment charge if the carrying amount of the investment

exceeds its fair value less cost to sell. In determining an investment’s carrying amount, we consider its net book value and any

cumulative unrealized foreign currency translation adjustment related to the investment.

The appropriate amounts of foreign exchange rate gains or losses classified in accumulated other comprehensive income

(loss) are reclassified to net income (loss) when realized upon the sale of our investment or upon the complete or substantially

complete liquidation of our investment.

Investments

We hold investments in publicly traded companies and privately held entities primarily involved in the life science industry. As a

REIT, we generally limit our ownership of each individual entity’s voting stock to less than 10%. We evaluate each investment to

determine whether we have the ability to exercise significant influence, but not control, over an investee. We evaluate investments in

which our ownership is equal to or greater than 20%, but less than or equal to 50%, of an investee’s voting stock with a presumption

that we have this ability. For our investments in limited partnerships that maintain specific ownership accounts, we presume that such

ability exists when our ownership interest exceeds 3% to 5%. In addition to our ownership interest, we consider whether we have a

board seat or whether we participate in the investee’s policymaking process, among other criteria, to determine if we have the ability to

exert significant influence, but not control, over an investee. If we determine that we have such ability, we account for the investment

under the equity method, as described below. From time to time, we may hold equity investments in publicly traded companies that are

subject to contractual sale restrictions. We do not recognize a discount related to a contractual sale restriction.

13

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

Investments accounted for under the equity method

Under the equity method of accounting, we initially recognize our investment at cost and subsequently adjust the carrying

amount of the investment for our share of earnings or losses reported by the investee, distributions received, and other-than-temporary

impairments. For additional information about our investments accounted for under the equity method, refer to Note 7 – “Investments” to

our unaudited consolidated financial statements.

Investments that do not qualify for the equity method of accounting

For investees over which we determine that we do not have the ability to exercise significant influence or control, we account

for each investment depending on whether it is an investment in a (i) publicly traded company, (ii) privately held entity that reports NAV

per share, or (iii) privately held entity that does not report NAV per share, as described below.

Investments in publicly traded companies

Our investments in publicly traded companies are classified as investments with readily determinable fair values and are

presented at fair value in our consolidated balance sheets, with changes in fair value classified in investment income (loss) in our

consolidated statements of operations. The fair values for our investments in publicly traded companies are determined based on sales

prices or quotes available on securities exchanges.

Investments in privately held companies

Our investments in privately held entities without readily determinable fair values consist of (i) investments in privately held

entities that report NAV per share and (ii) investments in privately held entities that do not report NAV per share. These investments are

accounted for as follows:

Investments in privately held entities that report NAV per share

Investments in privately held entities that report NAV per share, such as our privately held investments in limited partnerships,

are presented at fair value using NAV as a practical expedient, with changes in fair value classified in investment income (loss) in our

consolidated statements of operations. We use NAV per share reported by limited partnerships generally without adjustment, unless we

are aware of information indicating that the NAV reported by a limited partnership does not accurately reflect the fair value of the

investment at our reporting date.

Investments in privately held entities that do not report NAV per share

Investments in privately held entities that do not report NAV per share are accounted for using a measurement alternative

under which these investments are measured at cost, adjusted for observable price changes and impairments, with changes classified

in investment income (loss) in our consolidated statements of operations.

An observable price arises from an orderly transaction for an identical or similar investment of the same issuer, which is

observed by an investor without expending undue cost and effort. Observable price changes result from, among other things, equity

transactions of the same issuer executed during the reporting period, including subsequent equity offerings or other reported equity

transactions related to the same issuer. To determine whether these transactions are indicative of an observable price change, we

evaluate, among other factors, whether these transactions have similar rights and obligations, including voting rights, distribution

preferences, and conversion rights to the investments we hold.

Impairment evaluation of equity method investments and investments in privately held entities that do not report NAV per share

We monitor equity method investments and investments in privately held entities that do not report NAV per share for new

developments, including operating results, prospects and results of clinical trials, new product initiatives, new collaborative agreements,

capital-raising events, and merger and acquisition activities. These investments are evaluated on the basis of a qualitative assessment

for indicators of impairment by monitoring the presence of the following triggering events or impairment indicators:

(i)a significant deterioration in the earnings performance, credit rating, asset quality, or business prospects of the investee;

(ii)a significant adverse change in the regulatory, economic, or technological environment of the investee;

(iii)a significant adverse change in the general market condition, including the research and development of technology and

products that the investee is bringing or attempting to bring to the market;

(iv)significant concerns about the investee’s ability to continue as a going concern; and/or

(v)a decision by investors to cease providing support or reduce their financial commitment to the investee.

If such indicators are present, we are required to estimate the investment’s fair value and immediately recognize an

impairment charge in an amount equal to the investment’s carrying value in excess of its estimated fair value.

14

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

Investment income/loss recognition and classification

We recognize both realized and unrealized gains and losses in our consolidated statements of operations, classified in

investment income (loss) in our consolidated statements of operations. Unrealized gains and losses represent:

(i)changes in fair value for investments in publicly traded companies;

(ii)changes in NAV for investments in privately held entities that report NAV per share;

(iii)observable price changes for investments in privately held entities that do not report NAV per share; and

(iv)our share of unrealized gains or losses reported by our equity method investees.

Realized gains and losses on our investments represent the difference between proceeds received upon disposition of

investments and their historical or adjusted cost basis. For our equity method investments, realized gains and losses represent our

share of realized gains or losses reported by the investee. Impairments are realized losses, which result in an adjusted cost basis, and

represent charges to reduce the carrying values of investments in privately held entities that do not report NAV per share and equity

method investments, if impairments are deemed other than temporary, to their estimated fair value.

Revenues

The table below provides details of our consolidated total revenues for the three months ended March 31, 2025 and 2024

(in thousands):

Three Months Ended March 31,
2025 2024
Income from rentals:
Revenues subject to the lease accounting standard:
Operating leases $731,421 $746,061
Direct financing and sales-type leases 810 659
Revenues subject to the lease accounting standard 732,231 746,720
Revenues subject to the revenue recognition accounting standard 10,944 8,831
Income from rentals 743,175 755,551
Other income 14,983 13,557
Total revenues $758,158 $769,108

During the three months ended March 31, 2025 and 2024, revenues that were subject to the lease accounting standard

aggregated $732.2 million, or 96.6%, and $746.7 million, or 97.1%, of our total revenue, respectively. Our other income consisted

primarily of management fees and interest income earned during each period presented. For a detailed discussion related to our

revenue streams, refer to “Lease accounting” and “Recognition of revenue arising from contracts with customers” in Note 2 – “Summary

of significant accounting policies” to our unaudited consolidated financial statements.

Lease accounting

Definition and classification of a lease

When we enter into a contract or amend an existing contract, we evaluate whether the contract meets the definition of a lease.

To meet the definition of a lease, the contract must meet all three criteria:

(i)One party (lessor) must hold an identified asset;

(ii)The counterparty (lessee) must have the right to obtain substantially all of the economic benefits from the use of the asset

throughout the period of the contract; and

(iii)The counterparty (lessee) must have the right to direct the use of the identified asset throughout the period of the contract.

15

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

We classify our leases as either finance leases or operating leases if we are the lessee, or sales-type, direct financing, or

operating leases if we are the lessor. We use the following criteria to determine if a lease is a finance lease (as a lessee) or sales-type

or direct financing lease (as a lessor):

(i)Ownership is transferred from lessor to lessee by the end of the lease term;

(ii)An option to purchase is reasonably certain to be exercised;

(iii)The lease term is for the major part of the underlying asset’s remaining economic life;

(iv)The present value of lease payments equals or exceeds substantially all of the fair value of the underlying asset; or

(v)The underlying asset is specialized and is expected to have no alternative use at the end of the lease term.

If we meet any of the above criteria, we account for the lease as a finance, a sales-type, or a direct financing lease. If we do

not meet any of the criteria, we account for the lease as an operating lease.

A lease is accounted for as a sales-type lease if it is considered to transfer control of the underlying asset to the lessee. A

lease is accounted for as a direct financing lease if risks and rewards are conveyed without the transfer of control, which is normally

indicated by the existence of a residual value guarantee from an unrelated third party other than the lessee.

This classification will determine the method of recognition of the lease:

•For an operating lease, we recognize income from rentals if we are the lessor, or rental operations expense if we are the

lessee, over the term of the lease on a straight-line basis.

•For a sales-type lease or a direct financing lease, we recognize the income from rentals, or for a finance lease, we

recognize rental operations expense, over the term of the lease using the effective interest method.

•At inception of a sales-type lease or a direct financing lease, if we determine the fair value of the leased property is lower

than its carrying amount, we recognize a selling loss immediately at lease commencement. If fair value exceeds the

carrying amount of a lease, a gain is recognized at lease commencement on a sales-type lease. For a direct financing

lease, a gain is deferred at lease commencement and amortized over the lease term.

Lessor accounting

Costs to execute leases

We capitalize initial direct costs, which represent only incremental costs to execute a lease that would not have been incurred

if the lease had not been obtained. Costs that we incur to negotiate or arrange a lease, regardless of its outcome, such as for fixed

employee compensation, tax or legal advice to negotiate lease terms, and other costs, are expensed as incurred.

Operating leases

We account for the revenue from our lease contracts by utilizing the single component accounting policy. This policy requires

us to account for, by class of underlying asset, the lease component and nonlease component(s) associated with each lease as a single

component if two criteria are met:

(i)The timing and pattern of transfer of the lease component and the nonlease component(s) are the same; and

(ii)The lease component would be classified as an operating lease if it were accounted for separately.

Lease components consist primarily of fixed rental payments, which represent scheduled rental amounts due under our

leases, and contingent rental payments. Nonlease components consist primarily of tenant recoveries representing reimbursements of

rental operating expenses under our triple net lease structure, including recoveries for property taxes, insurance, utilities, repairs and

maintenance, and common area expenses.

If the lease component is the predominant component, we account for all revenues under such lease as a single component in

accordance with the lease accounting standard. Conversely, if the nonlease component is the predominant component, all revenues

under such lease are accounted for in accordance with the revenue recognition accounting standard. Our operating leases qualify for

the single component accounting, and the lease component in each of our leases is predominant. Therefore, we account for all

revenues from our operating leases under the lease accounting standard and classify these revenues as income from rentals in our

consolidated statements of operations.

We commence recognition of income from rentals related to the operating leases at the date the property is ready for its

intended use by the tenant and the tenant takes possession or controls the physical use of the leased asset. When a lease includes

construction of improvements, we determine whether the improvements are landlord or tenant assets. In determining if the

improvements are landlord or tenant improvements, we consider various factors, including, but not limited to, the following:

16

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

•Which party retains legal title to the improvements upon lease expiration;

•Whether the improvements are expected to have significant residual value at the end of the lease term;

•Whether the improvements are unique to the tenant;

•What happens to the improvements upon lease expiration (i.e., whether they are removed or preserved for the landlord);

•Which party bears all costs of the improvements (including the risk of cost overruns); and

•Which party supervises the construction of the improvements.

If the improvements are landlord assets, we capitalize such improvements. If the improvements are tenant assets, we do not

capitalize these assets. Improvements that qualify as tenant assets, if funded by us, are accounted for as lease incentives and

amortized as a reduction of revenue over the term of the lease. If the tenant funds improvements without reimbursement from us, and

we determine these improvements to be landlord assets, we consider the amount associated with the improvements to be non-cash

lease payments, which are recognized as incremental revenue over the term of the lease.

Income from rentals related to fixed rental payments under operating leases is recognized on a straight-line basis over the

respective operating lease terms. We classify amounts expected to be received in later periods as deferred rent in our consolidated

balance sheets. Amounts received currently but recognized as revenue in future periods are classified in accounts payable, accrued

expenses, and other liabilities in our consolidated balance sheets.

Income from rentals related to variable payments includes tenant recoveries and contingent rental payments. Tenant

recoveries, including reimbursements of utilities, repairs and maintenance, common area expenses, real estate taxes and insurance,

and other operating expenses, are recognized as revenue in the period during which the applicable expenses are incurred and the

tenant’s obligation to reimburse us arises. Income from rentals related to other variable payments is recognized when associated

contingencies are removed.

We assess collectibility from our tenants of future lease payments for each of our operating leases. If we determine that

collectibility is probable, we recognize income from rentals based on the methodology described above. If we determine that

collectibility is not probable, we recognize an adjustment to lower our income from rentals. Furthermore, we may recognize a general

allowance at a portfolio level (not the individual level) if we do not expect to collect future lease payments in full.

For each lease for which we determine that collectibility of future lease payments is not probable, we cease the recognition of

income from rentals on a straight-line basis and limit the recognition of income to the lesser of payments collected from the lessee or

lease income that would have been recognized on a straight-line basis. We do not resume straight-line recognition of income from

rentals for these leases until we determine that the collectibility of future payments related to these leases is probable. We also record a

general allowance related to the deferred rent balances that at the portfolio level (not the individual level) are not expected to be

collected in full through the lease term. As of March 31, 2025 and December 31, 2024, our general allowance balance aggregated

$14.3 million and $21.3 million, respectively.

Direct financing and sales-type leases

Income from rentals related to direct financing and sales-type leases is recognized over the lease term using the effective

interest rate method. At lease commencement, we derecognize the underlying asset classified within investments in real estate and

record net investment in a lease within other assets in our consolidated balance sheets. This initial net investment is determined by

aggregating the present values of the total future lease payments and the estimated residual value of the property, less any unearned

income related to a direct financing lease. Over the lease term, the investment in the lease accretes in value, producing a constant

periodic rate of return on the net investment in the lease. Income from these leases is classified in income from rentals in our

consolidated statements of operations. Our net investment is reduced over time as lease payments are received.

We evaluate our net investment in direct financing and sales-type leases for impairment under the current expected credit

losses accounting standard. For more information, refer to “Provision for expected credit losses” in Note 2 – “Summary of significant

accounting policies” to our unaudited consolidated financial statements.

As a lessor, we classify a lease with variable lease payments that do not depend on an index or a rate as an operating lease

on the commencement date of the lease if both of the following criteria are met:

(i)The lease would have been classified as a sales-type lease or direct financing lease under the current lease accounting

standard; and

(ii)The sales-type lease or direct financing lease classification would have resulted in a selling loss at lease commencement.

We do not derecognize the underlying asset and do not recognize a loss upon lease commencement but continue to

depreciate the underlying asset over its useful life.

17

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

Lessee accounting

We have operating lease agreements in which we are the lessee consisting of ground and office leases. At the lease

commencement date (or at the acquisition date if the lease is acquired as part of a real estate acquisition), we are required to recognize

a liability to account for our future obligations under these operating leases, and a corresponding right-of-use asset.

The lease liability is measured based on the present value of the future lease payments, including payments during the term

under our extension options that we are reasonably certain to exercise. The present value of the future lease payments is calculated for

each operating lease using each respective remaining lease term and a corresponding estimated incremental borrowing rate, which is

the interest rate that we estimate we would have to pay to borrow on a collateralized basis over a similar term for an amount equal to

the lease payments. Subsequently, the lease liability is accreted by applying a discount rate established at the lease commencement

date to the lease liability balance as of the beginning of the period and is reduced by the payments made during the period. We classify

the operating lease liability in accounts payable, accrued expenses, and other liabilities in our consolidated balance sheets.

The right-of-use asset is measured based on the corresponding lease liability, adjusted for initial direct leasing costs and any

other consideration exchanged with the landlord prior to the commencement of the lease, as well as adjustments to reflect favorable or

unfavorable terms of an acquired lease when compared with market terms at the time of acquisition. Subsequently, the right-of-use

asset is amortized on a straight-line basis during the lease term. We classify the right-of-use asset in other assets in our consolidated

balance sheets.

Recognition of revenue arising from contracts with customers

We recognize revenues associated with transactions arising from contracts with customers, excluding revenues subject to the

lease accounting standard discussed in “Lease accounting” above, in accordance with the revenue recognition accounting standard. A

customer is distinguished from a noncustomer by the nature of the goods or services that are transferred. Customers are provided with

goods or services that are generated by a company’s ordinary output activities, whereas noncustomers are provided with nonfinancial

assets that are outside of a company’s ordinary output activities.

We generally recognize revenue representing the transfer of goods and services to customers in an amount that reflects the

consideration to which we expect to be entitled in the exchange. In order to determine the recognition of revenue from customer

contracts, we use a five-step model to (i) identify the contract with the customer, (ii) identify the performance obligations in the contract,

(iii) determine the transaction price, including variable consideration to the extent that it is probable that a significant future reversal will

not occur, (iv) allocate the transaction price to the performance obligations in the contract, and (v) recognize revenue when (or as) we

satisfy the performance obligation.

We identify contractual performance obligations and determine whether revenue should be recognized at a point in time or

over time, based on when control of goods and services transfers to a customer. We consider whether we control the goods or services

prior to the transfer to the customer in order to determine whether we should account for the arrangement as a principal or agent. If we

determine that we control the goods or services provided to the customer, then we are the principal to the transaction, and we recognize

the gross amount of consideration expected in the exchange. If we simply arrange but do not control the goods or services being

transferred to the customer, then we are considered to be an agent to the transaction, and we recognize the net amount of

consideration we are entitled to retain in the exchange.

Total revenues subject to the revenue recognition accounting standard and classified within income from rentals in our

consolidated statements of operations for the three months ended March 31, 2025 included $10.9 million primarily related to short-term

parking revenues associated with long-term lease agreements. Short-term parking revenues do not qualify for the single component

accounting policy, as discussed in “Lessor accounting” in Note 2 – “Summary of significant accounting policies,” due to the difference in

the timing and pattern of transfer of our parking service obligations and associated lease components within the same lease agreement.

We recognize short-term parking revenues in accordance with the revenue recognition accounting standard when the service is

provided and the performance obligation is satisfied, which normally occurs at a point in time.

Monitoring of tenant credit quality

During the term of each lease, we monitor the credit quality and any related material changes of our tenants by (i) monitoring

the credit rating of tenants that are rated by a nationally recognized credit rating agency, (ii) reviewing financial statements of the

tenants that are publicly available or that are required to be delivered to us pursuant to the applicable lease, (iii) monitoring news

reports regarding our tenants and their respective businesses, and (iv) monitoring the timeliness of lease payments.

18

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

Notes receivable

We carry notes receivable at amortized cost, adjusted for an estimated provision for expected credit losses. Interest income on

notes receivable is recognized using the effective interest rate method and is classified within other income in our consolidated

statements of operations. Direct costs incurred in originating notes, along with any premium or discount, are deferred and amortized as

an adjustment to interest income over the note’s term using the effective interest rate method. Notes receivable are classified within

other assets in our consolidated balance sheets. Refer to Note 8 – “Other assets” to our unuaudited consolidated financial statements

for additional details.

Provision for expected credit losses

We are required to estimate and recognize lifetime expected losses, rather than incurred losses, for most of our financial

assets measured at amortized cost and certain other instruments, including trade, notes, and other receivables (excluding receivables

arising from operating leases), loans, held-to-maturity debt securities, net investments in leases arising from sales-type and direct

financing leases, and off-balance-sheet credit exposures (e.g., loan commitments). The recognition of such expected losses, even if the

expected risk of credit loss is remote, typically results in earlier recognition of credit losses. At each reporting date, we reassess our

provision for expected credit losses, and, if necessary, we recognize an adjustment for our current estimate of expected credit losses.

Refer to Note 5 – “Leases” and Note 8 – “Other assets” to our unaudited consolidated financial statements for additional details.

An assessment of the collectibility of operating lease payments and the recognition of an adjustment to lease income based on

this assessment is governed by the lease accounting standard discussed in “Lease accounting” earlier in Note 2 — “Summary of

significant accounting policies” to our unaudited consolidated financial statements.

Income taxes

We are organized and operate as a REIT pursuant to the Internal Revenue Code (the “Code”). Under the Code, a REIT that

distributes at least 90% of its REIT taxable income to its stockholders annually (excluding net capital gains) and meets certain other

conditions is not subject to federal income tax on its distributed taxable income, but could be subject to certain federal, foreign, state,

and local taxes. We distribute 100% of our taxable income annually; therefore, a provision for federal income taxes is not required. In

addition to our REIT returns, we file federal, foreign, state, and local tax returns for our subsidiaries. We file with jurisdictions located in

the U.S., Canada, and other international locations. Our tax returns are subject to routine examination in various jurisdictions for the

2019 through 2024 calendar years.

Employee and non-employee share-based payments

We have implemented an entity-wide accounting policy to account for forfeitures related to unmet service conditions of share-

based awards granted to employees and non-employees when they occur. Under this policy, when forfeitures occur, any previously

recognized expense related to those forfeited awards is reversed in the period of forfeiture.

Our employee and non-employee share-based awards are measured at fair value on the grant date and recognized over the

recipient’s required service period. For share-based awards with performance conditions, we continue to assess the probability of

achieving the performance conditions and recognize expense only when it becomes probable that the performance targets will be met.

Conversely, for share-based awards with market conditions, expense is recognized regardless of whether the market condition is met.

Dividends paid on share-based awards with nonforfeitable dividends are initially classified in retained earnings and reclassified

to compensation cost only if the underlying awards are forfeited. Conversely, for share-based awards with forfeitable dividends,

declared dividends are initially classified in retained earnings and in dividends payable within our consolidated balance sheets. If the

underlying awards are forfeited, the corresponding accrued dividend is reversed in the period of forfeiture. Upon vesting of the

underlying share-based awards with forfeitable dividends, the accumulated dividend payment is made and the dividend payable liability

is settled.

Forward equity sales agreements

From time to time, we enter into forward equity sales agreements and account for them in accordance with the accounting

guidance governing financial instruments and derivatives. Under the accounting guidance, our forward equity sales agreements are not

deemed to be liabilities as they do not embody obligations to repurchase our shares, nor do they embody obligations to issue a variable

number of shares for which the monetary value is predominantly fixed, varied with something other than the fair value of our shares, or

varied inversely in relation to our shares. We also evaluate whether the agreements meet the derivatives and hedging guidance scope

exception to be accounted for as equity instruments. Our forward equity sales agreements are classified as equity contracts based on

the following assessment: (i) none of the agreements’ exercise contingencies are based on observable markets or indices besides

those related to the market for our own stock price and operations; and (ii) none of the settlement provisions preclude the agreements

from being indexed to our own stock.

19

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

Issuer and guarantor subsidiaries of guaranteed securities

Generally, a parent entity of an issuer that holds guaranteed securities must provide separate subsidiary issuer or guarantor

financial statements, unless it qualifies for disclosure exceptions. A parent entity may be eligible for disclosure exceptions if it meets the

following criteria:

(i)The subsidiary issuer or guarantor is a consolidated subsidiary of the parent company, and

(ii)The subsidiary issues a registered security that is:

•issued jointly and severally with the parent company, or

•fully and unconditionally guaranteed by the parent company.

A parent entity that meets the above criteria may instead present summarized financial information (“alternative disclosures”)

either within the consolidated financial statements or in “Management’s discussion and analysis of financial condition and results of

operations” in Item 2. We evaluated the criteria and determined that we are eligible for the disclosure exceptions, which allow us to

provide alternative disclosures; as such, we present alternative disclosures in “Management’s discussion and analysis of financial

condition and results of operations” in Item 2.

Loan fees

Fees incurred in obtaining long-term financing are capitalized and classified with the corresponding debt instrument appearing

on our consolidated balance sheets. Loan fees related to our unsecured senior line of credit are capitalized and classified within other

assets. Capitalized amounts are amortized over the term of the related loan, and the amortization is classified in interest expense in our

consolidated statements of operations.

Distributions from equity method investments

We use the “nature of the distribution” approach to determine the classification within our consolidated statements of cash

flows of cash distributions received from equity method investments, including our unconsolidated real estate joint ventures and equity

method non-real estate investments. Under this approach, distributions are classified based on the nature of the underlying activity that

generated the cash distributions. If we lack the information necessary to apply this approach in the future, we will be required to apply

the “cumulative earnings” approach as an accounting change on a retrospective basis. Under the cumulative earnings approach,

distributions up to the amount of cumulative equity in earnings recognized are classified as cash inflows from operating activities, and

those in excess of that amount are classified as cash inflows from investing activities.

Restricted cash

We present cash and cash equivalents separately from restricted cash within our consolidated balance sheets. However, we

include restricted cash with cash and cash equivalents when reconciling the beginning-of-period and end-of-period total amounts shown

in the consolidated statements of cash flows. We provide a reconciliation between the consolidated balance sheets and the

consolidated statements of cash flows, which is required when the balance includes greater than one line item for cash, cash

equivalents, and restricted cash. We also provide a disclosure of the nature of the restrictions related to material restricted cash

balances.

Recent accounting pronouncements

On November 4, 2024, the FASB issued ASU 2024-03, Disaggregation of Income Statement Expenses, that will require

entities to provide enhanced disclosures related to certain expense categories included in income statement captions. The ASU aims to

increase transparency and provide investors with additional detailed information about the nature of expenses reported on the face of

the income statement. The new standard does not change the requirements for the presentation of expenses on the face of the income

statement.

Under this ASU, entities are required to disaggregate, in a tabular format, expense captions presented on the face of the

income statement — excluding earnings or losses from equity method investments — if they include any of the following expense

categories: purchases of inventory, employee compensation, depreciation, intangible asset amortization, and depreciation or depletion.

For any remaining items within each relevant expense caption, entities must provide a qualitative description of the nature of those

expenses. The new ASU is effective for annual reporting periods beginning after December 15, 2026 and interim reporting periods

beginning after December 15, 2027. Early adoption is permitted. We expect to adopt this ASU on January 1, 2027. Although the

adoption is not expected to have an impact on our financial statements, it is expected to result in incremental disclosures within the

footnotes to our consolidated financial statements.

20

3.INVESTMENTS IN REAL ESTATE

Our consolidated investments in real estate consisted of the following as of March 31, 2025 and December 31, 2024 (in

thousands):

March 31, 2025 December 31, 2024
Rental properties:
Land (related to rental properties) $3,897,927 $3,863,027
Buildings and building improvements 20,788,541 20,377,935
Other improvements 4,530,136 4,354,785
Rental properties 29,216,604 28,595,747
Current and future development and redevelopment projects 8,456,971 8,618,727
Gross investments in real estate 37,673,575 37,214,474
Less: accumulated depreciation (5,768,114) (5,477,082)
Investments in real estate assets held for sale(1) 216,251 372,647
Investments in real estate $32,121,712 $32,110,039

(1)See “Assets held for sale” below.

Assets held for sale

As of March 31, 2025, we had six operating properties aggregating 382,527 RSF and land parcels aggregating 1.9 million SF

that were classified as held for sale.

The disposal of properties classified as held for sale does not represent a strategic shift that has (or will have) a major effect

on our operations or financial results and therefore does not meet the criteria for classification as a discontinued operation. We cease

depreciation of our properties upon their classification as held for sale.

The following is a summary of net assets as of March 31, 2025 and December 31, 2024 for our real estate investments that

were classified as held for sale as of each respective date (in thousands):

March 31, 2025 December 31, 2024
Investments in real estate $216,251 $372,647
Other assets 8,333 9,488
Total assets 224,584 382,135
Total liabilities (13,068) (13,462)
Total accumulated other comprehensive income 2,429 2,584
Net assets classified as held for sale $213,945 $371,257

For additional information, refer to “Real estate sales” in Note 2 – “Summary of significant accounting policies” to our unaudited

consolidated financial statements.

Sales of real estate assets

Our completed dispositions of real estate assets during the three months ended March 31, 2025 consisted of the following

(dollars in thousands):

Square Footage Gain on<br><br>Sales of<br><br>Real Estate
Property Submarket/Market Date of<br><br>Sale Interest<br><br>Sold Operating Land and<br><br>Future Sales Price
Costa Verde by Alexandria University Town Center/<br><br>San Diego 1/31/25 100% 8,730 537,000 124,000
Other 52,352 13,165
176,352 $13,165

All values are in US Dollars.

(1)Represents the aggregate contractual sales price of our dispositions, which differs from proceeds from sales of real estate and contributions from and sales of

noncontrolling interests in our consolidated statement of cash flows under “Investing activities” and “Financing activities,” respectively, primarily due to the timing of

payment, closing costs, and other sales adjustments such as prorations of rents and expenses.

21

3.INVESTMENTS IN REAL ESTATE (continued)

Other

In 2006, ARE-East River Science Park, LLC, a subsidiary of Alexandria Real Estate Equities, Inc., was granted an option to

incorporate a land parcel adjacent to and north of the Alexandria Center® for Life Science – New York City (“ACLS-NYC”) campus

(“Option Parcel”) into the existing ground lease of that campus. The Option Parcel will allow ARE-East River Science Park, LLC to

develop a future world-class life science building within the ACLS-NYC campus. ARE-East River Science Park, LLC’s investment in pre-

construction costs related to the development of the Option Parcel, including costs related to design, engineering, environmental,

survey/title, and permitting and legal costs, aggregated $171.1 million as of March 31, 2025.

On August 6, 2024, ARE-East River Science Park, LLC filed a lawsuit in the United States District Court for the Southern

District of New York against its landlord, New York City Health + Hospitals Corporation (“H+H”), and the New York City Economic

Development Corporation (“EDC”). On January 24, 2025, ARE-East River Science Park, LLC filed a first amended complaint. The

lawsuit alleges two principal claims against H+H and EDC: fraud in the inducement, and, in the alternative, breach of contract in

violation of the implied covenant of good faith and fair dealing. As alleged in the complaint, ARE-East River Science Park, LLC’s claims

arise from H+H’s and EDC’s misrepresentations and concealment of material facts in connection with a floodwall, which H+H and EDC

are seeking to require ARE-East River Science Park, LLC to integrate into the development of the Option Parcel. ARE-East River

Science Park, LLC alleges that H+H’s and EDC’s misconduct have prevented it from commencing the development of the Option

Parcel. In light of the pending litigation, the closing date for our option and thus the commencement date for construction of the third

tower at the campus are presently indeterminate. Among other things, ARE-East River Science Park, LLC is seeking significant

damages and equitable relief from the court to confirm our understanding that the option is in full force and effect.

This matter exposes us to potential losses ranging from zero to the full amount of the investment in the project aggregating

$171.1 million as of March 31, 2025, depending on any collection of damages and/or the ability to develop the project. We performed a

probability-weighted recoverability analysis based on estimates of various possible outcomes and determined no impairment was

present as of March 31, 2025.

22

4.CONSOLIDATED AND UNCONSOLIDATED REAL ESTATE JOINT VENTURES

From time to time, we enter into joint venture agreements through which we own a partial interest in real estate entities that

own, develop, and operate real estate properties. As of March 31, 2025, our real estate joint ventures held the following properties:

Property(1) Market Submarket Our Ownership<br><br>Interest
Consolidated real estate joint ventures:
50 and 60 Binney Street Greater Boston Cambridge/Inner Suburbs 34.0%
75/125 Binney Street Greater Boston Cambridge/Inner Suburbs 40.0%
100 and 225 Binney Street and 300 Third Street Greater Boston Cambridge/Inner Suburbs 30.0%
99 Coolidge Avenue Greater Boston Cambridge/Inner Suburbs 75.7%
15 Necco Street Greater Boston Seaport Innovation District 56.7%
285, 299, 307, and 345 Dorchester Avenue Greater Boston Seaport Innovation District 60.0%
Alexandria Center® for Science and Technology –<br><br>Mission Bay(2) San Francisco Bay Area Mission Bay 25.0%
601, 611, 651, 681, 685, and 701 Gateway<br><br>Boulevard San Francisco Bay Area South San Francisco 50.0%
751 Gateway Boulevard San Francisco Bay Area South San Francisco 51.0%
211 and 213 East Grand Avenue San Francisco Bay Area South San Francisco 30.0%
500 Forbes Boulevard San Francisco Bay Area South San Francisco 10.0%
Alexandria Center® for Life Science – Millbrae San Francisco Bay Area South San Francisco 48.3%
3215 Merryfield Row San Diego Torrey Pines 30.0%
Campus Point by Alexandria(3) San Diego University Town Center 55.0%
5200 Illumina Way San Diego University Town Center 51.0%
9625 Towne Centre Drive San Diego University Town Center 30.0%
SD Tech by Alexandria(4) San Diego Sorrento Mesa 50.0%
Pacific Technology Park San Diego Sorrento Mesa 50.0%
Summers Ridge Science Park(5) San Diego Sorrento Mesa 30.0%
1201 and 1208 Eastlake Avenue East Seattle Lake Union 30.0%
199 East Blaine Street Seattle Lake Union 30.0%
400 Dexter Avenue North Seattle Lake Union 30.0%
800 Mercer Street Seattle Lake Union 60.0%
Unconsolidated real estate joint ventures(6):
1655 and 1725 Third Street San Francisco Bay Area Mission Bay 10.0%
1450 Research Boulevard Maryland Rockville 73.2% (7)
101 West Dickman Street Maryland Beltsville 58.4% (7)

(1)Refer to the table on the next page that shows the categorization of our joint ventures under the consolidation framework.

(2)Includes 409 and 499 Illinois Street, 1450, 1500, and 1700 Owens Street, and 455 Mission Bay Boulevard South.

(3)Includes 10210, 10260, 10290, and 10300 Campus Point Drive and 4110, 4135, 4155, 4161, 4165, 4224, and 4242 Campus Point Court.

(4)Includes 9605, 9645, 9675, 9725, 9735, 9805, 9808, 9855, and 9868 Scranton Road and 10055, 10065, and 10075 Barnes Canyon Road.

(5)Includes 9965, 9975, 9985, and 9995 Summers Ridge Road.

(6)In addition to the real estate joint ventures listed, we hold an interest in one insignificant unconsolidated real estate joint venture in North America.

(7)Represents a joint venture with a local real estate operator in which our joint venture partner manages the day-to-day activities that significantly affect the economic

performance of the joint venture.

Our consolidation policy is described under “Consolidation” in Note 2 – “Summary of significant accounting policies” to our

unaudited consolidated financial statements. Consolidation accounting is highly technical, but its framework is primarily based on the

controlling financial interests and benefits of the joint ventures. We generally consolidate a joint venture that is a legal entity that we

control (i.e., we have the power to direct the activities of the joint venture that most significantly affect its economic performance)

through contractual rights, regardless of our ownership interest, and where we determine that we have benefits through the allocation of

earnings or losses and fees paid to us that could be significant to the joint venture (the “VIE model”).

23

4.CONSOLIDATED AND UNCONSOLIDATED REAL ESTATE JOINT VENTURES (continued)

We also generally consolidate joint ventures when we have a controlling financial interest through voting rights and where our

voting interest is greater than 50% (the “voting model”). Voting interest differs from ownership interest for some joint ventures. We

account for joint ventures that do not meet the consolidation criteria under the equity method of accounting by recognizing our share of

income and losses.

The table below shows the categorization of our real estate joint ventures under the consolidation framework:

Property(1) Consolidation<br><br>Model Voting Interest Consolidation Analysis Conclusion
50 and 60 Binney Street VIE model Not applicable<br><br>under VIE<br><br>model Consolidated
75/125 Binney Street We have:
100 and 225 Binney Street and 300<br><br>Third Street
99 Coolidge Avenue (i) The power to direct the<br><br>activities of the joint venture<br><br>that most significantly affect its<br><br>economic performance; and
15 Necco Street
285, 299, 307, and 345 Dorchester<br><br>Avenue
Alexandria Center® for Science and<br><br>Technology – Mission Bay
601, 611, 651, 681, 685, and 701<br><br>Gateway Boulevard
751 Gateway Boulevard
211 and 213 East Grand Avenue (ii) Benefits that can be significant<br><br>to the joint venture.
500 Forbes Boulevard
Alexandria Center® for Life Science –<br><br>Millbrae
3215 Merryfield Row
Campus Point by Alexandria
5200 Illumina Way Therefore, we are the primary<br><br>beneficiary of each VIE
9625 Towne Centre Drive
SD Tech by Alexandria
Pacific Technology Park
Summers Ridge Science Park
1201 and 1208 Eastlake Avenue East
199 East Blaine Street
400 Dexter Avenue North
800 Mercer Street
1450 Research Boulevard We do not control the joint venture<br><br>and are therefore not the primary<br><br>beneficiary. Equity method<br><br>of accounting
101 West Dickman Street
1655 and 1725 Third Street Voting model Does not<br><br>exceed 50% Our voting interest is 50% or less.

(1)In addition to the real estate joint ventures listed, we hold an interest in one insignificant unconsolidated real estate joint venture in North America.

24

4.CONSOLIDATED AND UNCONSOLIDATED REAL ESTATE JOINT VENTURES (continued)

Consolidated VIEs’ balance sheet information

We, together with joint venture partners, hold interests in real estate joint ventures that we consolidate in our financial

statements. These existing joint ventures provide significant equity capital to fund a portion of our future construction spend, and our

joint venture partners may also contribute equity into these entities for financing-related activities.

The table below aggregates the balance sheet information of our consolidated VIEs (in thousands):

March 31, 2025 December 31, 2024
Investments in real estate $7,756,140 $8,917,718
Cash and cash equivalents 226,533 335,223
Other assets 736,953 777,033
Total assets $8,719,626 $10,029,974
Secured notes payable $150,220 $149,321
Other liabilities 432,154 626,460
Total liabilities 582,374 775,781
Redeemable noncontrolling interests 10,360
Alexandria Real Estate Equities, Inc.’s share of equity 3,611,953 4,754,386
Noncontrolling interests’ share of equity 4,525,299 4,489,447
Total liabilities and equity $8,719,626 $10,029,974

In determining whether to aggregate the balance sheet information of consolidated VIEs, we considered the similarity of each

VIE, including the primary purpose of these entities to own, manage, operate, and lease real estate properties owned by the VIEs, and

the similar nature of our involvement in each VIE as a managing member. Due to the similarity of the characteristics, we present the

balance sheet information of these entities on an aggregated basis. None of our consolidated VIEs’ assets have restrictions that limit

their use to settle specific obligations of the VIE. There are no creditors or other partners of our consolidated VIEs that have recourse to

our general credit, and our maximum exposure to our consolidated VIEs is limited to our variable interests in each VIE, except for our

99 Coolidge Avenue real estate joint venture in which the VIE’s secured construction loan is guaranteed by us. Refer to Note 10 –

“Secured and unsecured senior debt” to our unaudited consolidated financial statements for additional information.

Unconsolidated real estate joint ventures

Our maximum exposure to our unconsolidated VIEs is limited to our investment in each VIE, except for our 1450 Research

Boulevard and 101 West Dickman Street unconsolidated real estate joint ventures in which we guarantee up to $6.7 million of the

outstanding balance related to each VIE’s secured loan. Our investments in unconsolidated real estate joint ventures, accounted for

under the equity method and classified in investments in unconsolidated real estate joint ventures in our consolidated balance sheets,

consisted of the following as of March 31, 2025 and December 31, 2024 (in thousands):

Property March 31, 2025 December 31, 2024
1655 and 1725 Third Street $21,224 $10,574
1450 Research Boulevard 8,941 9,193
101 West Dickman Street 9,629 9,749
Other 10,292 10,357
$50,086 $39,873

Below are key terms of unconsolidated real estate joint ventures’ secured loans as of March 31, 2025 (dollars in thousands):

Interest<br><br>Rate(1) At 100%
Unconsolidated Joint Venture Maturity Date Stated Rate Aggregate<br><br>Commitment Debt Balance(2)
101 West Dickman Street 11/10/26 SOFR+1.95% (3) 6.35% $26,750 19,139
1450 Research Boulevard 12/10/26 SOFR+1.95% (3) 6.41% 13,000 8,998
1655 and 1725 Third Street(4) 2/10/35 6.37% 6.44% 500,000 496,658
$539,750 524,795

All values are in US Dollars.

(1)Includes interest expense and amortization of loan fees.

(2)Represents outstanding principal, net of unamortized deferred financing costs, as of March 31, 2025.

(3)This loan is subject to a fixed SOFR floor of 0.75%.

(4)During the three months ended March 31, 2025, the unconsolidated real estate joint venture refinanced $500 million of its $600 million fixed-rate debt with a new

secured note payable maturing in 2035. The remaining debt balance of approximately $100 million was repaid through contributions from the unconsolidated joint

venture partners, including our share of $10.8 million. As of March 31, 2025, our investment in this unconsolidated real estate joint venture was $21.2 million.

25

5.LEASES

Refer to “Lease accounting” in Note 2 – “Summary of significant accounting policies” to our unaudited consolidated financial

statements for information about lease accounting standards that set principles for the recognition, measurement, presentation, and

disclosure of leases for both parties to a lease agreement (i.e., lessees and lessors).

Leases in which we are the lessor

As of March 31, 2025, we had 386 properties aggregating 39.6 million operating RSF in key cluster locations, including

Greater Boston, the San Francisco Bay Area, San Diego, Seattle, Maryland, Research Triangle, and New York City. We primarily focus

on developing Class A/A+ properties in AAA life science innovation cluster locations that offer the scale and strategic design integral to

our Megacampus strategy. Strategically located near top academic and medical research institutions and equipped with curated

amenities and services, and convenient access to transit, our Megacampus ecosystems are designed to support our tenants in

attracting and retaining top talent, which we believe is a key driver of tenant demand for our properties.

As of March 31, 2025, all leases in which we are the lessor were classified as operating leases, with the exception of one

direct financing and one sales-type lease. Our leases are described below.

Operating leases

As of March 31, 2025, our 386 properties were subject to operating lease agreements. Seven of these properties are subject

to operating lease agreements that each contain a purchase option as described below:

(i)Two of these properties, representing two land parcels in our San Francisco Bay Area market, are subject to lease

agreements that each contain an option for the lessee to purchase the underlying asset from us at fair market value during

each of the 30-day periods commencing on the dates that are 15 years, 30 years, and 74.5 years after the rent

commencement date of October 1, 2017. The remaining lease term related to each of the two land parcels is 67.7 years.

(ii)Two operating properties in our Seattle market, held by a consolidated real estate joint venture, are subject to purchase

options held by our partner in this joint venture, which is also a tenant at these properties. One purchase option allows our

partner to purchase our 30% interest in one property for $40.0 million in 2031. Contingent upon the exercise of this option,

the second purchase option allows our partner to purchase our 30% interest in one property for $69.1 million in 2034. Our

partner’s remaining lease terms for these operating leases are 5.9 years and 19.5 years, respectively.

(iii)Three properties subject to operating lease agreements that contain purchase options with a weighted-average (based on

property RSF) exercise date in October 2027.

Certain operating leases contain options for the tenant to extend their lease at prevailing market rates at the time of expiration.

In addition, certain operating leases contain an early termination option that requires advance notification and payment of an early

termination fee by the tenant.

At the commencement of each lease, we establish the lease term comprising the noncancelable period for each lease together

with periods covered by options to extend or terminate the lease that we determine the lessee is reasonably certain to exercise. Our

assessment of whether a lessee is reasonably certain to exercise or not exercise an option considers all economic factors relevant to

the assessment, including property-based, market-based, and tenant-based factors. We do not reassess the lease term or a lessee

option to purchase the underlying asset unless there is a lease modification that is not accounted for as a separate contract.

Future lease payments to be received under the terms of our operating lease agreements, excluding expense

reimbursements, in effect as of March 31, 2025 are outlined in the table below (in thousands):

Year Amount
2025 $1,408,110
2026 1,804,811
2027 1,737,372
2028 1,609,987
2029 1,486,702
Thereafter 9,741,742
Total $17,788,724

Refer to Note 3 – “Investments in real estate” to our unaudited consolidated financial statements for additional information

about our owned real estate assets, which are the underlying assets under our operating leases.

26

5.LEASES (continued)

Direct financing and sales-type leases

As of March 31, 2025, we have one direct financing lease agreement, with a net investment balance of $41.7 million, for a

parking structure with a remaining lease term of 67.7 years. The lessee has an option to purchase the underlying asset at fair market

value during each of the 30-day periods commencing on the dates that are 15 years, 30 years, and 74.5 years after the rent

commencement date of October 1, 2017.

As of March 31, 2025, we also have one sales-type lease for an operating property located in the Seattle market with the lease

term through August 2025. As of March 31, 2025, the net investment in this lease is $18.6 million. Upon recognition of the sales-type

lease during the three months ended March 31, 2025, we recognized a gain on sale of real estate aggregating $12.7 million classified in

gain on sales of real estate within our unaudited consolidated statement of operations for the three months ended March 31, 2025.

The components of our aggregate net investment in our direct financing lease and our sales-type lease as of March 31, 2025

and December 31, 2024 are summarized in the table below (in thousands):

March 31, 2025 December 31, 2024
Gross investment in direct financing and sales-type leases $269,532 $251,405
Less: unearned income on direct financing lease (207,064) (207,734)
Less: provision for expected credit losses (2,217) (2,168)
Net investment in leases $60,251 $41,503

As of March 31, 2025, our estimated provision for expected credit loss related to our direct financing lease aggregated

$2.2 million, unchanged from December 31, 2024. We estimate the provision for expected credit loss related to our direct financing

lease using a probability of default methodology, which incorporates the borrower’s investment-grade credit rating from S&P, to evaluate

the probability of default. Additionally, we incorporate the projected value of the real estate securing the investments to estimate

potential recoveries in the event of default, among other inputs.

During the three months ended March 31, 2025, we recognized an estimated provision for expected credit loss aggregating

$49 thousand related to our sales-type lease. This estimate was determined using historical industry losses and considering

transaction-specific information, including the estimated fair value of the underlying real estate asset securing this transaction, the short-

term nature of this lease, and other available information.

We classify adjustments to estimated provision for expected credit loss related to our direct financing and sales-type leases

within other income in our consolidated statement of operations. For further details, refer to “Provision for expected credit losses” in

Note 2 – “Summary of significant accounting policies” to our unaudited consolidated financial statements.

Future lease payments to be received under the terms of our direct financing lease and our sales-type lease as of March 31,

2025 are outlined in the table below (in thousands):

Year Total
2025 $20,103
2026 2,036
2027 2,097
2028 2,160
2029 2,224
Thereafter 240,912
Total $269,532

27

5.LEASES (continued)

Income from rentals

Our income from rentals includes revenue related to agreements for the rental of our real estate, which primarily includes

revenues subject to the lease accounting standard and the revenue recognition accounting standard as shown below (in thousands):

Three Months Ended March 31,
2025 2024
Income from rentals:
Revenues subject to the lease accounting standard:
Operating leases $731,421 $746,061
Direct financing and sales-type leases 810 659
Revenues subject to the lease accounting standard 732,231 746,720
Revenues subject to the revenue recognition accounting standard 10,944 8,831
Income from rentals $743,175 $755,551

Our revenues that are subject to the revenue recognition accounting standard and are classified in income from rentals consist

primarily of short-term parking revenues that are not considered lease revenues under the lease accounting standard. Refer to

“Revenues” and “Recognition of revenue arising from contracts with customers” in Note 2 – “Summary of significant accounting policies”

to our unaudited consolidated financial statements for additional information.

Residual value risk management strategy

Our leases do not have guarantees of residual value on the underlying assets. We manage risk associated with the residual

value of our leased assets by (i) evaluating each potential acquisition of real estate to determine whether it meets our business

objective to invest primarily in high-demand markets, (ii) directly managing our leased properties, conducting frequent property

inspections, proactively addressing potential maintenance issues before they arise, and/or timely resolving any occurring issues, and

(iii) carefully selecting our tenants and monitoring their credit quality throughout their respective lease terms.

Leases in which we are the lessee

Operating lease agreements

We have operating lease agreements in which we are the lessee consisting of ground and office leases. Certain of these

leases have options to extend or terminate the contract terms upon meeting certain criteria. There are no notable restrictions or

covenants imposed by the leases, nor guarantees of residual value.

We recognize a right-of-use asset, which is classified within other assets in our consolidated balance sheets, and a related

liability, which is classified within accounts payable, accrued expenses, and other liabilities in our consolidated balance sheets, to

account for our future obligations under ground and office lease arrangements in which we are the lessee. Refer to “Lessee accounting”

in Note 2 – “Summary of significant accounting policies” to our unaudited consolidated financial statements.

As of March 31, 2025, the present value of the remaining contractual payments aggregating $809.0 million under our operating

lease agreements, including our extension options that we are reasonably certain to exercise, was $371.4 million. Our corresponding

operating lease right-of-use assets, adjusted for initial direct leasing costs and other consideration exchanged with the landlord prior to

the commencement of the lease, aggregated $728.9 million. As of March 31, 2025, the weighted-average remaining lease term of

operating leases in which we are the lessee was approximately 54 years, including extension options that we are reasonably certain to

exercise, and the weighted-average discount rate was 4.7%. The weighted-average discount rate is based on the incremental

borrowing rate estimated for each lease, which is the interest rate that we estimate we would have to pay to borrow on a collateralized

basis over a similar term for an amount equal to the lease payments.

Ground lease obligations as of March 31, 2025 included leases for 32 of our properties, which accounted for approximately 8%

of our total number of properties. Excluding one ground lease that expires in 2036 related to one operating property with a net book

value of $5.5 million as of March 31, 2025, our ground lease obligations have remaining lease terms ranging from approximately 29 to

82 years, including extension options that we are reasonably certain to exercise.

28

5.LEASES (continued)

The reconciliation of future lease payments under noncancelable operating leases in which we are the lessee to the operating

lease liability reflected in our unaudited consolidated balance sheet as of March 31, 2025 is in the table below (in thousands):

Year Total
2025 $17,434
2026 23,081
2027 22,162
2028 21,830
2029 21,338
Thereafter 703,148
Total future payments under our operating leases in which we are the lessee 808,993
Effect of discounting (437,581)
Operating lease liability $371,412

Lessee operating costs

Operating lease costs relate to our ground and office leases in which we are the lessee. Ground leases generally require fixed

annual rent payments and may also include escalation clauses and renewal options. For the three months ended March 31, 2025 and

2024, amounts paid and classified as operating activities in our unaudited consolidated statements of cash flows for leases in which we

are the lessee aggregated $144.6 million and $7.3 million, respectively. The increase is primarily due to the ground lease prepayment of

$135.0 million made in January 2025 for a 24-year lease term extension to our existing ground lease agreement at the Alexandria

Technology Square® Megacampus in our Cambridge submarket.

Our operating lease obligations related to our office leases have remaining terms of up to 11 years, exclusive of extension

options. For the three months ended March 31, 2025 and 2024, our costs of operating leases in which we are the lessee were as

follows (in thousands):

Three Months Ended March 31,
2025 2024
Gross operating lease costs $12,359 $9,211
Capitalized lease costs (693) (528)
Expenses for operating leases in which we are the lessee $11,666 $8,683

During the three months ended March 31, 2025, we recognized an impairment charge aggregating $32.2 million, related to a

ground lease entered into in 2021 for a future development site in our San Francisco Bay Area market. As of December 31, 2024, we

had a right-of-use asset aggregating $32.4 million related to our investment into this ground lease. During the three months ended

March 31, 2025, based on our current financial outlook for this project, we made the determination to no longer proceed with this

project. Consequently, we recognized an impairment charge aggregating $32.2 million to write off our remaining right-of-use asset

balance. As of March 31, 2025 and December 31, 2024, we had no operating lease liability associated with this ground lease, as the

related lease obligation had been fully prepaid. We do not expect to make additional future payments in connection with this ground

lease.

6. CASH, CASH EQUIVALENTS, AND RESTRICTED CASH

Cash, cash equivalents, and restricted cash consisted of the following as of March 31, 2025 and December 31, 2024 (in

thousands):

March 31, 2025 December 31, 2024
Cash and cash equivalents $476,430 $552,146
Restricted cash:
Funds held in escrow for real estate acquisitions 2,954 2,954
Other 4,370 4,747
7,324 7,701
Total $483,754 $559,847

29

7.INVESTMENTS

We hold investments in publicly traded companies and privately held entities primarily involved in the life science industry. As a

REIT, we generally limit our ownership of each individual entity’s voting stock to less than 10%. We evaluate each investment to

determine whether we have the ability to exercise significant influence, but not control, over an investee. We evaluate investments in

which our ownership is equal to or greater than 20%, but less than or equal to 50%, of an investee’s voting stock with a presumption

that we have this ability. For our investments in limited partnerships that maintain specific ownership accounts, we presume that such

ability exists when our ownership interest exceeds 3% to 5%. In addition to our ownership interest, we consider whether we have a

board seat or whether we participate in the investee’s policy-making process, among other criteria, to determine if we have the ability to

exert significant influence, but not control, over an investee. If we determine that we have such ability, we account for the investment

under the equity method, as described below.

From time to time, we may hold equity investments in publicly traded companies that are subject to temporary contractual sale

restrictions. We do not recognize a discount related to such contractual sale restrictions.

Investments accounted for under the equity method

Under the equity method of accounting, we initially recognize our investment at cost and subsequently adjust the carrying

amount of the investment for our share of earnings or losses reported by the investee, distributions received, and other-than-temporary

impairments.

As of March 31, 2025, we had nine investments in limited partnerships maintaining specific ownership accounts for each

investor, which were accounted for under the equity method. These investments aggregated $224.6 million. Our ownership interest in

each of these nine investments was greater than 5%.

Investments that do not qualify for the equity method of accounting

For investees over which we determine that we do not have the ability to exercise significant influence or control, we account

for each investment depending on whether it is an investment in a (i) publicly traded company, (ii) privately held entity that reports NAV

per share, or (iii) privately held entity that does not report NAV per share, as described below.

Investments in publicly traded companies

Our investments in publicly traded companies are classified as investments with readily determinable fair values and are

presented at fair value in our consolidated balance sheets, with changes in fair value classified in investment income (loss) in our

consolidated statements of operations. The fair values for our investments in publicly traded companies are determined based on sales

prices or quotes available on securities exchanges.

Investments in privately held companies

Our investments in privately held entities without readily determinable fair values consist of (i) investments in privately held

entities that report NAV per share and (ii) investments in privately held entities that do not report NAV per share. These investments are

accounted for as follows:

Investments in privately held entities that report NAV per share

Investments in privately held entities that report NAV per share, such as our privately held investments in limited partnerships,

are presented at fair value using NAV as a practical expedient, with changes in fair value classified in investment income (loss) in our

consolidated statements of operations. We use NAV per share reported by limited partnerships generally without adjustment, unless we

are aware of information indicating that the NAV reported by a limited partnership does not accurately reflect the fair value of the

investment at our reporting date.

Investments in privately held entities that do not report NAV per share

Investments in privately held entities that do not report NAV per share are accounted for using a measurement alternative

under which these investments are measured at cost, adjusted for observable price changes and impairments, with changes classified

in investment income (loss) in our consolidated statements of operations.

An observable price arises from an orderly transaction for an identical or similar investment of the same issuer, which is

observed by an investor without expending undue cost and effort. Observable price changes result from, among other things, equity

transactions of the same issuer executed during the reporting period, including subsequent equity offerings or other reported equity

transactions related to the same issuer. To determine whether these transactions are indicative of an observable price change, we

evaluate, among other factors, whether these transactions have similar rights and obligations, including voting rights, distribution

preferences, and conversion rights to the investments we hold.

30

7.INVESTMENTS (continued)

Impairment evaluation of equity method investments and investments in privately held entities that do not report NAV per share

We monitor equity method investments and investments in privately held entities that do not report NAV per share for new

developments, including operating results, prospects and results of clinical trials, new product initiatives, new collaborative agreements,

capital-raising events, and merger and acquisition activities. These investments are evaluated on the basis of a qualitative assessment

for indicators of impairment by monitoring the presence of the following triggering events or impairment indicators:

(i)a significant deterioration in the earnings performance, credit rating, asset quality, or business prospects of the investee;

(ii)a significant adverse change in the regulatory, economic, or technological environment of the investee;

(iii)a significant adverse change in the general market condition, including the research and development of technology and

products that the investee is bringing or attempting to bring to the market;

(iv)significant concerns about the investee’s ability to continue as a going concern; and/or

(v)a decision by investors to cease providing support or reduce their financial commitment to the investee.

If such indicators are present, we are required to estimate the investment’s fair value and immediately recognize an

impairment charge in an amount equal to the investment’s carrying value in excess of its estimated fair value.

Investment income/loss recognition and classification

We recognize both realized and unrealized gains and losses in our consolidated statements of operations, classified in

investment income (loss) in our consolidated statements of operations. Unrealized gains and losses represent:

(i)changes in fair value for investments in publicly traded companies;

(ii)changes in NAV for investments in privately held entities that report NAV per share;

(iii)observable price changes for investments in privately held entities that do not report NAV per share; and

(iv)our share of unrealized gains or losses reported by our equity method investees.

Realized gains and losses on our investments represent the difference between proceeds received upon disposition of

investments and their historical or adjusted cost basis. For our equity method investments, realized gains and losses represent our

share of realized gains or losses reported by the investee. Impairments are realized losses, which result in an adjusted cost basis, and

represent charges to reduce the carrying values of investments in privately held entities that do not report NAV per share and equity

method investments, if impairments are deemed other than temporary, to their estimated fair value.

Funding commitments to investments in privately held entities that report NAV

We are committed to funding approximately $363.9 million for our investments in privately held entities that report NAV. Our

funding commitments expire at various dates over the next 12 years, with a weighted-average expiration of 8.1 years as of March 31,

  1. These investments are not redeemable by us, but we may receive distributions from these investments throughout their terms.

Our investments in privately held entities that report NAV generally have expected initial terms in excess of 10 years. The weighted-

average remaining term during which these investments are expected to be liquidated was 5.2 years as of March 31, 2025.

31

7.INVESTMENTS (continued)

The following tables summarize our investments as of March 31, 2025 and December 31, 2024 (in thousands):

March 31, 2025
Cost Unrealized<br><br>Gains Unrealized<br><br>Losses Carrying<br><br>Amount
Publicly traded companies $182,797 $24,425 $(122,472) $84,750
Entities that report NAV 511,907 105,405 (42,327) 574,985
Entities that do not report NAV:
Entities with observable price changes 106,465 75,087 (8,255) 173,297
Entities without observable price changes 422,052 422,052
Investments accounted for under the equity method N/A N/A N/A 224,604
Total investments $1,223,221 $204,917 $(173,054) $1,479,688 December 31, 2024
--- --- --- --- ---
Cost Unrealized<br><br>Gains Unrealized<br><br>Losses Carrying<br><br>Amount
Publicly traded companies $188,653 $24,262 $(107,248) $105,667
Entities that report NAV 518,074 126,077 (34,285) 609,866
Entities that do not report NAV:
Entities with observable price changes 99,932 77,761 (2,956) 174,737
Entities without observable price changes 400,487 400,487
Investments accounted for under the equity method N/A N/A N/A 186,228
Total investments $1,207,146 $228,100 $(144,489) $1,476,985

Cumulative gains and losses (realized and unrealized) on investments in privately held entities that do not report NAV still held

as of March 31, 2025 aggregated to a loss of $86.3 million, which consisted of upward adjustments aggregating $75.1 million,

downward adjustments aggregating $8.3 million, and impairments aggregating $153.1 million.

Our investment income (loss) for the three months ended March 31, 2025 and 2024 consisted of the following (in thousands):

Three Months Ended March 31,
2025 2024
Realized gains 18,153 $14,126
Unrealized (losses) gains (68,145) 29,158
Investment (loss) income (49,992) $43,284

All values are in US Dollars.

(1)Consists of realized gains of $29.3 million, partially offset by impairment charges of $11.2 million during the three months ended March 31, 2025.

During the three months ended March 31, 2025, gains and losses on investments in privately held entities that do not report

NAV still held as of March 31, 2025 aggregated to a loss of $9.0 million, which consisted of upward adjustments aggregating

$8.1 million and downward adjustments and impairments aggregating $17.1 million.

During the three months ended March 31, 2024, gains and losses on investments in privately held entities that do not report

NAV still held as of March 31, 2024 aggregated to a loss of $10.5 million, which consisted of upward adjustments aggregating

$8.6 million and downward adjustments and impairments aggregating $19.1 million.

Unrealized gains or losses related to investments still held (excluding investments accounted for under the equity method) as

of March 31, 2025 and 2024 aggregated to a loss of $37.4 million and a gain of $47.8 million during the three months ended

March 31, 2025 and 2024, respectively.

Our investment loss of $50.0 million for the three months ended March 31, 2025 also included $895 thousand of equity in

losses of our equity method investments.

Refer to “Investments” in Note 2 – “Summary of significant accounting policies” to our unaudited consolidated financial

statements for additional information.

32

8. OTHER ASSETS

The following table summarizes the components of other assets as of March 31, 2025 and December 31, 2024 (in thousands):

March 31, 2025 December 31, 2024
Acquired in-place leases 280,421 $305,144
Deferred compensation plan 47,746 47,727
Deferred financing costs – unsecured senior line of credit 46,644 49,056
Deposits 31,112 21,768
Furniture, fixtures, equipment, and software 47,279 39,558
Net investment in leases 60,251 41,503
Notes receivable 213,040 120,546
Operating lease right-of-use assets 728,873 764,472
Other assets 100,289 96,690
Prepaid expenses 62,405 33,567
Property, plant, and equipment 140,382 141,275
Total 1,758,442 $1,661,306

All values are in US Dollars.

(1)Refer to “Leases in which we are the lessee" section within Note 5 – Leases for information about the decrease in this balance since December 31, 2024.

Notes receivable

Our notes receivable as of March 31, 2025 and December 31, 2024 consisted of the following (dollars in thousands):

As of March 31, 2025
Weighted Average
Notes Receivable Effective<br><br>Interest Rate Maturity<br><br>Date Balance December 31, 2024
Secured by real estate assets in San Diego 10.1% 11/4/28 $195,969 $103,427
Secured by real estate assets in Greater Boston 4.6% 12/16/29 17,544 17,356
Less: provision for expected credit losses (473) (237)
Notes receivable $213,040 $120,546

Our notes receivable represent held-to-maturity debt securities carried at amortized costs and are generally secured by real

estate. Under the current expected credit losses accounting standard, we are required to estimate and, if necessary, recognize a

provision for expected credit losses related to these notes. We do not have a history of losses on such securities; therefore, we utilize

available information on historical losses for the commercial real estate industry. We determine expected credit losses for our notes

receivable using historical industry losses and considering loan-specific information, including credit ratings of the borrowers, estimated

fair values of underlying real estate assets, loan-to-value ratios, the presence of guarantors, and/or other available information. During

the three months ended March 31, 2025, we recognized a provision for expected credit losses of $236 thousand related to our notes

receivable, reducing the notes receivable balance in our unaudited consolidated balance sheet and other income in our unaudited

consolidated statement of operations accordingly. This provision is reevaluated on an ongoing basis, with any necessary adjustments

recognized in the corresponding period.

33

9.FAIR VALUE MEASUREMENTS

We provide fair value information about all financial instruments for which it is practicable to estimate fair value. We measure

and disclose the estimated fair value of financial assets and liabilities by utilizing a fair value hierarchy that distinguishes between data

obtained from sources independent of the reporting entity and the reporting entity’s own assumptions about market participant

assumptions. This hierarchy consists of three broad levels, as follows: (i) quoted prices in active markets for identical assets or liabilities

(Level 1), (ii) significant other observable inputs (Level 2), and (iii) significant unobservable inputs (Level 3). Significant other observable

inputs can include quoted prices for similar assets or liabilities in active markets, as well as inputs that are observable for the asset or

liability, such as interest rates, foreign exchange rates, and yield curves. Significant unobservable inputs are typically based on an

entity’s own assumptions, since there is little, if any, related market activity. In instances in which the determination of the fair value

measurement is based on inputs from different levels of the fair value hierarchy, the level in the fair value hierarchy within which the

entire fair value measurement falls is based on the lowest level of input that is significant to the fair value measurement in its entirety.

Our assessment of the significance of a particular input to the fair value measurement in its entirety requires judgment and considers

factors specific to the asset or liability.

Assets and liabilities measured at fair value on a recurring basis

The following table sets forth the assets that we measure at fair value on a recurring basis by level in the fair value hierarchy

(in thousands). There were no liabilities measured at fair value on a recurring basis as of March 31, 2025 and December 31, 2024.

There were no transfers of assets measured at fair value on a recurring basis to or from Level 3 in the fair value hierarchy during the

three months ended March 31, 2025.

Fair Value Measurement Using
Description Total Quoted Prices in<br><br>Active Markets<br><br>for Identical Assets<br><br>(Level 1) Significant<br><br>Other<br><br>Observable Inputs<br><br>(Level 2) Significant<br><br>Unobservable<br><br>Inputs<br><br>(Level 3)
Investments in publicly traded companies:
As of March 31, 2025 $84,750 $84,750 $— $—
As of December 31, 2024 $105,667 $105,667 $— $—

Our investments in publicly traded companies represent investments with readily determinable fair values, and are carried at

fair value, with changes in fair value classified in investment income (loss) in our consolidated financial statements. We also hold

investments in privately held entities, which consist of (i) investments that report NAV and (ii) investments that do not report NAV, as

further described below.

Our investments in privately held entities that report NAV, such as our privately held investments in limited partnerships, are

carried at fair value using NAV as a practical expedient, with changes in fair value classified in net income. As of March 31, 2025 and

December 31, 2024, the carrying values of investments in privately held entities that report NAV aggregated $575.0 million and

$609.9 million, respectively. These investments are excluded from the fair value hierarchy above as required by the fair value

accounting standard. We estimate the fair value of each of our investments in limited partnerships based on the most recent NAV

reported by each limited partnership. As a result, the determination of fair values of our investments in privately held entities that report

NAV generally does not involve significant estimates, assumptions, or judgments.

34

9.FAIR VALUE MEASUREMENTS (continued)

Assets and liabilities measured at fair value on a nonrecurring basis

The following table sets forth the assets measured at fair value on a nonrecurring basis by level within the fair value hierarchy

as of March 31, 2025 and December 31, 2024 (in thousands).

Fair Value Measurement Using
Description Carrying Amount Quoted Prices in<br><br>Active Markets<br><br>for Identical Assets<br><br>(Level 1) SignificantOtherObservable Inputs(Level 2) Significant<br><br>Unobservable<br><br>Inputs<br><br>(Level 3)
Real estate assets held for sale with carrying<br><br>values adjusted to fair value less costs to sell:
As of March 31, 2025 169,745 $— $169,745 (2)
As of December 31, 2024 322,662 $— $322,662 (2)
Investments in privately held entities that do not<br><br>report NAV:
As of March 31, 2025 184,126 $— 173,297 $10,829 (4)
As of December 31, 2024 184,236 $— 174,737 $9,499 (4)

All values are in US Dollars.

(1)These amounts are included in the total balances of our net assets classified as held for sale aggregating $213.9 million and $371.3 million as of March 31, 2025 and

December 31, 2024, respectively, disclosed in Note 3 – “Investments in real estate,” and represent assets held for sale as of March 31, 2025 and December 31, 2024,

respectively, for which impairments were recognized.

(2)These amounts represent the aggregate carrying amounts of assets held for sale after adjustments to their respective fair values less costs to sell based on executed

purchase and sale agreements, letters of intent, or valuations provided by third-party real estate brokers.

(3)These amounts represent the total carrying amounts of our equity investments in privately held entities with observable price changes, which are included in the

investments balances of $1.5 billion in our unaudited consolidated balance sheets as of March 31, 2025 and December 31, 2024, disclosed in Note 7 – “Investments” to

our unaudited consolidated financial statements.

(4)These amounts are included in the investments in privately held entities without observable price changes balances aggregating $422.1 million and $400.5 million as of

March 31, 2025 and December 31, 2024, respectively, disclosed in Note 7 – “Investments” to our unaudited consolidated financial statements. The aforementioned

balances represent the carrying amounts of investments in privately held entities that do not report NAV for which impairments have been recognized in accordance with

the measurement alternative guidance described in “Investments” in Note 2 – “Summary of significant accounting policies” to our unaudited consolidated financial

statements.

Real estate assets classified as held for sale measured at fair value less costs to sell

Our real estate assets classified as held for sale and measured at fair value less costs to sell are presented in the table above.

These properties are subsets of our total real estate assets classified as held for sale as of March 31, 2025 and December 31, 2024,

respectively. The fair values for these real estate assets were estimated based on executed purchase and sale agreements, letters of

intent, or valuations provided by third-party real estate brokers. Refer to “Investments in real estate” in Note 2 – “Summary of significant

accounting policies” and “Assets held for sale” in Note 3 – “Investments in real estate” to our unaudited consolidated financial

statements for additional information.

Investments in privately held entities that do not report NAV

Our investments in privately held entities that do not report NAV are measured at cost, adjusted for observable price changes

and impairments, with changes recognized in net income (loss). These investments are adjusted based on the observable price

changes in orderly transactions for the identical or similar investment of the same issuer. Further adjustments are not made until

another observable transaction occurs. Therefore, the determination of fair values of our investments in privately held entities that do

not report NAV does not involve significant estimates and assumptions or subjective and complex judgments.

We also subject our investments in privately held entities that do not report NAV to a qualitative assessment for indicators of

impairment. If indicators of impairment are present, we are required to estimate the investment’s fair value and immediately recognize

an impairment charge in an amount equal to the investment’s carrying value in excess of its estimated fair value.

The estimates of fair value typically incorporate valuation techniques that include an income approach reflecting a discounted

cash flow analysis, and a market approach that includes a comparative analysis of acquisition multiples and pricing multiples generated

by market participants. In certain instances, we may use multiple valuation techniques for a particular investment and estimate its fair

value based on an average of multiple valuation results.

Refer to Note 7 – “Investments” to our unaudited consolidated financial statements for additional information.

35

9.FAIR VALUE MEASUREMENTS (continued)

Assets and liabilities not measured at fair value in the statement of financial position but for which the fair value is disclosed

The fair values of our secured notes payable and unsecured senior notes payable, and the amounts outstanding on our

unsecured senior line of credit and commercial paper program, were estimated using widely accepted valuation techniques, including

discounted cash flow analyses using significant other observable inputs such as available market information on discount and

borrowing rates with similar terms, maturities, and credit ratings. Because the valuations of our financial instruments are based on these

types of estimates, the actual fair value of our financial instruments may differ materially if our estimates do not prove to be accurate.

Additionally, the use of different market assumptions or estimation methods may have a material effect on the estimated fair value

amounts.

As of March 31, 2025 and December 31, 2024, the book and estimated fair values of our secured notes payable and

unsecured senior notes payable and the amounts outstanding under our unsecured senior line of credit and commercial paper program,

including the level within the fair value hierarchy for which the estimates were derived, were as follows (in thousands):

March 31, 2025
Book Value Fair Value Hierarchy Estimated<br><br>Fair Value
Quoted Prices in<br><br>Active Markets<br><br>for Identical Assets<br><br>(Level 1) Significant<br><br>Other<br><br>Observable Inputs<br><br>(Level 2) Significant<br><br>Unobservable<br><br>Inputs<br><br>(Level 3)
Liabilities:
Secured notes payable $150,807 $— $150,331 $— $150,331
Unsecured senior notes payable $12,640,144 $— $11,094,905 $— $11,094,905
Unsecured senior line of credit $— $— $— $— $—
Commercial paper program $299,883 $— $299,729 $— $299,729 December 31, 2024
--- --- --- --- --- ---
Book Value Fair Value Hierarchy Estimated<br><br>Fair Value
Quoted Prices in<br><br>Active Markets<br><br>for Identical Assets<br><br>(Level 1) Significant<br><br>Other<br><br>Observable Inputs<br><br>(Level 2) Significant<br><br>Unobservable<br><br>Inputs<br><br>(Level 3)
Liabilities:
Secured notes payable $149,909 $— $149,413 $— $149,413
Unsecured senior notes payable $12,094,465 $— $10,472,993 $— $10,472,993
Unsecured senior line of credit $— $— $— $— $—
Commercial paper program $— $— $— $— $—

The carrying values of cash and cash equivalents, restricted cash, tenant receivables, deposits, notes receivable, accounts

payable, accrued expenses, and other short-term liabilities approximate their fair value.

36

10.SECURED AND UNSECURED SENIOR DEBT

The following table summarizes our outstanding indebtedness and respective principal payments remaining as of March 31, 2025 (dollars in thousands):

Stated<br><br>Rate Interest<br><br>Rate(1) Maturity<br><br>Date(2) Principal Payments Remaining for the Periods Ending December 31, Unamortized<br><br>(Deferred<br><br>Financing<br><br>Cost),<br><br>(Discount)/<br><br>Premium
Debt 2025 2026 2027 2028 2029 Thereafter Principal Total
Secured notes payable
Greater Boston(3) SOFR+2.70% 7.20% 11/19/26 $— $150,418 $— $— $— $— $150,418 $(199) $150,219
San Francisco Bay Area 6.50% 6.50 7/1/36 34 36 38 41 44 395 588 588
Secured debt weighted-average interest rate/<br><br>subtotal 7.20 34 150,454 38 41 44 395 151,006 (199) 150,807
Unsecured senior line of credit and<br><br>commercial paper program(4) (4) 4.69 (4) 1/22/30 (4) 300,000 300,000 (117) 299,883
Unsecured senior notes payable 3.45% 3.62 4/30/25 (5) 600,000 600,000 (74) 599,926
Unsecured senior notes payable 4.30% 4.50 1/15/26 300,000 300,000 (408) 299,592
Unsecured senior notes payable 3.80% 3.96 4/15/26 350,000 350,000 (531) 349,469
Unsecured senior notes payable 3.95% 4.13 1/15/27 350,000 350,000 (940) 349,060
Unsecured senior notes payable 3.95% 4.07 1/15/28 425,000 425,000 (1,206) 423,794
Unsecured senior notes payable 4.50% 4.60 7/30/29 300,000 300,000 (971) 299,029
Unsecured senior notes payable 2.75% 2.87 12/15/29 400,000 400,000 (1,962) 398,038
Unsecured senior notes payable 4.70% 4.81 7/1/30 450,000 450,000 (1,964) 448,036
Unsecured senior notes payable 4.90% 5.05 12/15/30 700,000 700,000 (4,535) 695,465
Unsecured senior notes payable 3.375% 3.48 8/15/31 750,000 750,000 (4,188) 745,812
Unsecured senior notes payable 2.00% 2.12 5/18/32 900,000 900,000 (6,737) 893,263
Unsecured senior notes payable 1.875% 1.97 2/1/33 1,000,000 1,000,000 (6,892) 993,108
Unsecured senior notes payable 2.95% 3.07 3/15/34 800,000 800,000 (7,047) 792,953
Unsecured senior notes payable 4.75% 4.88 4/15/35 500,000 500,000 (4,844) 495,156
Unsecured senior notes payable 5.50% 5.66 10/1/35 550,000 550,000 (6,777) 543,223
Unsecured senior notes payable 5.25% 5.38 5/15/36 400,000 400,000 (4,024) 395,976
Unsecured senior notes payable 4.85% 4.93 4/15/49 300,000 300,000 (2,843) 297,157
Unsecured senior notes payable 4.00% 3.91 2/1/50 700,000 700,000 9,951 709,951
Unsecured senior notes payable 3.00% 3.08 5/18/51 850,000 850,000 (11,130) 838,870
Unsecured senior notes payable 3.55% 3.63 3/15/52 1,000,000 1,000,000 (13,561) 986,439
Unsecured senior notes payable 5.15% 5.26 4/15/53 500,000 500,000 (7,537) 492,463
Unsecured senior notes payable 5.625% 5.71 5/15/54 600,000 600,000 (6,636) 593,364
Unsecured debt weighted-average interest<br><br>rate/subtotal 3.91 600,000 650,000 350,000 425,000 700,000 10,300,000 13,025,000 (84,973) 12,940,027
Weighted-average interest rate/total 3.95% $600,034 $800,454 $350,038 $425,041 $700,044 $10,300,395 $13,176,006 $(85,172) $13,090,834

(1)Represents the weighted-average interest rate as of the end of the applicable period, including amortization of loan fees, amortization of debt premiums (discounts), and other bank fees.

(2)Reflects any extension options that we control.

(3)Represents a secured construction loan held by our consolidated real estate joint venture for 99 Coolidge Avenue, of which we own a 75.7% interest. As of March 31, 2025, this joint venture has $44.9 million available under existing lender

commitments. The interest rate shall be reduced from SOFR+2.70% to SOFR+2.10% over time upon the completion of certain leasing, construction, and financial covenant milestones.

(4)Refer to “$5.0 billion unsecured senior line of credit” and “$2.5 billion commercial paper program” on the following page.

(5)Upon maturity on April 30, 2025, we expect to repay $600.0 million of our 3.45% unsecured senior notes payable.

37

10.SECURED AND UNSECURED SENIOR DEBT (continued)

The following table summarizes our secured and unsecured senior debt and amounts outstanding under our unsecured senior

line of credit and commercial paper program as of March 31, 2025 (dollars in thousands):

Fixed-Rate<br><br>Debt Variable-Rate<br><br>Debt Weighted-Average
Interest Remaining<br><br>Term<br><br>(in years)
Total Percentage Rate(1)
Secured notes payable $588 $150,219 150,807 1.2% 7.20% 1.7
Unsecured senior notes payable 12,640,144 12,640,144 96.5 3.89 12.5
Unsecured senior line of credit<br><br>and commercial paper program 299,883 299,883 2.3 4.69 (2) 4.8 (3)
Total/weighted average $12,640,732 $450,102 13,090,834 100.0% 3.95% 12.2 (3)
Percentage of total debt 96.6% 3.4% 100%

All values are in US Dollars.

(1)Represents the weighted-average interest rate as of the end of the applicable period, including expense/income related to the amortization of loan fees, amortization of

debt premiums (discounts), and other bank fees.

(2)As of March 31, 2025, we had no outstanding balance on our unsecured senior line of credit and $299.9 million of commercial paper notes outstanding.

(3)We calculate the weighted-average remaining term of our commercial paper notes by using the maturity date of our unsecured senior line of credit. Using the maturity

date of our outstanding commercial paper notes, the consolidated weighted-average maturity of our debt is 12.0 years. The commercial paper notes sold during the

three months ended March 31, 2025 were issued at a weighted-average yield to maturity of 4.60% and had a weighted-average maturity term of 13 days.

Unsecured senior notes payable

In February 2025, we issued $550.0 million of unsecured senior notes payable, due 2035, with an interest rate of 5.50%.

$5.0 billion unsecured senior line of credit

As of March 31, 2025, our unsecured senior line of credit had aggregate commitments of $5.0 billion and bore an interest rate

of SOFR plus 0.855%. In addition to the cost of borrowing, the unsecured senior line of credit is subject to an annual facility fee of

0.145% based on the aggregate commitments outstanding. Based upon our ability to achieve certain annual sustainability metrics, the

interest rate and facility fee rate are also subject to upward or downward adjustments of up to four basis points with respect to the

interest rate and up to one basis point with respect to the facility fee rate.

Based on certain sustainability metrics achieved in accordance with the terms of our unsecured senior line of credit

agreement, the borrowing rate was reduced for a one-year period by two basis points to SOFR plus 0.855%, from SOFR plus 0.875%,

and the facility fee was reduced by 0.5 basis point to 0.145% from 0.15%. As of March 31, 2025, we had no outstanding balance on our

unsecured line of credit.

$2.5 billion commercial paper program

Our commercial paper program provides us with the ability to issue up to $2.5 billion of commercial paper notes that bear

interest at short-term fixed rates with a maturity of generally 30 days or less and a maximum maturity of 397 days from the date of

issuance. Our commercial paper program is backed by our unsecured senior line of credit, and at all times we expect to retain a

minimum undrawn amount of borrowing capacity under our unsecured senior line of credit equal to any outstanding notes issued under

our commercial paper program. We use the net proceeds from the issuances of the notes for general working capital and other general

corporate purposes. General corporate purposes may include, but are not limited to, the repayment of other debt and selective

development, redevelopment, or acquisition of properties. During the three months ended March 31, 2025, the commercial paper notes

were issued at a weighted-average yield to maturity of 4.60% and had a weighted-average maturity term of 13 days. As of March 31,

2025, we had a $299.9 million outstanding balance on our commercial paper program.

Interest expense

The following table summarizes interest expense for the three months ended March 31, 2025 and 2024 (in thousands):

Three Months Ended March 31,
2025 2024
Interest incurred $130,941 $122,680
Capitalized interest (80,065) (81,840)
Interest expense $50,876 $40,840

38

11. ACCOUNTS PAYABLE, ACCRUED EXPENSES, AND OTHER LIABILITIES

The following table summarizes the components of accounts payable, accrued expenses, and other liabilities as of March 31,

2025 and December 31, 2024 (in thousands):

March 31, 2025 December 31, 2024
Accounts payable and accrued expenses $382,956 $534,803
Accrued construction 400,317 500,890
Acquired below-market leases 164,020 180,407
Conditional asset retirement obligations 53,886 53,968
Deferred rent liabilities 11,297 11,461
Operating lease liability 371,412 507,127
Unearned rent and tenant security deposits 733,394 691,873
Other liabilities 164,132 173,822
Total $2,281,414 $2,654,351

As of March 31, 2025 and December 31, 2024, our conditional asset retirement obligations liability primarily consisted of the

soil and groundwater remediation liabilities associated with certain of our properties. Some of our properties may contain asbestos or

may be subjected to other hazardous or toxic substances, which, under certain conditions, requires remediation. We engage

independent environmental consultants to conduct Phase I or similar environmental assessments at our properties. This type of

assessment generally includes a site inspection, interviews, and a public records review; asbestos, lead-based paint, and mold surveys;

subsurface sampling; and other testing. We recognize a liability for the fair value of a conditional asset retirement obligation (including

asbestos) when the fair value of the liability can be reasonably estimated. In addition, environmental laws and regulations subject our

tenants, and potentially us, to liability that may result from our tenants’ routine handling of hazardous substances and wastes as part of

their operations at our properties. These assessments and investigations of our properties have not to date revealed any additional

environmental liability we believe would have a material adverse effect on our business and financial statements or that would require

additional disclosures or recognition in our consolidated financial statements.

39

12.EARNINGS PER SHARE

With respect to dividend rights, we have granted two types of restricted stock awards: (i) restricted stock awards with

nonforfeitable dividends and (ii) restricted stock awards with forfeitable dividends.

We account for unvested restricted stock awards (“RSAs”) with nonforfeitable dividends as participating securities and include

these securities in the computation of EPS using the two-class method. Under the two-class method, we allocate net income (after

amounts attributable to noncontrolling interests) to common stockholders and unvested RSAs with nonforfeitable dividends by using the

weighted-average shares of each class outstanding for quarter-to-date and year-to-date periods independently, based on their

respective participation rights to dividends declared (or accumulated) and undistributed earnings.

Unvested RSAs with forfeitable dividends do not qualify as participating securities under the two-class method because the

dividends are forfeited if the awards do not vest. As a result, undistributed earnings are not allocated to these awards prior to vesting,

and these awards have no effect on the computation of basic EPS while unvested. Once these awards vest, they are included in the

denominator of basic EPS, weighted for the portion of the reporting period they were vested. Prior to vesting, these awards are included

in the denominator of diluted EPS if they are dilutive, which is determined using the treasury stock method. Under this method,

incremental shares are calculated as the difference between the total unvested shares and the number of shares that could

hypothetically be repurchased using the assumed proceeds (including unrecognized compensation cost related to these awards).

These incremental shares are weighted for the portion of the reporting period they were unvested, and are included in the diluted EPS

denominator only if their inclusion reduces EPS (i.e., if they are not antidilutive).

In addition, from time to time, we enter into forward equity sales agreements. We consider the potential dilution resulting from

the forward equity sales agreements on the EPS calculations. At inception, the agreements do not have an effect on the computation of

basic EPS as no shares are delivered until settlement. The common shares issued upon the settlement of the forward equity sales

agreements, weighted for the period these common shares were outstanding, are included in the denominator of basic EPS. To

determine the dilution resulting from the forward equity sales agreements during the period of time prior to settlement, we calculate the

number of weighted-average shares outstanding – diluted using the treasury stock method. As of March 31, 2025, no forward equity

sales agreements were outstanding.

The table below reconciles the numerators and denominators of the basic and diluted EPS computations for the three months

ended March 31, 2025 and 2024 (in thousands, except per share amounts):

Three Months Ended March 31,
2025 2024
Net income $38,662 $219,176
Net income attributable to noncontrolling interests (47,601) (48,631)
Net income attributable to unvested RSAs with nonforfeitable dividends (2,660) (3,659)
Numerator for basic and diluted EPS – net (loss) income attributable to Alexandria Real Estate<br><br>Equities, Inc.’s common stockholders $(11,599) $166,886
Denominator for basic EPS – weighted-average shares of common stock outstanding 170,522 171,949
Dilutive effect of unvested RSAs with forfeitable dividends
Denominator for diluted EPS – weighted-average shares of common stock outstanding 170,522 171,949
Net (loss) income per share attributable to Alexandria Real Estate Equities, Inc.’s common<br><br>stockholders:
Basic $(0.07) $0.97
Diluted $(0.07) $0.97

40

13.STOCKHOLDERS’ EQUITY

Common equity transactions

Common stock repurchase program

Under our common stock repurchase program authorized in December 2024, we may repurchase up to $500.0 million of our

common stock in the open market, in privately negotiated transactions, or otherwise through December 31, 2025.

During the three months ended March 31, 2025, we repurchased 2.2 million shares of common stock under this repurchase

program at an average price per share of $96.71. As of March 31, 2025, the approximate value of shares that may yet be purchased

under this program was $241.8 million.

ATM common stock offering program

In February 2024, we entered into a new ATM common stock offering program that allows us to sell up to an aggregate of

$1.5 billion of our common stock. As of March 31, 2025, the remaining aggregate amount available under our ATM program for future

sales of common stock was $1.47 billion.

Dividends

During the three months ended March 31, 2025, we declared cash dividends on our common stock aggregating $228.3 million,

or $1.32 per share.

Accumulated other comprehensive loss

The change in accumulated other comprehensive loss attributable to Alexandria Real Estate Equities, Inc.’s stockholders

during the three months ended March 31, 2025 was entirely due to net unrealized gains of $50 thousand on foreign currency translation

related to our operations primarily in Canada.

Common stock, preferred stock, and excess stock authorizations

Our charter authorizes the issuance of 400.0 million shares of common stock, of which 170.1 million shares were issued and

outstanding as of March 31, 2025. Our charter also authorizes the issuance of up to 100.0 million shares of preferred stock, none of

which were issued and outstanding as of March 31, 2025. In addition, 200.0 million shares of “excess stock” (as defined in our charter)

are authorized, none of which were issued and outstanding as of March 31, 2025.

14.NONCONTROLLING INTERESTS

Noncontrolling interests represent the third-party interests in certain entities in which we have a controlling interest. As of

March 31, 2025, these entities owned 63 properties, which are included in our consolidated financial statements. Noncontrolling

interests are adjusted for additional contributions and distributions, the proportionate share of the net earnings or losses, and other

comprehensive income or loss. Distributions, profits, and losses related to these entities are allocated in accordance with the respective

operating agreements. During the three months ended March 31, 2025 and 2024, we distributed $66.0 million and $59.8 million,

respectively, to our consolidated real estate joint venture partners.

Certain of our noncontrolling interests have the right to require us to redeem their ownership interests in the respective entities.

We classify these ownership interests in the entities as redeemable noncontrolling interests outside of total equity in our consolidated

balance sheets. Redeemable noncontrolling interests are adjusted for additional contributions and distributions, the proportionate share

of the net earnings or losses, and other comprehensive income or loss. If the amount of a redeemable noncontrolling interest is less

than the maximum redemption value at the balance sheet date, such amount is adjusted to the maximum redemption value.

Subsequent declines in the redemption value are recognized only to the extent that previous increases have been recognized.

In March 2025, we redeemed our partner’s entire noncontrolling interests in three real estate joint ventures in our Greater

Boston market, with a book value aggregating $10.4 million, and recognized $7.0 million of consideration in excess of the book value in

additional paid-in capital.

Refer to Note 4 – “Consolidated and unconsolidated real estate joint ventures” to our unaudited consolidated financial

statements for additional information.

41

15.SEGMENT INFORMATION

We are a life science REIT focused on developing, redeveloping, and operating properties that provide space for lease to

tenants primarily in the life science industry. Our properties are leased predominantly through triple-net lease agreements and share

key characteristics, including generic and reusable improvements, consistent lease structures, and business strategy. All properties are

located within North America, predominantly in the U.S., and operate within a comparable regulatory environment.

Operating segments

Our Chief Operating Decision Maker (“CODM”), represented by our Executive Chairman and our Chief Executive Officer,

evaluates operating results at the geographic market level to assess performance and allocate resources. Our operating segments align

with our markets, including Greater Boston, San Francisco Bay Area, San Diego, and Seattle, among others. Regular market

performance updates are provided directly to the CODM. These updates include each market’s net operating income (“NOI”), which

serves as the profit or loss measure used by the CODM for performance assessment and resource allocation. NOI provides useful

information regarding performance of each market as it reflects income and expenses incurred in connection with real estate operations

in each market. This metric enables the CODM to evaluate the profitability and performance of each market on a consistent and

comparable basis, supporting decisions on capital resource allocation, including in connection with development, redevelopment,

acquisition, and disposition activities in each market.

Evaluation of economic similarity and aggregation of operating segments

In accordance with the segment reporting accounting standard, we evaluated the economic similarity of our operating

segments. Seven of our nine operating segments exhibit consistent long-term economic characteristics, including similar historical long-

term NOI margins, which are also expected to remain similar in the future. Additionally, these markets share similar operational

characteristics, including nature of services provided (i.e., leasing, operating, developing, and redeveloping life science properties),

tenant base (i.e., a variety of tenants involved in the life science industry), methods of operation (i.e., consistent lease structures,

property management practices, and business strategies), nature of the regulatory environment (consistent across North America,

where all our operating segments are located). Based on shared economic characteristics, we have aggregated our seven operating

segments into one reportable segment for segment reporting purposes. Two of our operating segments, specifically our New York City

and Canada markets, do not meet the aggregation criteria and individually do not meet the quantitative thresholds to qualify as

reportable segments. Therefore, these operating segments are included in the “all other” category in the tables below.

The following table presents the reportable segment profit or loss measure, net operating income, for the three months ended

March 31, 2025 and 2024:

Three Months Ended March 31,
(in thousands) 2025 2024
Reportable segment revenues:
Revenues from external customers $699,199 $720,565
Other income 6,527 5,676
Reportable segment total revenues 705,726 726,241
Reportable segment total rental operating expenses (212,436) (191,090)
Reportable segment net operating income (reportable segment profit or loss) $493,290 $535,151

Significant expenses included in the reportable segment profit or loss measure (i.e., net operating income) are represented by

the reportable segment total rental operating expenses and are disclosed in the table above. These expenses primarily include property

taxes, utilities, repairs and maintenance, engineering, janitorial, and other costs.

42

15.SEGMENT INFORMATION (continued)

Presented below are reconciliations of the reportable segment total revenues to the consolidated revenues, the reportable

segment total rental operating expenses to consolidated rental operations, the reportable segment NOI to the consolidated net income,

and the reportable segment investments in real estate assets to the consolidated investments in real estate assets:

Three Months Ended March 31,
(in thousands) 2025 2024
Reconciliation of reportable segment revenues to consolidated total revenues:
Reportable segment total revenues $705,726 $726,241
All other revenues 52,432 42,867
Consolidated revenues $758,158 $769,108
Reconciliation of reportable segment total rental operating expenses to consolidated rental<br><br>operations:
Reportable segment total rental operating expenses $(212,436) $(191,090)
All other rental operating expenses (13,959) (27,224)
Consolidated rental operations $(226,395) $(218,314)
Reconciliation of reportable segment net operating income to consolidated net income:
Reportable segment net operating income (reportable segment profit or loss) $493,290 $535,151
All other revenues 52,432 42,867
All other rental operating expenses (13,959) (27,224)
Other items not allocated to segments:
General and administrative (30,675) (47,055)
Interest expense (50,876) (40,840)
Depreciation and amortization (342,062) (287,554)
Impairment of real estate (32,154)
Equity in (losses) earnings of unconsolidated real estate joint ventures (507) 155
Investment (loss) income (49,992) 43,284
Gain on sale of real estate 13,165 392
Consolidated net income $38,662 $219,176 (in thousands) March 31, 2025 December 31, 2024
--- --- ---
Reconciliation of reportable segment assets to consolidated investments in real<br><br>estate assets
Reportable segment investments in real estate $30,405,384 $30,393,144
All other investments in real estate 1,716,328 1,716,895
Consolidated investments in real estate $32,121,712 $32,110,039

43

16.SUBSEQUENT EVENTS

In April 2025, an office property aggregating 182,276 RSF, located in Carlsbad, San Diego, met the criteria for classification as

held for sale based on current negotiations with the prospective buyer and our decision to dispose of this property for an estimated

sales price of approximately $72.0 million. We expect to complete the sale within 12 months. Upon our decision to commit to sell this

property, we recognized an impairment charge of $35.4 million to reduce the carrying amount of this asset to its estimated fair value

less costs to sell.

44

ITEM 2. MANAGEMENT’S DISCUSSION AND ANALYSIS OF FINANCIAL CONDITION AND RESULTS OF OPERATIONS

Forward-looking statements

Certain information and statements included in this quarterly report on Form 10-Q, including, without limitation, statements

containing the words “forecast,” “guidance,” “goals,” “projects,” “estimates,” “anticipates,” “believes,” “expects,” “intends,” “may,” “plans,”

“seeks,” “should,” “targets,” or “will,” or the negative of those words or similar words, constitute “forward-looking statements” within the

meaning of Section 27A of the Securities Act of 1933, as amended, and Section 21E of the Securities Exchange Act of 1934, as

amended. Forward-looking statements involve inherent risks and uncertainties regarding events, conditions, and financial trends that

may affect our future plans of operations, business strategy, results of operations, and financial position. A number of important factors

could cause actual results to differ materially from those included within or contemplated by the forward-looking statements, including,

but not limited to, the following:

•Operating factors, such as a failure to operate our business successfully in comparison to market expectations or in

comparison to our competitors, our inability to obtain capital when desired or refinance debt maturities when desired, and/

or a failure to maintain our status as a REIT for federal tax purposes;

•Market and industry factors, such as adverse developments concerning the life science industry and/or our tenants;

•Government factors, such as any unfavorable effects resulting from federal, state, local, and/or foreign government

policies, laws, and/or funding levels;

•Global factors, such as negative economic, social, political, financial, credit market, banking conditions, and/or regional

armed hostilities; and

•Other factors, such as climate change, cyber intrusions, and/or changes in laws, regulations, and financial accounting

standards.

Global Trade Policies

We have been monitoring and will continue to monitor macroeconomic trends and uncertainties. In particular, we are

assessing how recent fluctuations in international trade relations and trade policies could adversely affect our business or the

businesses of our tenants.

In early March 2025, the U.S. government imposed or indicated that it would impose a series of tariffs on certain goods from

Canada and Mexico as well as raise tariffs on Chinese imports. President Trump has also indicated his intent to impose a “major”

pharmaceutical-specific tariff, which could adversely affect our business and/or the business of our tenants. As a result of these

developments, the global securities and trade markets have reacted with volatility, and trade tensions remain high.

The imposition of tariffs or the potential future imposition of additional or modified tariffs in the current geopolitical climate could

have material adverse effects on the net profitability, revenues, or operations of Alexandria and many other companies. While we are

evaluating the potential impacts of such tariffs, as well as our ability to mitigate such impacts, these recent trends may in the meantime

interrupt supply chains, fragment international business relationships, and create unknown risks that would thereby affect our or our

tenants’ business operations.

This list of risks and uncertainties is not exhaustive. Additional information regarding risk factors that may affect us is included

under Part I; “Item 1A. Risk factors”; and “Item 7. Management’s discussion and analysis of financial condition and results of

operations” in our annual report on Form 10-K for the year ended December 31, 2024 and under respective sections in this quarterly

report on Form 10-Q. Readers of this quarterly report on Form 10-Q should also read our other documents filed publicly with the SEC

for further discussion regarding such factors.

45

Overview

We are a Maryland corporation formed in October 1994 that has elected to be taxed as a REIT for federal income tax

purposes. Alexandria Real Estate Equities, Inc. (NYSE: ARE), an S&P 500® company, is a best-in-class, mission-driven life science

REIT making a positive and lasting impact on the world. With our founding in 1994, Alexandria pioneered the life science real estate

niche. Alexandria is the preeminent and longest-tenured owner, operator, and developer of collaborative Megacampus™ ecosystems in

AAA life science innovation cluster locations, including Greater Boston, the San Francisco Bay Area, San Diego, Seattle, Maryland,

Research Triangle, and New York City. As of March 31, 2025, Alexandria has a total market capitalization of $28.8 billion and an asset

base in North America that includes 39.6 million RSF of operating properties and 4.0 million RSF of Class A/A+ properties undergoing

construction.

We develop dynamic Megacampus ecosystems that enable and inspire some of the world’s most brilliant minds and innovative

companies to create life-changing scientific and technological innovations. We believe in the utmost professionalism, humility, and

teamwork. Our tenants include multinational pharmaceutical companies; public and private biotechnology companies; life science

product, service, and medical device companies; digital health, technology, and agtech companies; academic and medical research

institutions; U.S. government research agencies; non-profit organizations; and venture capital firms. Alexandria has a longstanding and

proven track record of developing Class A/A+ properties clustered in highly dynamic and collaborative Megacampus environments that

enhance our tenants’ ability to successfully recruit and retain world-class talent and inspire productivity, efficiency, creativity, and

success. Alexandria also provides strategic capital to transformative life science companies through our venture capital platform. We

believe our unique business model and diligent underwriting ensure a high-quality and diverse tenant base that results in higher

occupancy levels, longer lease terms, higher rental income, higher returns, and greater long-term asset value.

As of March 31, 2025:

•Investment-grade or publicly traded large cap tenants represented 51% of our annual rental revenue;

•Approximately 98% of our leases (on an annual rental revenue basis) contained effective annual rent escalations

approximating 3% that were either fixed or indexed based on a consumer price index or other index;

•Approximately 91% of our leases (on an annual rental revenue basis) were triple net leases, which require tenants to pay

substantially all real estate taxes, insurance, utilities, repairs and maintenance, common area expenses, and other

operating expenses (including increases thereto) in addition to base rent;

•Approximately 93% of our leases (on an annual rental revenue basis) provided for the recapture of capital expenditures

(such as HVAC maintenance and/or replacement, roof replacement, and parking lot resurfacing) that we believe would

typically be borne by the landlord in traditional office leases; and

•89% of our leasing activity during the three months ended March 31, 2025 was generated from our existing tenant base.

A key element of our business strategy is our unique focus on Class A/A+ properties primarily located in collaborative

Megacampus ecosystems in AAA life science innovation clusters. Our Megacampus ecosystems are designed for optionality and

scalability, offering our tenants a clear path to address their growth requirements, including through our future developments and

redevelopments. Strategically located near top academic and medical research institutions and equipped with curated amenities and

services, and convenient access to transit, our Megacampus ecosystems are designed to support our tenants in attracting and retaining

top talent and in meeting our tenants’ growth needs, which we believe is a key driver of tenant demand for our properties. Our strategy

also includes drawing upon our deep, broad, and longstanding real estate and life science industry relationships in order to retain

tenants, identify and attract new and leading tenants, and source additional real estate.

Executive summary

Operating results

Three Months Ended March 31,
2025 2024
Net (loss) income attributable to Alexandria’s common stockholders – diluted:
In millions $(11.6) $166.9
Per share $(0.07) $0.97
Funds from operations attributable to Alexandria’s common stockholders – diluted, as adjusted:
In millions $392.0 $403.9
Per share $2.30 $2.35

For additional information, refer to “Funds from operations and funds from operations, as adjusted, attributable to Alexandria

Real Estate Equities, Inc.’s common stockholders” under “Definitions and reconciliations” and to the tabular presentation of these items

in “Results of operations” in Item 2.

46

A sector-leading REIT with a high-quality, diverse tenant base and strong margins

(As of March 31, 2025, unless stated otherwise)
Occupancy of operating properties in North America 91.7% (1)
Percentage of total annual rental revenue in effect from Megacampus platform 75%
Percentage of total annual rental revenue in effect from investment-grade or publicly traded large cap tenants 51%
Adjusted EBITDA margin for the three months ended March 31, 2025 71%
Percentage of leases containing annual rent escalations 98%
Weighted-average remaining lease term:
Top 20 tenants 9.6 years
All tenants 7.6 years
Sustained strength in tenant collections:
April 2025 tenant rents and receivables collected as of the date of this report 99.8%
Tenant rents and receivables for the three months ended March 31, 2025 collected as of the date of this<br><br>report 99.9%

(1)Refer to “Summary of occupancy percentages in North America” in Item 2 for additional details.

Strong and flexible balance sheet with significant liquidity; top 10% credit rating ranking among all publicly traded U.S. REITs

As of March 31, 2025, unless stated otherwise:

•Net debt and preferred stock to Adjusted EBITDA of 5.9x and fixed-charge coverage ratio of 4.3x for the three months ended

March 31, 2025 annualized, with targets for the three months ended December 31, 2025 annualized of less than or equal to

5.2x and 4.0x to 4.5x, respectively.

•Significant liquidity of $5.3 billion.

•Only 13% of our total debt matures through 2027.

•12.2 years weighted-average remaining term of debt, longest among S&P 500 REITs.

•Since 2021, an average of 97.9% of our year-end debt balances have been fixed rate.

•Total debt and preferred stock to gross assets of 30%.

•$414.9 million of capital contribution commitments from existing consolidated real estate joint venture partners to fund

construction from April 1, 2025 through 2027 and beyond, including $166.8 million from April 1, 2025 to December 31, 2025.

Continued solid leasing volume and rental rate increases

•Continued solid leasing volume aggregating 1.0 million RSF during the three months ended March 31, 2025, the fifth

consecutive quarter with leasing volume exceeding 1 million RSF.

•Solid rental rate increases on lease renewals and re-leasing of space of 18.5% and 7.5% (cash basis) for the three months

ended March 31, 2025.

•89% of our leasing activity during the three months ended March 31, 2025 was generated from our existing tenant base.

Three Months Ended<br><br>March 31, 2025
Total leasing activity – RSF 1,030,553
Lease renewals and re-leasing of space:
RSF (included in total leasing activity above) 884,408
Rental rate increase 18.5%
Rental rate increase (cash basis) 7.5%
Leasing of development and redevelopment space – RSF 6,430 (1)

(1)As of March 31, 2025, our construction projects expected to stabilize in 2025 and 2026 were 75% leased/negotiating.

47

Maintained solid operating metrics

•Total revenues of $758.2 million, down 1.4%, for the three months ended March 31, 2025, compared to $769.1 million for the

three months ended March 31, 2024. Excluding dispositions completed after January 1, 2024, total revenues would have

increased by 3.9% for the three months ended March 31, 2025, from $730.0 million for the three months ended March 31,

2024.

•Net operating income (cash basis) of $2.0 billion for the three months ended March 31, 2025 annualized increased by

$83.8 million, or 4.4%, compared to the three months ended March 31, 2024 annualized. Refer to “Net operating income, net

operating income (cash basis), and operating margin” under “Definitions and reconciliations” in Item 2 for a reconciliation of our

net income to net operating income (cash basis).

•Same property net operating income changes of (3.1)% and 5.1% (cash basis) for the three months ended March 31, 2025,

compared to the three months ended March 31, 2024 includes certain lease expirations during the three months ended March

31, 2025, aggregating 768,080 RSF at six properties across four submarkets. Excluding the impact of these lease expirations,

same property net operating income changes for the three months ended March 31, 2025 would have been 0.1% and 9.0%

(cash basis). Refer to the “Summary of occupancy percentages in North America” in Item 2 for additional details.

•General and administrative expenses of $30.7 million, savings of $16.4 million or 35%, for the three months ended March 31,

2025, compared to three months ended March 31, 2024, is primarily the result of cost-control and efficiency initiatives on

personnel-related costs and streamlining of business processes.

•As a percentage of net operating income, our general and administrative expenses for the trailing twelve months ended

March 31, 2025 were 6.9%, representing the lowest level in the past ten years, compared to 9.5% for the trailing twelve

months ended March 31, 2024.

Dividend strategy to share net cash flows from operating activities with stockholders while retaining a significant portion for reinvestment

•Common stock dividend declared for the three months ended March 31, 2025 of $1.32 per share aggregating $5.24 per

common share for the twelve months ended March 31, 2025, up 22 cents, or 4%, over the twelve months ended March 31,

2024.

•Dividend yield of 5.7% as of March 31, 2025.

•Dividend payout ratio of 57% for the three months ended March 31, 2025.

•Average annual dividend per-share growth of 4.5% from 2021 through the three months ended March 31, 2025 annualized.

•Significant net cash flows provided by operating activities after dividends retained for reinvestment aggregating $2.3 billion for

the years ended December 31, 2021 through 2024 and the midpoint of our 2025 guidance range for net cash provided by

operating activities after dividends.

Ongoing execution of Alexandria’s 2025 capital recycling strategy

We plan to continue funding a significant portion of our capital requirements for the year ending December 31, 2025 through

dispositions of non-core assets, land, partial interest sales, and sales to owner/users (in millions):

Completed dispositions 176
Our share of pending transactions subject to non-refundable deposits, signed letters of intent, and/or<br><br>purchase and sale agreement negotiations 433
Our share of completed and pending 2025 dispositions 609
Additional targeted dispositions 1,341
2025 guidance midpoint for dispositions and sales of partial interests 1,950

All values are in US Dollars.

Significant leasing progress on temporary vacancy for the three months ended March 31, 2025, including previously disclosed key

lease expirations during the three months ended March 31, 2025

Occupancy as of December 31, 2024 94.6%
Lease expirations which became vacant as of March 31, 2025:
Re-leased with future delivery or subject to ongoing negotiations (1.3) (1)
Marketing (1.6) (2.9) (2)
Occupancy as of March 31, 2025 91.7%

(1)Includes 0.7% of RSF that is re-leased with a weighted-average commencement date around the end of 2025 and 0.6% of RSF that is subject to ongoing negotiations.

(2)Includes 768,080 RSF of previously disclosed key lease expirations for the three months ended March 31, 2025. Refer to “Summary of properties and occupancy” in

Item 2 for additional details. The balance of lease expirations for the three months ended March 31, 2025 that became vacant was spread across multiple submarkets,

with no individual space aggregating greater than 62,000 RSF.

48

Strong and flexible balance sheet

Key capital metrics as of or for the three months ended March 31, 2025

•$28.8 billion in total market capitalization.

•$15.7 billion in total equity capitalization.

•Non-real estate investments aggregating $1.5 billion:

•Unrealized gains presented in our consolidated balance sheet were $31.9 million, comprising gross unrealized gains and

losses aggregating $204.9 million and $173.1 million, respectively.

•Investment loss of $50.0 million for the three months ended March 31, 2025 presented in our consolidated statement of

operations consisted of $29.3 million of realized gains, $68.1 million of unrealized losses, and $11.2 million of impairment

charges.

Key capital events

•In February 2025, we issued $550.0 million of unsecured senior notes payable, due in 2035, with an interest rate of 5.50%.

This issuance marked our tightest-ever spread to the 10-year treasury rate, surpassing our previous record in September 2019

by 25 bps.

•Upon maturity on April 30, 2025, we expect to repay $600.0 million of our 3.45% unsecured senior notes payable.

•During the three months ended March 31, 2025, our unconsolidated real estate joint venture at 1655 and 1725 Third Street, in

which we own a 10% interest, located in our Mission Bay submarket, refinanced $500 million of an existing fixed-rate secured

note payable with a new secured note payable, which bears a fixed weighted-average interest rate of 6.37% and matures in

2035.

•Under our common stock repurchase program authorized in December 2024, we may repurchase up to $500.0 million of our

common stock through December 31, 2025.

•During the three months ended March 31, 2025, we repurchased 2.2 million shares of common stock for an aggregate

value of $208.1 million at an average price per share of $96.71.

•As of the date of this report, the approximate value of shares authorized and remaining under this program was

$241.8 million

External growth and investments in real estate

Alexandria’s development and redevelopment pipeline delivered incremental annual net operating income of $37 million, commencing

during the three months ended March 31, 2025, with an additional $171 million of incremental annual net operating income anticipated

to deliver by the fourth quarter of 2026

•During the three months ended March 31, 2025, we placed into service development and redevelopment projects aggregating

309,494 RSF that are 100% leased across multiple submarkets and delivered incremental annual net operating income of

$37 million. A significant delivery during the three months ended March 31, 2025 was 285,346 RSF at 230 Harriet Tubman

Way located at the Alexandria Center® for Life Science – Millbrae in our South San Francisco submarket.

•Our active development and redevelopment projects under construction, primarily related to our Megacampus ecosystems,

have an estimated $2.4 billion of remaining costs to complete, of which $1.3 billion is not under contract as of March 31, 2025.

Additionally, we estimate that 30%–40% of the costs not under contract represent costs of materials that may be subject to

inflationary pressure and/or potential tariffs. As such, we estimate that each 10% increase in these costs of materials may

result in incremental costs aggregating $40 million–$50 million and a corresponding decline in initial stabilized yields of

approximately 2.5 to 3.5 basis points for our existing active development and redevelopment projects. This estimate does not

account for the cost of potential delays that may occur in receiving or replacing materials subject to tariffs.

•Annual net operating income (cash basis) from recently delivered projects is expected to increase by $61 million by the fourth

quarter of 2025 upon the burn-off of initial free rent, which have a weighted-average burn-off period of approximately four

months.

•71% of RSF in our total development and redevelopment pipeline is within our Megacampus ecosystems.

(dollars in millions) IncrementalAnnual Net Operating Income RSF Leased/<br><br>Negotiating<br><br>Percentage
Placed into service during the three months ended March 31, 2025 37 309,494 100%
Expected to be placed into service:
Second quarter of 2025 through fourth quarter of 2026 171 1,597,920 (2) 75% (3)
2027 through second quarter of 2028 179 2,449,862 16

All values are in US Dollars.

(1)Includes expected partial deliveries through the fourth quarter of 2026 from projects expected to stabilize in 2027 and beyond. Refer to the initial and stabilized

occupancy years under “New Class A/A+ development and redevelopment properties: current projects” in Item 2 for additional information.

(2)Represents the RSF related to projects expected to stabilize by fourth quarter of 2026. Does not include partial deliveries through fourth quarter of 2026 from

projects expected to stabilize in 2027 and beyond.

(3)Represents the leased/negotiating percentage of development and redevelopment projects that are expected to stabilize during 2025 and 2026.

49

Operating summary

Same Property Net<br><br>Operating Income Performance Rental Rate Growth:<br><br>Renewed/Re-Leased Space
Margins(2) Favorable Lease Structure(3)
Operating Adjusted EBITDA Strategic Lease Structure by Owner and<br><br>Operator of Collaborative Megacampus Ecosystems
70% 71% Increasing cash flows
Percentage of leases containing annual<br><br>rent escalations 98%
Stable cash flows
Historical Weighted-Average<br><br>Lease Term of Executed Leases(4) Percentage of triple<br><br>net leases 91%
Lower capex burden
8.9 Years Percentage of leases providing for the<br><br>recapture of capital expenditures 93%
Net Debt and Preferred Stock<br><br>to Adjusted EBITDA(5) Fixed-Charge Coverage Ratio(5)

1

13

25

37

49

61

4.0x to 4.5x

(1)

(3.1)%

2024 3/31/25

Refer to “Same properties” and “Definitions and reconciliations” in Item 2 for additional details. “Definitions and reconciliations” contains the definitions of “Fixed-charge

coverage ratio,” “Net debt and preferred stock to Adjusted EBITDA,” and “Net operating income” and their respective reconciliations from the most directly comparable

financial measures presented in accordance with GAAP.

(1)Refer to footnote 1 in “Same properties” in Item 2 for additional details.

(2)For the three months ended March 31, 2025.

(3)Percentages calculated based on our annual rental revenue in effect as of March 31, 2025.

(4)Represents the weighted-average lease term of executed leases based on annual rental revenue for the approximate 10-year period for the years ended December 31,

2016 through 2024 and the three months ended March 31, 2025.

(5)Quarter annualized.

50

Stable Cash Flows From Our High-Quality and Diverse Mix of<br><br>Approximately 750 Tenants
Investment-Grade or Publicly Traded<br><br>Large Cap Tenants
87%
of ARE’s Top 20 Tenant<br><br>Annual Rental Revenue
51%
of ARE’s Annual<br><br>Rental Revenue
Percentage of ARE’s<br><br>Annual Rental Revenue

25

Life Science

Product,

Service, and

Device

Multinational

Pharmaceutical

Public

Biotechnology –

Approved or

Marketed

Product

Public

Biotechnology –

Preclinical or

Clinical Stage

Private

Biotechnology

Other(2)

Other Investment-Grade

or Large Cap Tech

Biomedical

Institutions(1)

Government

Institutions

As of March 31, 2025. Annual rental revenue represents amounts in effect as of March 31, 2025. Refer to “Definitions and reconciliations” in Item 2 for additional information.

(1)79% of our annual rental revenue from biomedical institutions are from investment-grade or publicly traded large cap tenants.

(2)Represents the percentage of our annual rental revenue generated by technology, professional services, finance, telecommunications, construction/real estate

companies, and retail-related tenants.

51

Long-Duration and Stable Cash Flows From<br><br>High-Quality and Diverse Tenants
Long-Duration Lease Terms
9.6 Years
Top 20 Tenants
7.6 Years
All Tenants
Weighted-Average Remaining Term(1) Sustained Strength in Tenant Collections(2)
---
99.9%
For the Three Months Ended<br><br>March 31, 2025
99.8%
April 2025

(1)Based on annual rental revenue in effect as of March 31, 2025.

(2)Represents the portion of total receivables billed for each indicated period collected as of the date of this report.

52

Leasing Activity

The following table summarizes our leasing activity at our properties:

Three Months Ended Year Ended
March 31, 2025 December 31, 2024
(Dollars per RSF) Including Straight-Line Rent Cash Basis Including Straight-Line Rent
Leasing activity:
Renewed/re-leased space(1)
Rental rate changes 18.5% 7.5% 16.9%
New rates 57.56 $55.04 65.48
Expiring rates 48.57 $51.18 56.01
RSF 884,408 3,888,139
Tenant improvements/leasing commissions 83.09 46.89
Weighted-average lease term 10.1 years 8.5 years
Developed/redeveloped/previously vacant space leased(3)
New rates 49.80 $49.51 59.44
RSF 146,145 1,165,815
Weighted-average lease term 8.8 years 10.0 years
Leasing activity summary (totals):
New rates 56.46 $54.26 64.16
RSF 1,030,553 5,053,954
Weighted-average lease term 10.0 years 8.9 years
Lease expirations(1)
Expiring rates 49.93 $51.55 53.82
RSF 1,923,048 5,005,638

All values are in US Dollars.

Leasing activity includes 100% of results for properties in North America in which we have an investment.

(1)Excludes month-to-month leases aggregating 160,540 RSF and 136,131 RSF as of March 31, 2025 and December 31, 2024, respectively. During the trailing twelve

months ended March 31, 2025, we granted free rent concessions averaging 0.7 months per annum.

(2)Includes tenant improvements and leasing commissions for one 11.4-year lease at the Alexandria Technology Square® Megacampus in our Cambridge submarket

aggregating 119,280 RSF. Excluding this lease, tenant improvements and leasing commissions per RSF and as a percentage of total rents for the three months ended

March 31, 2025 were $40.93 and 9.1%, which are consistent with the five-year quarterly averages of $37.53 and 10.5%, respectively.

(3)Refer to “New Class A/A+ development and redevelopment properties: summary of pipeline” in Item 2 for additional information, including total project costs.

53

Summary of contractual lease expirations

The following table summarizes the contractual lease expirations at our properties as of March 31, 2025:

Year RSF Percentage of<br><br>Occupied RSF Annual Rental Revenue<br><br>(per RSF)(1)
2025 (2) 2,005,741 5.6% 46.91 4.6%
2026 3,043,760 8.5% 56.08 8.3%
2027 3,130,452 8.7% 51.23 7.8%
2028 4,060,412 11.3% 52.17 10.3%
2029 2,429,749 6.8% 50.67 6.0%
2030 3,064,307 8.6% 43.86 6.5%
2031 3,579,117 10.0% 54.84 9.5%
2032 1,023,407 2.9% 58.33 2.9%
2033 2,539,851 7.1% 48.14 5.9%
2034 3,280,121 9.2% 67.72 10.7%
Thereafter 7,673,811 21.3% 74.48 27.5%

All values are in US Dollars.

Contractual lease expirations for properties classified as held for sale as of March 31, 2025 are excluded from the information on this page.

(1)Represents amounts in effect as of March 31, 2025.

(2)Excludes month-to-month leases aggregating 160,540 RSF as of March 31, 2025.

The following tables present our lease expirations by market for the remainder of 2025 and for 2026 as of March 31, 2025:

2025 Contractual Lease Expirations (in RSF)
Market Leased Negotiating/<br><br>Anticipating Targeted for<br><br>Future<br><br>Development/<br><br>Redevelopment(1) Remaining<br><br>Expiring<br><br>Leases(2) Total(3) Annual Rental<br><br>Revenue<br><br>(per RSF)(4)
Greater Boston 136,506 5,597 25,312 261,540 428,955 $45.19
San Francisco Bay Area 293,051 110,549 346,927 750,527 71.21
San Diego 28,760 85,189 113,949 34.37
Seattle 67,114 67,114 31.33
Maryland 35,055 6,228 31,683 72,966 22.19
Research Triangle 173,888 78,625 252,513 27.98
New York City 42,002 42,002 99.58
Texas 198,972 (5) 198,972 N/A
Canada 22,991 54,752 77,743 18.35
Non-cluster/other markets 1,000 1,000 49.20
Total 690,251 122,374 224,284 968,832 2,005,741 $46.91
Percentage of expiring leases 34% 6% 11% 49% 100%
2026 Contractual Lease Expirations (in RSF) Annual Rental<br><br>Revenue<br><br>(per RSF)(4)
Market Leased Negotiating/<br><br>Anticipating Targeted for<br><br>Future<br><br>Development/<br><br>Redevelopment Remaining<br><br>Expiring<br><br>Leases(2) Total
Greater Boston 47,439 11,565 399,436 458,440 $94.58
San Francisco Bay Area 25,511 623,634 649,145 76.43
San Diego 28,827 873,855 902,682 47.04
Seattle 26,266 166,491 192,757 31.57
Maryland 15,489 276,969 292,458 20.20
Research Triangle 19,753 167,805 187,558 38.98
New York City 72,052 72,052 104.17
Texas
Canada 247,743 247,743 21.23
Non-cluster/other markets 40,925 40,925 75.98
Total 118,969 303,624 2,621,167 3,043,760 $56.08
Percentage of expiring leases 4% 10% 0% 86% 100%

Contractual lease expirations for properties classified as held for sale as of March 31, 2025 are excluded from the information on this page.

(1)Primarily represents assets that were recently acquired for future development or redevelopment opportunities, for which we expect, subject to market conditions and

leasing, to commence first-time conversion from non-laboratory space to laboratory space, or to commence future ground-up development. As of March 31, 2025, the

weighted-average annual rental revenue and expiration date of these leases expiring in 2025 is $1.6 million and May 27, 2025, respectively. Refer to “Investments in real

estate” under “Definitions and reconciliations” in Item 2 for additional details, including development and redevelopment square feet currently included in rental

properties.

(2)The largest remaining contractual lease expiration in 2025 is 88,179 RSF in our Cambridge/Inner Suburbs submarket and in 2026 is 163,648 RSF in our University Town

Center submarket, at a property in which we have an ownership interest of 30.0% and are evaluating options to re-lease or reposition the space from single tenancy to

multi-tenancy.

(3)Excludes month-to-month leases aggregating 160,540 RSF as of March 31, 2025.

(4)Represents amounts in effect as of March 31, 2025.

(5)Represents two properties with future development and redevelopment opportunities, located at 1001 Trinity Street and 1020 Red River Street in our Austin submarket,

with contractual lease expirations during the second quarter of 2025.

54

Top 20 tenants

87% of Top 20 Tenant Annual Rental Revenue Is From Investment-Grade

or Publicly Traded Large Cap Tenants(1)

Our properties are leased to a high-quality and diverse group of tenants, with no individual tenant accounting for greater than

4.3% of our annual rental revenue in effect as of March 31, 2025. The following table sets forth information regarding leases with our 20

largest tenants in North America based upon annual rental revenue in effect as of March 31, 2025 (dollars in thousands, except average

market cap amounts):

Remaining<br><br>Lease<br><br>Term(1)<br><br>(in Years) Aggregate<br><br>RSF Annual<br><br>Rental<br><br>Revenue(1) Percentage<br><br>of Annual<br><br>Rental<br><br>Revenue (1) Investment-Grade<br><br>Credit Ratings Average<br><br>Market<br><br>Cap<br><br>(in billions)
Tenant Moody’s S&P
1 Eli Lilly and Company 9.7 1,070,953 $ 89,599 4.3% Aa3 A+ $797.9
2 Moderna, Inc. 11.1 496,814 89,347 4.3 $29.1
3 Bristol-Myers Squibb Company 5.2 999,379 77,188 3.7 A2 A $104.1
4 Takeda Pharmaceutical Company Limited 10.2 549,759 47,899 2.3 Baa1 BBB+ $43.8
5 Eikon Therapeutics, Inc.(2) 13.7 311,806 36,783 1.8 $—
6 Roche 8.0 647,069 36,189 1.7 Aa2 AA $242.8
7 Illumina, Inc. 5.6 857,967 35,924 1.7 Baa3 BBB $19.5
8 Alphabet Inc. 2.6 625,015 34,899 1.7 Aa2 AA+ $2,143.6
9 2seventy bio, Inc.(3) 8.4 312,805 33,543 1.6 $0.2
10 United States Government 5.3 429,359 29,097 (4) 1.4 Aaa AA+ $—
11 Uber Technologies, Inc. 57.5 (5) 1,009,188 27,799 1.3 Baa2 BBB $148.3
12 Novartis AG 3.3 387,563 27,709 1.3 Aa3 AA- $234.5
13 AstraZeneca PLC 4.6 450,848 27,226 1.3 A1 A+ $231.1
14 Cloud Software Group, Inc. 1.2 (6) 292,013 26,446 1.3 $—
15 Boston Children’s Hospital 12.0 309,231 26,212 1.3 Aa2 AA $—
16 The Regents of the University of California 6.2 369,753 23,330 1.1 Aa2 AA $—
17 Sanofi 5.8 267,278 21,851 1.1 A1 AA $130.9
18 Charles River Laboratories, Inc. 10.1 256,066 21,202 1.0 $10.2
19 New York University 7.3 218,983 21,110 1.0 Aa2 AA- $—
20 Merck & Co., Inc. 8.4 333,124 21,001 1.0 Aa3 A+ $281.3
Total/weighted-average 9.6 (5) 10,194,973 $ 754,354 36.2%

Annual rental revenue and RSF include 100% of each property managed by us in North America. Refer to “Annual rental revenue” and “Investment-grade or publicly traded large

cap tenants” under “Definitions and reconciliations” in Item 2 for additional details, including our methodologies of calculating annual rental revenue from unconsolidated real

estate joint ventures and average market capitalization, respectively.

(1)Based on total annual rental revenue in effect as of March 31, 2025.

(2)Eikon Therapeutics, Inc. is a private biotechnology company led by renowned biopharma executive Roger Perlmutter, formerly an executive vice president at Merck & Co.,

Inc. As of February 25, 2025, the company has raised over $1.2 billion in private venture capital funding.

(3)In March 2025, 2seventy bio, Inc. announced a definitive merger agreement with Bristol-Myers Squibb Company, which is expected to close in the second quarter of 2025.

(4)Includes leases, which are not subject to annual appropriations, with governmental entities such as the National Institutes of Health and the General Services

Administration.  Approximately 3% of the annual rental revenue derived from our leases with the United States Government is cancellable prior to the lease expiration date.

(5)Includes (i) ground leases for land at 1455 and 1515 Third Street (two buildings aggregating 422,980 RSF) and (ii) leases at 1655 and 1725 Third Street (two buildings

aggregating 586,208 RSF) in our Mission Bay submarket owned by our unconsolidated real estate joint venture in which we have an ownership interest of 10%. Annual

rental revenue is presented using 100% of the annual rental revenue from our consolidated properties and our share of annual rental revenue from our unconsolidated real

estate joint ventures. Excluding these ground leases, the weighted-average remaining lease term for our top 20 tenants was 6.9 years as of March 31, 2025.

(6)Represents one lease at a property acquired in 2022 with potential future development and redevelopment opportunities. This lease with Cloud Software Group, Inc.

(formerly known as TIBCO Software, Inc.) was in place when we acquired the property.

55

Locations of properties

Our properties are strategically located in AAA life science innovation cluster markets. The following table sets forth the total

RSF, number of properties, and annual rental revenue in effect as of March 31, 2025 in each of our markets in North America (dollars in

thousands, except per RSF amounts):

RSF Number of<br><br>Properties Annual Rental Revenue
Market Operating Development Redevelopment Total % of Total Total % of Total Per RSF
Greater Boston 9,304,074 632,850 1,601,010 11,537,934 26% 65 $754,342 36% $88.20
San Francisco Bay Area 7,971,965 109,435 366,939 8,448,339 19 65 455,516 22 68.28
San Diego 7,140,194 903,792 8,043,986 18 77 323,222 16 47.98
Seattle 3,179,033 227,577 3,406,610 9 45 137,539 6 47.27
Maryland 3,848,870 3,848,870 9 50 141,895 7 39.70
Research Triangle 3,801,564 3,801,564 9 38 109,002 5 30.71
New York City 921,894 921,894 2 4 74,571 4 92.34
Texas 1,845,159 73,298 1,918,457 4 15 37,754 2 24.93
Canada 895,182 132,881 1,028,063 2 11 18,525 1 21.86
Non-cluster/other markets 349,099 349,099 1 10 15,413 1 60.52
Properties held for sale 382,527 382,527 1 6 9,031 49.82
North America 39,639,561 1,873,654 2,174,128 43,687,343 100% 386 $2,076,810 100% $58.38
4,047,782

Summary of occupancy percentages in North America

The following table sets forth the occupancy percentages for our operating properties and our operating and redevelopment

properties in each of our North America markets, excluding properties held for sale, as of the following dates:

Operating Properties Operating and Redevelopment Properties
Market 3/31/25 12/31/24 3/31/24 3/31/25 12/31/24 3/31/24
Greater Boston 91.8% (1) 94.8% 94.5% 78.4% 80.8% 83.3%
San Francisco Bay Area 90.3 (1) 93.3 94.4 86.3 89.1 91.2
San Diego 94.3 96.3 95.2 94.3 96.3 95.2
Seattle 91.5 92.4 94.9 91.5 92.4 93.9
Maryland 94.1 95.7 95.4 94.1 95.7 95.4
Research Triangle 93.4 (1) 97.4 97.8 93.4 97.4 97.8
New York City 87.6 (2) 88.4 84.4 87.6 88.4 84.4
Texas 82.1 (1) 95.5 95.1 78.9 91.8 91.5
Subtotal 91.8 94.8 94.9 87.1 90.0 90.6
Canada 94.6 95.9 91.8 82.4 82.9 77.8
Non-cluster/other markets 73.0 72.5 75.4 73.0 72.5 75.4
North America 91.7% (1)(3) 94.6% 94.6% 86.9% 89.7% 90.2%

(1)The decline in occupancy from December 31, 2024 includes certain previously disclosed lease expirations during the three months ended March 31, 2025 aggregating

768,080 RSF at six properties in four submarkets comprising the following: (i) 182,054 RSF at the Alexandria Technology Square® Megacampus in our Cambridge

submarket, (ii) 234,249 RSF at 409 Illinois Street in our Mission Bay submarket, (iii) one property aggregating 104,531 RSF in our Research Triangle market, and

(iv) two properties aggregating 247,246 RSF in our Austin submarket.

(2)The Alexandria Center® for Life Science – New York City Megacampus is 97.7% occupied as of March 31, 2025. Occupancy percentage in our New York City market

reflects vacancy at the Alexandria Center® for Life Science – Long Island City property, which was 45.7% occupied as of March 31, 2025.

(3)Includes vacant spaces aggregating 250,925 RSF, or 0.7% impact to occupancy, which are leased with a weighted-average expected delivery date around the end of

2025 and 242,035 RSF, or 0.6% impact to occupancy, which is subject to ongoing negotiations.

56

Investments in real estate

A key component of our business model is our disciplined allocation of capital to the development and redevelopment of new

Class A/A+ properties, and property enhancements identified during the underwriting of certain acquired properties, primarily located in

collaborative Megacampus ecosystems in AAA life science innovation clusters. These projects are focused on providing high-quality,

generic, and reusable spaces that meet the real estate requirements of a wide range of tenants. Upon completion, each development or

redevelopment project is expected to generate increases in rental income, net operating income, and cash flows. Our development and

redevelopment projects are generally in locations that are highly desirable to high-quality entities, which we believe results in higher

occupancy levels, longer lease terms, higher rental income, higher returns, and greater long-term asset value. Our pre-construction

activities are undertaken in order to prepare the property for its intended use and include entitlements, permitting, design, site work, and

other activities preceding commencement of construction of aboveground building improvements.

Our investments in real estate consisted of the following as of March 31, 2025 (dollars in thousands):

Development and Redevelopment
Under Construction
Operating 2025 and<br><br>2026 2027 and<br><br>Beyond Future Subtotal Total
Square footage
Operating 39,257,034 39,257,034
Future Class A/A+ development and redevelopment<br><br>properties 1,597,920 2,449,862 25,757,349 29,805,131 29,805,131
Future development and redevelopment square feet<br><br>currently included in rental properties(1) (2,780,364) (2,780,364) (2,780,364)
Total square footage, excluding properties held for sale 39,257,034 1,597,920 2,449,862 22,976,985 27,024,767 66,281,801
Properties held for sale 382,527 1,853,856 1,853,856 2,236,383
Total square footage 39,639,561 1,597,920 2,449,862 24,830,841 28,878,623 68,518,184
Investments in real estate
Gross book value as of March 31, 2025(2) $29,411,505 $1,549,293 $2,139,008 $4,908,467 $8,596,768 $38,008,273

(1)Refer to “Investments in real estate” under “Definitions and reconciliations” in Item 2 for additional details, including future development and redevelopment square feet

currently included in rental properties.

(2)Balances exclude accumulated depreciation and our share of the cost basis associated with our properties held by our unconsolidated real estate joint ventures, which is

classified as investments in unconsolidated real estate joint ventures in our consolidated balance sheet.

57

Dispositions and sales of partial interests

Our completed dispositions and sales of partial interests of real estate assets during the three months ended March 31, 2025 and pending as of the date of this report consisted of

the following (dollars in thousands):

Property Submarket/Market Date of<br><br>Sale Interest<br><br>Sold Future Development RSF Gain on<br><br>Sales of<br><br>Real Estate
Completed during the three months ended March 31, 2025:
Land and other
Costa Verde by Alexandria University Town Center/San Diego 1/31/25 100% 537,000 $—
Other 13,165
$13,165
Our share of pending 2025 dispositions and sales of partial interests expected to<br><br>close subsequent to April 28, 2025:
Subject to non-refundable deposits:
Pending San Diego 2H25 100%
Pending Texas 2Q25 100%
Other
Subject to executed letters of intent and/or purchase and sale agreement negotiations
Our share of completed and pending 2025 dispositions and sales of partial interests
2025 guidance range for dispositions and sales of partial interests 1,450,000 – 2,450,000

All values are in US Dollars.

(1)As part of a completed transaction, we provided seller financing of $91.0 million, due 2028, with an interest rate of 12.0%.

58

New Class A/A+ development and redevelopment properties

pipelinepagev2.jpg

ALEXANDRIA’S DEVELOPMENT AND REDEVELOPMENT

DELIVERIES ARE EXPECTED TO PROVIDE INCREMENTAL

GROWTH IN ANNUAL NET OPERATING INCOME

Placed Into<br><br>Service Near-Term<br><br>Deliveries Intermediate-Term<br><br>Deliveries
1Q25 2Q25–4Q26 2027–2Q28
$37M $171M $179M
309,494 RSF 1.6 million RSF 2.4 million RSF
100%<br><br>Leased 75%<br><br>Leased/Negotiating 16%<br><br>Leased/Negotiating

(1)

(1)

(2)

(4)

(3)

For the definition of “Net operating income” and a reconciliation from the most directly comparable GAAP measure, refer to the “Definitions and reconciliations” in Item 2.

(1)Our share of incremental annual net operating income from development and redevelopment projects expected to be placed into service primarily commencing from second quarter of 2025 through second quarter of 2028 is projected to be

$311 million.

(2)Includes expected partial deliveries through fourth quarter of 2026 from projects expected to stabilize in 2027 and beyond. Refer to the initial and stabilized occupancy years under “New Class A/A+ development and redevelopment

properties: current projects” in Item 2 for additional details.

(3)Represents the RSF related to projects expected to stabilize by fourth quarter of 2026. Does not include partial deliveries through fourth quarter of 2026 from projects expected to stabilize in 2027 and beyond.

(4)Represents the leased/negotiating percentage of development and redevelopment projects that are expected to stabilize during 2025 and 2026.

59

New Class A/A+ development and redevelopment properties: recent deliveries

The following table presents development and redevelopment of new Class A/A+ projects placed into service during the three months ended March 31, 2025 (dollars in thousands):

Incremental Annual Net Operating Income Generated From 1Q25 Deliveries Aggregated $37 Million

230 Harriet Tubman Way 10075 Barnes Canyon Road
San Francisco Bay Area/<br><br>South San Francisco San Diego/Sorrento Mesa
285,346 RSF 17,718 RSF
100% Occupancy 100% Occupancy
Property/Market/Submarket 1Q25<br><br>Delivery<br><br>Date(1) Our<br><br>Ownership<br><br>Interest RSF Placed in Service Occupancy<br><br>Percentage(2) Total Project
--- --- --- --- --- --- --- --- --- --- --- --- --- ---
Prior to<br><br>1/1/25 1Q25 Total Initial<br><br>Stabilized
RSF Investment
Development projects
230 Harriet Tubman Way/San Francisco Bay Area/South San Francisco 3/1/25 48.3% 285,346 285,346 100% 285,346 476,000 7.5% 6.2%
10075 Barnes Canyon Road/San Diego/Sorrento Mesa 2/6/25 50.0% 17,718 17,718 100% 253,079 321,000 5.5 5.7
Redevelopment projects
Canada 3/27/25 100% 78,487 6,430 84,917 100% 250,790 115,000 6.0 6.0
Weighted average/total 2/28/25 78,487 309,494 387,981 789,215 912,000 6.6% 6.0%

All values are in US Dollars.

(1)Represents the average delivery date for deliveries that occurred during the three months ended March 31, 2025, weighted by annual rental revenue.

(2)Occupancy relates to total operating RSF placed in service as of the most recent delivery.

60

New Class A/A+ development and redevelopment properties: 2025 and 2026 stabilization

99 Coolidge Avenue 500 North Beacon Street and<br><br>4 Kingsbury Avenue(1) 401 Park Drive 1450 Owens Street
Greater Boston/<br><br>Cambridge/Inner Suburbs Greater Boston/<br><br>Cambridge/Inner Suburbs Greater Boston/Fenway San Francisco Bay Area/<br><br>Mission Bay
204,395 RSF 36,444 RSF 137,675 RSF 109,435 RSF(2)
76% Leased/Negotiating 92% Leased/Negotiating Marketing Marketing 10935, 10945, and 10955<br><br>Alexandria Way(3) 4135 Campus Point Court 10075 Barnes Canyon Road 8800 Technology Forest Place
--- --- --- ---
San Diego/Torrey Pines San Diego/<br><br>University Town Center San Diego/Sorrento Mesa Texas/Greater Houston
241,504 RSF 426,927 RSF 235,361 RSF 73,298 RSF
100% Leased 100% Leased 68% Leased/Negotiating 41% Leased/Negotiating

(1)Image represents 500 North Beacon Street on The Arsenal on the Charles Megacampus.

(2)Image represents a multi-tenant project expanding the Alexandria Center® for Science and Technology – Mission Bay Megacampus, where we have a 25% interest. During the three months ended December 31, 2024, we

executed a letter of intent with a biomedical institution for the sale of a condominium interest aggregating 103,361 RSF, or approximately 49% of the development project. We expect to complete the transaction in the second half

of 2025. Accordingly, we adjusted the development project RSF and its related book value to reflect 109,435 RSF.

(3)Image represents 10955 Alexandria Way on the One Alexandria Square Megacampus.

61

New Class A/A+ development and redevelopment properties: 2027 and beyond stabilization

311 Arsenal Street 421 Park Drive 40, 50, and 60 Sylvan Road(1)
Greater Boston/<br><br>Cambridge/Inner Suburbs Greater Boston/Fenway Greater Boston/Route 128
308,446 RSF 392,011 RSF 596,064 RSF 651 Gateway Boulevard 269 East Grand Avenue 701 Dexter Avenue North
--- --- ---
San Francisco Bay Area/<br><br>South San Francisco San Francisco Bay Area/<br><br>South San Francisco Seattle/Lake Union
259,689 RSF 107,250 RSF 227,577 RSF

(1)Image represents 60 Sylvan Road on the Alexandria Center® for Life Science – Waltham Megacampus. The project is expected to capture demand in our Route 128 submarket.

62

New Class A/A+ development and redevelopment properties: current projects

The following tables set forth a summary of our new Class A/A+ development and redevelopment properties under construction as of March 31, 2025 (dollars in thousands):

Property/Market/Submarket Square Footage Percentage Occupancy(1)
In Service CIP Total Leased Leased/<br><br>Negotiating Initial Stabilized
Under construction
2025 and 2026 stabilization
99 Coolidge Avenue/Greater Boston/Cambridge/Inner Suburbs 116,414 204,395 320,809 40% 76% 4Q23 2026
500 North Beacon Street and 4 Kingsbury Avenue/Greater Boston/Cambridge/Inner Suburbs 211,574 36,444 248,018 92 92 1Q24 2025
401 Park Drive/Greater Boston/Fenway 137,675 137,675 2026 2026
1450 Owens Street/San Francisco Bay Area/Mission Bay(2) 109,435 109,435 2026 2026
10935, 10945, and 10955 Alexandria Way/San Diego/Torrey Pines 93,492 241,504 334,996 100 100 4Q24 2026
4135 Campus Point Court/San Diego/University Town Center 426,927 426,927 100 100 2026 2026
10075 Barnes Canyon Road/San Diego/Sorrento Mesa 17,718 235,361 253,079 68 68 1Q25 2026
8800 Technology Forest Place/Texas/Greater Houston 50,094 73,298 123,392 41 41 2Q23 2026
Canada 117,909 132,881 250,790 78 80 3Q23 2025
607,201 1,597,920 2,205,121 70 75
2027 and beyond stabilization
One Hampshire Street/Greater Boston/Cambridge 104,956 104,956 2027 2028
311 Arsenal Street/Greater Boston/Cambridge/Inner Suburbs 82,216 (3) 308,446 390,662 12 12 2027 2027
421 Park Drive/Greater Boston/Fenway 392,011 392,011 13 13 2026 2027
40, 50, and 60 Sylvan Road/Greater Boston/Route 128 596,064 596,064 31 31 2026 2027
Other/Greater Boston 453,869 453,869 (4) 2027 2027
651 Gateway Boulevard/San Francisco Bay Area/South San Francisco(5) 67,017 259,689 326,706 21 21 1Q24 2027
269 East Grand Avenue/San Francisco Bay Area/South San Francisco 107,250 107,250 2026 2027
701 Dexter Avenue North/Seattle/Lake Union 227,577 227,577 23 2026 2027
149,233 2,449,862 2,599,095 14 16
756,434 4,047,782 4,804,216 39% 43%
(1)Initial occupancy dates are subject to leasing and/or market conditions. Stabilized occupancy may vary depending on single tenancy versus multi-tenancy. Multi-tenant projects may increase in occupancy over a period of time.(2)Represents a multi-tenant project expanding the Alexandria Center® for Science and Technology – Mission Bay Megacampus, where we have a 25% interest. During the three months ended December 31, 2024, we executed a letter of intent with a biomedical institution for the sale of a condominium interest aggregating 103,361 RSF, or approximately 49% of the development project. We expect to complete the transaction in the second half of 2025. Accordingly, we adjusted the development project RSF and its related book value to reflect 109,435 RSF.(3)We expect to redevelop an additional 25,312 RSF of space occupied as of March 31, 2025 into laboratory space upon expiration of the existing leases during the second quarter of 2025. Refer to “Investments in real estate” under “Definitions and reconciliations” in Item 2 for additional information. (4)Represents a project focused on demand from our existing tenants in our adjacent properties/campuses that will address demand from other non-Alexandria properties/campuses.(5)We continue to build out this project on a floor-by-floor basis.  As of March 31, 2025, the remaining cost to complete is 138 million, or 28% of the total cost at completion.

All values are in US Dollars.

63

New Class A/A+ development and redevelopment properties: current projects (continued)

At 100% Unlevered Yields
Property/Market/Submarket In Service Initial Stabilized<br><br>(Cash Basis)
Under construction
2025 and 2026 stabilization(1)
99 Coolidge Avenue/Greater Boston/Cambridge/Inner Suburbs 136,658 203,904 103,438 444,000 6.0% 6.8%
500 North Beacon Street and 4 Kingsbury Avenue/Greater Boston/Cambridge/Inner Suburbs 378,211 41,649 7,140 427,000 6.2% 5.5%
401 Park Drive/Greater Boston/Fenway 167,606 TBD
1450 Owens Street/San Francisco Bay Area/Mission Bay 123,380
10935, 10945, and 10955 Alexandria Way/San Diego/Torrey Pines 105,766 367,114 30,120 503,000 6.2% 5.8%
4135 Campus Point Court/San Diego/University Town Center 369,624 154,376 524,000 6.6% 6.2%
10075 Barnes Canyon Road/San Diego/Sorrento Mesa 16,126 179,471 125,403 321,000 5.5% 5.7%
8800 Technology Forest Place/Texas/Greater Houston 60,225 46,300 5,475 112,000 6.3% 6.0%
Canada 55,503 50,245 9,252 115,000 6.0% 6.0%
752,489 1,549,293
2027 and beyond stabilization(1)
One Hampshire Street/Greater Boston/Cambridge 167,381 TBD
311 Arsenal Street/Greater Boston/Cambridge/Inner Suburbs 60,742 246,329
421 Park Drive/Greater Boston/Fenway 502,007
40, 50, and 60 Sylvan Road/Greater Boston/Route 128 466,334
Other/Greater Boston 155,305
651 Gateway Boulevard/San Francisco Bay Area/South San Francisco 87,515 261,199 138,286 487,000 5.0% 5.1%
269 East Grand Avenue/San Francisco Bay Area/South San Francisco 77,223 TBD
701 Dexter Avenue North/Seattle/Lake Union 263,230
148,257 2,139,008
900,746 3,688,301 2,390,000 6,980,000
Our share of investment(2)(3) 810,000 3,160,000 2,130,000 6,100,000
Refer to “Initial stabilized yield (unlevered)” under “Definitions and reconciliations” in Item 2 for additional information.(1)We expect to provide total estimated costs and related yields for each project with estimated stabilization in 2026 and beyond over the next several quarters. (2)Represents dollar amount rounded to the nearest 10 million and includes preliminary estimated amounts for projects listed as TBD. Total cost to complete for our development and redevelopment projects under construction have not been adjusted for the potential impact related to higher materials costs associated with potential tariffs. We are still evaluating the potential impact on costs and returns that can be significantly impacted by tariffs, the amount of foreign materials required, and/or the higher cost on domestic materials. Refer to “Executive summary” in Item 2 for additional details.(3)Represents our share of investment based on our ownership percentage upon completion of development or redevelopment projects.

All values are in US Dollars.

64

New Class A/A+ development and redevelopment properties: summary of pipeline

71% of Our Total Development and Redevelopment Pipeline RSF Is Within Our Megacampus™ Ecosystems

The following table summarizes the key information for all our development and redevelopment projects in North America as of March 31, 2025 (dollars in thousands):

Market<br><br>Property/Submarket Our<br><br>Ownership<br><br>Interest Book Value Square Footage
Total(1)
Future
Greater Boston
Megacampus: Alexandria Center® at One Kendall Square/Cambridge 100% 167,381 104,956 104,956
One Hampshire Street
Megacampus: The Arsenal on the Charles/Cambridge/Inner Suburbs 100% 299,765 344,890 59,469 404,359
311 Arsenal Street, 500 North Beacon Street, and 4 Kingsbury Avenue
Megacampus: 480 Arsenal Way and 446, 458, 500, and 550 Arsenal Street, and 99 Coolidge Avenue/<br><br>Cambridge/Inner Suburbs (2) 294,250 204,395 902,000 1,106,395
446, 458, 500, and 550 Arsenal Street, and 99 Coolidge Avenue
Megacampus: Alexandria Center® for Life Science – Fenway/Fenway 100% 669,613 529,686 529,686
401 and 421 Park Drive
Megacampus: Alexandria Center® for Life Science – Waltham/Route 128 100% 529,233 596,064 515,000 1,111,064
40, 50, and 60 Sylvan Road, and 35 Gatehouse Drive
Megacampus: Alexandria Center® at Kendall Square/Cambridge 100% 206,847 174,500 174,500
100 Edwin H. Land Boulevard
Megacampus: Alexandria Technology Square®/Cambridge 100% 8,064 100,000 100,000
Megacampus: 285, 299, 307, and 345 Dorchester Avenue/Seaport Innovation District 60.0% 290,685 1,040,000 1,040,000
10 Necco Street/Seaport Innovation District 100% 105,260 175,000 175,000
215 Presidential Way/Route 128 100% 6,816 112,000 112,000
Other development and redevelopment projects 100% 368,337 453,869 1,348,541 1,802,410
2,946,251 2,233,860 4,426,510 6,660,370
Refer to “Megacampus” under “Definitions and reconciliations” in Item 2 for additional information.<br><br>(1)Represents total square footage upon completion of development or redevelopment of one or more new Class A/A+ properties. Square footage presented includes the RSF of buildings currently in operation at properties that also have<br><br>future development or redevelopment opportunities. Upon expiration of existing in-place leases, we have the intent to demolish or redevelop the existing property subject to market conditions and leasing. Refer to “Investments in real<br><br>estate” under “Definitions and reconciliations” in Item 2 for additional information, including development and redevelopment square feet currently included in rental properties.<br><br>(2)We have a 75.7% interest in 99 Coolidge Avenue aggregating 204,395 RSF and a 100% interest in 446, 458, 500, and 550 Arsenal Street aggregating 902,000 RSF.

All values are in US Dollars.

65

New Class A/A+ development and redevelopment properties: summary of pipeline (continued)

Market<br><br>Property/Submarket Our<br><br>Ownership<br><br>Interest Book Value Square Footage
Total(1)
Future
San Francisco Bay Area
Megacampus: Alexandria Center® for Science and Technology – Mission Bay/Mission Bay 25.0% 123,380 109,435 (2) 109,435
1450 Owens Street
Megacampus: Alexandria Technology Center® – Gateway/South San Francisco 50.0% 287,764 259,689 291,000 550,689
651 Gateway Boulevard
Megacampus: Alexandria Center® for Advanced Technologies – South San Francisco/South San<br><br>Francisco 100% 83,878 107,250 90,000 197,250
211(3) and 269 East Grand Avenue
Megacampus: Alexandria Center® for Advanced Technologies – Tanforan/South San Francisco 100% 413,864 1,930,000 1,930,000
1122, 1150, and 1178 El Camino Real
Alexandria Center® for Life Science – Millbrae/South San Francisco 48.3% 156,100 348,401 348,401
201 and 231 Adrian Road and 30 Rollins Road
Megacampus: Alexandria Center® for Life Science – San Carlos/Greater Stanford 100% 464,630 1,497,830 1,497,830
960 Industrial Road, 987 and 1075 Commercial Street, and 888 Bransten Road
3825 and 3875 Fabian Way/Greater Stanford 100% 159,029 478,000 478,000
2100, 2200, 2300, and 2400 Geng Road/Greater Stanford 100% 37,999 240,000 240,000
Megacampus: 88 Bluxome Street/SoMa 100% 402,468 1,070,925 1,070,925
2,129,112 476,374 5,946,156 6,422,530
Refer to “Megacampus” under “Definitions and reconciliations” in Item 2 for additional information.<br><br>(1)Represents total square footage upon completion of development or redevelopment of one or more new Class A/A+ properties. Square footage presented includes the RSF of buildings currently in operation at properties that also have<br><br>future development or redevelopment opportunities. Upon expiration of existing in-place leases, we have the intent to demolish or redevelop the existing property subject to market conditions and leasing. Refer to “Investments in real<br><br>estate” under “Definitions and reconciliations” in Item 2 for additional information, including development and redevelopment square feet currently included in rental properties.<br><br>(2)During the three months ended December 31, 2024, we executed a letter of intent with a biomedical institution for the sale of a condominium interest aggregating 103,361 RSF, or approximately 49% of the development project, with the<br><br>transaction expected to close in the second half of 2025. Accordingly, we adjusted the development project RSF and its related book value to reflect 109,435 RSF.<br><br>(3)We own a partial interest in this property through a real estate joint venture. Refer to Note 4 – “Consolidated and unconsolidated real estate joint ventures” to our unaudited consolidated financial statements in Item 1 for additional details.

All values are in US Dollars.

66

New Class A/A+ development and redevelopment properties: summary of pipeline (continued)

Market<br><br>Property/Submarket Our<br><br>Ownership<br><br>Interest Book Value Square Footage
Total(1)
Future
San Diego
Megacampus: One Alexandria Square/Torrey Pines 100% 428,104 241,504 125,280 366,784
10935 and 10945 Alexandria Way and 10975 and 10995 Torreyana Road
Megacampus: Campus Point by Alexandria/University Town Center 55.0% 547,241 426,927 967,457 1,394,384
10010(2), 10140(2), 10210, and 10260 Campus Point Drive and 4135, 4161, 4165,<br><br>and 4224 Campus Point Court
Megacampus: SD Tech by Alexandria/Sorrento Mesa 50.0% 347,577 235,361 493,845 729,206
9805 Scranton Road and 10075 Barnes Canyon Road
11255 and 11355 North Torrey Pines Road/Torrey Pines 100% 156,640 215,000 215,000
Megacampus: 5200 Illumina Way/University Town Center 51.0% 17,469 451,832 451,832
9625 Towne Centre Drive/University Town Center 30.0% 837 100,000 100,000
Megacampus: Sequence District by Alexandria/Sorrento Mesa 100% 46,865 1,798,915 1,798,915
6260, 6290, 6310, 6340, 6350, and 6450 Sequence Drive
Scripps Science Park by Alexandria/Sorrento Mesa 100% 42,465 154,308 154,308
10256 and 10260 Meanley Drive
4075 Sorrento Valley Boulevard/Sorrento Valley 100% 19,508 144,000 144,000
Other development and redevelopment projects (3) 77,878 475,000 475,000
1,684,584 903,792 4,925,637 5,829,429
Refer to “Megacampus” under “Definitions and reconciliations” in Item 2 for additional information.<br><br>(1)Represents total square footage upon completion of development or redevelopment of one or more new Class A/A+ properties. Square footage presented includes RSF of buildings currently in operation at properties that also have<br><br>inherent future development or redevelopment opportunities. Upon expiration of existing in-place leases, we have the intent to demolish or redevelop the existing property subject to market conditions and leasing. Refer to “Investments in<br><br>real estate” under “Definitions and reconciliations” in Item 2 for additional information, including development and redevelopment square feet currently included in rental properties.<br><br>(2)We have a 100% interest in this property.<br><br>(3)Includes a property in which we own a partial interest through a real estate joint venture.

All values are in US Dollars.

67

New Class A/A+ development and redevelopment properties: summary of pipeline (continued)

Market<br><br>Property/Submarket Our<br><br>Ownership<br><br>Interest Book Value Square Footage
Total(1)
Future
Seattle
Megacampus: Alexandria Center® for Advanced Technologies – South Lake Union/Lake Union (2) 548,306 227,577 1,057,400 1,284,977
601 and 701 Dexter Avenue North and 800 Mercer Street
1010 4th Avenue South/SoDo 100% 60,921 544,825 544,825
410 West Harrison Street/Elliott Bay 100% 91,000 91,000
Megacampus: Alexandria Center® for Advanced Technologies – Canyon Park/Bothell 100% 18,521 230,000 230,000
21660 20th Avenue Southeast
Other development and redevelopment projects 100% 146,711 706,087 706,087
774,459 227,577 2,629,312 2,856,889
Maryland
Megacampus: Alexandria Center® for Life Science – Shady Grove/Rockville 100% 23,041 296,000 296,000
9830 Darnestown Road
23,041 296,000 296,000
Research Triangle
Megacampus: Alexandria Center® for Life Science – Durham/Research Triangle 100% 160,292 2,060,000 2,060,000
Megacampus: Alexandria Center® for Advanced Technologies and Agtech – Research Triangle/<br><br>Research Triangle 100% 108,266 1,170,000 1,170,000
4 and 12 Davis Drive
Megacampus: Alexandria Center® for NextGen Medicines/Research Triangle 100% 110,826 1,055,000 1,055,000
3029 East Cornwallis Road
Megacampus: Alexandria Center® for Sustainable Technologies/Research Triangle 100% 54,534 750,000 750,000
120 TW Alexander Drive, 2752 East NC Highway 54, and 10 South Triangle Drive
100 Capitola Drive/Research Triangle 100% 65,965 65,965
Other development and redevelopment projects 100% 4,185 76,262 76,262
438,103 5,177,227 5,177,227
Refer to “Megacampus” under “Definitions and reconciliations” in Item 2 for additional information.<br><br>(1)Represents total square footage upon completion of development or redevelopment of one or more new Class A/A+ properties. Square footage presented includes the RSF of buildings currently in operation at properties that also have<br><br>inherent future development or redevelopment opportunities. Upon expiration of existing in-place leases, we have the intent to demolish or redevelop the existing property. Refer to “Investments in real estate” under “Definitions and<br><br>reconciliations” in Item 2 for additional information, including development and redevelopment square feet currently included in rental properties.<br><br>(2)We have a 100% interest in 601 and 701 Dexter Avenue North aggregating 415,977 RSF and a 60% interest in the future development project at 800 Mercer Street aggregating 869,000 RSF.

All values are in US Dollars.

68

New Class A/A+ development and redevelopment properties: summary of pipeline (continued)

Market<br><br>Property/Submarket Our<br><br>Ownership<br><br>Interest Book Value Square Footage
Total(1)
Future
New York City
Megacampus: Alexandria Center® for Life Science – New York City/New York City 100% 171,060 550,000 (2) 550,000
171,060 550,000 550,000
Texas
Alexandria Center® for Advanced Technologies at The Woodlands/Greater Houston 100% 49,198 73,298 116,405 189,703
8800 Technology Forest Place
1001 Trinity Street and 1020 Red River Street/Austin 100% 10,694 250,010 250,010
Other development and redevelopment projects 100% 57,669 344,000 344,000
117,561 73,298 710,415 783,713
Canada 100% 50,245 132,881 371,743 504,624
Other development and redevelopment projects 100% 122,555 724,349 724,349
Total pipeline as of March 31, 2025, excluding properties held for sale 8,456,971 4,047,782 25,757,349 29,805,131
Properties held for sale 139,797 1,853,856 1,853,856
Total pipeline as of March 31, 2025 8,596,768 4,047,782 27,611,205 31,658,987

All values are in US Dollars.

Refer to “Megacampus” under “Definitions and reconciliations” in Item 2 for additional information.

(1)Total square footage includes 2,780,364 RSF of buildings currently in operation that we expect to demolish or redevelop and commence future construction subject to market conditions and leasing. Refer to “Investments in real estate”

under “Definitions and reconciliations” in Item 2 for additional information, including development and redevelopment square feet currently included in rental properties.

(2)During the three months ended September 30, 2024, we filed a lawsuit against the New York City Health + Hospitals Corporation and the New York City Economic Development Corporation for fraud and breach of contract concerning our

option to ground lease a land parcel to develop a future world-class life science building within the Alexandria Center® for Life Science – New York City Megacampus. Refer to “Legal proceedings” in Item 1 under Part II – Other Information

for additional details.

(3)Includes $3.7 billion of projects that are currently under construction.

69

Results of operations

We present a tabular comparison of items, whether gain or loss, that may facilitate a high-level understanding of our results

and provide context for the disclosures included in our annual report on Form 10-K for the year ended December 31, 2024 and our

subsequent quarterly reports on Form 10-Q. We believe that such tabular presentation promotes a better understanding for investors of

the corporate-level decisions made and activities performed that significantly affect comparison of our operating results from period to

period. We also believe that this tabular presentation will supplement for investors an understanding of our disclosures and real estate

operating results. Gains or losses on sales of real estate and impairments of real estate are related to corporate-level decisions to

dispose of real estate. Gains or losses on early extinguishment of debt are related to corporate-level financing decisions focused on our

capital structure strategy. Significant realized and unrealized gains or losses on non-real estate investments, impairments of real estate

and non-real estate investments, and acceleration of stock compensation expense due to the resignations of executive officers are not

related to the operating performance of our real estate assets as they result from strategic, corporate-level non-real estate investment

decisions and external market conditions. Impairments of non-real estate investments and changes in the provision for expected credit

losses on financial instruments are not related to the operating performance of our real estate as they represent the write-down of non-

real estate investments when their fair values decrease below their respective carrying values due to changes in general market or

other conditions outside of our control. Significant items, whether a gain or loss, included in the tabular disclosure for current periods

are described in further detail in Item 2. Key items included in net income attributable to Alexandria’s common stockholders for the three

months ended March 31, 2025 and 2024 and the related per share amounts were as follows (in millions, except per share amounts):

Three Months Ended March 31,
2025 2024 2025 2024
Amount Per Share – Diluted
Unrealized (losses) gains on non-real estate investments $(68.1) $29.2 $(0.40) $0.17
Gain on sales of real estate 13.2 0.4 0.08
Impairment of non-real estate investments (11.2) (14.7) (0.07) (0.09)
Impairment of real estate (32.2) (0.19)
Increase in provision for expected credit losses on financial instruments (0.3)
Total $(98.6) $14.9 $(0.58) $0.08

Refer to Note 3 – “Investments in real estate,”  Note 5 – “Leases,” and Note 7 – “Investments” to our unaudited consolidated

financial statements in Item 1 for additional information.

70

Same properties

We supplement an evaluation of our results of operations with an evaluation of operating performance of certain of our

properties, referred to as “Same Properties.” For additional information on the determination of our Same Properties portfolio, refer to

“Same property comparisons” under “Definitions and reconciliations” in Item 2. The following table presents information regarding our

Same Properties for the three months ended March 31, 2025:

Three Months Ended<br><br>March 31, 2025
Percentage change in net operating income over comparable period from prior year (3.1)% (1)
Percentage change in net operating income (cash basis) over comparable period from prior year 5.1% (1)(2)
Operating margin 68%
Number of Same Properties 333
RSF 34,099,158
Occupancy – current-period average 93.3%
Occupancy – same-period prior-year average 94.3%

(1)Includes certain leases expiring during the three months ended March 31, 2025 aggregating 768,080 RSF at six properties across four submarkets. Excluding the

impact of the properties with these leases, same property net operating income changes for the three months ended March 31, 2025 would have been 0.1% and

9.0% (cash basis). Refer to “Summary of occupancy percentages in North America” in Item 2 for additional details.

(2)Includes the impact of expiring initial free rent concessions that burned off after January 1, 2024 in connection with the development and redevelopment projects that

were placed into service in 2023 and, accordingly are part of our same property pool in the first quarter of 2025, including 15 Necco Street in our Seaport Innovation

District submarket and 751 Gateway Boulevard in our South San Francisco submarket. Excluding the impact of these expiring initial free rent concessions, same

property net operating income change (cash basis) for the three months ended March 31, 2025 would have been 0.4%.

The following table reconciles the number of Same Properties to total properties for the three months ended March 31, 2025:

Development – under construction Properties
99 Coolidge Avenue 1
500 North Beacon Street and 4 Kingsbury Avenue 2
1450 Owens Street 1
10935, 10945, and 10955 Alexandria Way 3
10075 Barnes Canyon Road 1
421 Park Drive 1
4135 Campus Point Court 1
701 Dexter Avenue North 1
11
Development – placed into service after<br><br>January 1, 2024 Properties
9810 Darnestown Road 1
9820 Darnestown Road 1
1150 Eastlake Avenue East 1
4155 Campus Point Court 1
201 Brookline Avenue 1
9808 Medical Center Drive 1
230 Harriet Tubman Way 1
7
Redevelopment – under construction Properties
40, 50, and 60 Sylvan Road 3
269 East Grand Avenue 1
651 Gateway Boulevard 1
401 Park Drive 1
8800 Technology Forest Place 1
311 Arsenal Street 1
One Hampshire Street 1
Canada 4
Other 2
15 Redevelopment – placed into service after<br><br>January 1, 2024 Properties
--- ---
840 Winter Street 1
Alexandria Center® for Advanced Technologies –<br><br>Monte Villa Parkway 6
7
Acquisitions after January 1, 2024 Properties
Other 3
3
Unconsolidated real estate JVs 4
Properties held for sale 6
Total properties excluded from Same Properties 53
Same Properties 333
Total properties in North America as of<br><br>March 31, 2025 386

71

Comparison of results for the three months ended March 31, 2025 to the three months ended March 31, 2024

The following table presents a comparison of the components of net operating income for our Same Properties and Non-Same

Properties for the three months ended March 31, 2025, compared to the three months ended March 31, 2024 (dollars in thousands).

Refer to “Definitions and reconciliations” in Item 2 for definitions of “Tenant recoveries” and “Net operating income” and their

reconciliations from the most directly comparable financial measures presented in accordance with GAAP, income from rentals and net

income, respectively.

Three Months Ended March 31,
2025 2024 $ Change % Change
Income from rentals:
Same Properties $469,387 $476,074 $(6,687) (1.4)%
Non-Same Properties 82,725 105,326 (22,601) (21.5)
Rental revenues 552,112 581,400 (29,288) (5.0)
Same Properties 170,823 155,405 15,418 9.9
Non-Same Properties 20,240 18,746 1,494 8.0
Tenant recoveries 191,063 174,151 16,912 9.7
Income from rentals 743,175 755,551 (12,376) (1.6)
Same Properties 346 340 6 1.8
Non-Same Properties 14,637 13,217 1,420 10.7
Other income 14,983 13,557 1,426 10.5
Same Properties 640,556 631,819 8,737 1.4
Non-Same Properties 117,602 137,289 (19,687) (14.3)
Total revenues 758,158 769,108 (10,950) (1.4)
Same Properties 203,497 180,739 22,758 12.6
Non-Same Properties 22,898 37,575 (14,677) (39.1)
Rental operations 226,395 218,314 8,081 3.7
Same Properties 437,059 451,080 (14,021) (3.1)
Non-Same Properties 94,704 99,714 (5,010) (5.0)
Net operating income $531,763 $550,794 $(19,031) (3.5)% (1)
Net operating income – Same Properties $437,059 $451,080 $(14,021) (3.1)%
Straight-line rent revenue (6,396) (39,287) 32,891 (83.7)
Amortization of acquired below-market leases (10,002) (11,525) 1,523 (13.2)
Net operating income – Same Properties (cash basis) $420,661 $400,268 $20,393 5.1%

(1)Decrease in total net operating income includes the impact of operating properties disposed of after January 1, 2024. Excluding these dispositions, the increase in net

operating income for the three months ended March 31, 2025 would have been 2.2%.

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Income from rentals

Total income from rentals for the three months ended March 31, 2025 decreased by $12.4 million, or 1.6%, to $743.2 million,

compared to $755.6 million for the three months ended March 31, 2024, due to a decrease in rental revenues, as discussed below.

Rental revenues

Total rental revenues for the three months ended March 31, 2025 decreased by $29.3 million, or 5.0%, to $552.1 million,

compared to $581.4 million for the three months ended March 31, 2024. The decrease was primarily related to our Non-Same

Properties resulting from the dispositions of real estate assets since January 1, 2024.

Same Properties’ rental revenues for the three months ended March 31, 2025 decreased by $6.7 million, or 1.4%, to

$469.4 million, compared to $476.1 million for the three months ended March 31, 2024, primarily due to a decrease in rental revenues

from certain lease expirations aggregating 768,080 RSF during the three months ended March 31, 2025, comprising the following: (i)

182,054 RSF at the Alexandria Technology Square® Megacampus in our Cambridge submarket, (ii) 234,249 RSF at 409 Illinois Street in

our Mission Bay submarket, (iii) one property aggregating 104,531 RSF in our Research Triangle market, and (iv) two properties

aggregating 247,246 RSF in our Austin submarket. The decrease was partially offset by the increase in Same Properties’ rental

revenues due to rental rate increases of 18.5% and 7.5% (cash basis) on lease renewals and re-leasing for the three months ended

March 31, 2025.

Tenant recoveries

Tenant recoveries for the three months ended March 31, 2025 increased by $16.9 million, or 9.7%, to $191.1 million,

compared to $174.2 million for the three months ended March 31, 2024, primarily in connection with Same Properties.

Same Properties’ tenant recoveries for the three months ended March 31, 2025 increased by $15.4 million, or 9.9%, to

$170.8 million, compared to $155.4 million for the three months ended March 31, 2024, primarily due to higher operating expenses

during the three months ended March 31, 2025, as discussed under “Rental operations” below. As of March 31, 2025, 91% of our

leases (on an annual rental revenue basis) were triple net leases, which require tenants to pay substantially all real estate taxes,

insurance, utilities, repairs and maintenance, common area expenses, and other operating expenses (including increases thereto) in

addition to base rent.

Rental operations

Total rental operating expenses for the three months ended March 31, 2025 increased by $8.1 million, or 3.7%, to

$226.4 million, compared to $218.3 million for the three months ended March 31, 2024. The increase was primarily due to higher rental

operating expenses related to our Same Properties, as discussed below, partially offset by the decrease in Non-Same Properties’ rental

operating expenses of $14.7 million primarily as a result of dispositions of real estate assets since January 1, 2024.

Same Properties’ rental operating expenses increased by $22.8 million, or 12.6%, to $203.5 million during the three months

ended March 31, 2025, compared to $180.7 million for the three months ended March 31, 2024, primarily as the result of increases in

(i) utility expenses aggregating $5.6 million due to higher utility consumption related to certain tenants’ increased operations in our

Greater Boston and San Diego markets, (ii) property taxes aggregating $3.9 million due to higher assessed property values in our

Greater Boston and San Francisco Bay Area markets, and (iii) repairs and maintenance expenses aggregating $3.7 million due to a

more severe winter in 2025 compared to that in 2024 in our Greater Boston market.

Depreciation and amortization

Depreciation and amortization expense for the three months ended March 31, 2025 increased by $54.5 million, or 19.0%, to

$342.1 million, compared to $287.6 million for the three months ended March 31, 2024. The increase primarily reflects the change in

useful lives related to certain projects expected to be redeveloped from office to laboratory use prior to the end of their previous useful

lives. In addition, the increase relates to 1.9 million RSF of development and redevelopment projects placed into service subsequent to

January 1, 2024 and three operating properties aggregating 401,560 RSF acquired subsequent to January 1, 2024, partially offset by

the decrease in depreciation and amortization related to properties that were sold or classified as held for sale subsequent to January 1,

2024.

Impairment of real estate

During the three months ended March 31, 2025, we recognized an impairment charge aggregating $32.2 million, related to a

ground lease entered into in 2021 for a future development site in our San Francisco Bay Area market. As of December 31, 2024, we

had a right-of-use asset aggregating $32.4 million related to our investment into this ground lease. During the three months ended

March 31, 2025, based on our current financial outlook for this project, we made the determination to no longer proceed with this

project. Consequently, we recognized an impairment charge aggregating $32.2 million to write off our remaining right-of-use asset

balance. We do not expect to make additional future payments in connection with this project.

73

General and administrative expenses

General and administrative expenses for the three months ended March 31, 2025 decreased by $16.4 million, or 34.8%, to

$30.7 million, compared to $47.1 million for the three months ended March 31, 2024, primarily due to cost-control and efficiency

initiatives implemented in prior reporting periods, including reduction in headcount, restructuring of compensation plans, systems

upgrades, and process improvements. As a percentage of net operating income, our general and administrative expenses for the

trailing twelve months ended March 31, 2025 and 2024 were 6.9% and 9.5%, respectively.

Interest expense

Interest expense for the three months ended March 31, 2025 and 2024 consisted of the following (dollars in thousands):

Three Months Ended March 31,
Component 2025 2024 Change
Gross interest $130,941 $122,680 $8,261
Capitalized interest (80,065) (81,840) 1,775
Interest expense $50,876 $40,840 $10,036
Average debt balance outstanding(1) $12,815,953 $12,056,184 $759,769
Weighted-average annual interest rate(2) 4.1% 4.1% —%

(1)Represents the average debt balance outstanding during the respective periods.

(2)Represents annualized total interest incurred divided by the average debt balance outstanding during the respective periods.

The net change in interest expense during the three months ended March 31, 2025, compared to the three months ended

March 31, 2024, resulted from the following (dollars in thousands):

Component Effective Date Change
Increases in interest incurred due to:
Issuances of debt:
550 million of unsecured senior notes payable due 2035 5.66% February 2025 $4,047
600 million of unsecured senior notes payable due 2054 5.71% February 2024 4,127
400 million of unsecured senior notes payable due 2036 5.38% February 2024 2,574
Increases in construction borrowings and interest rates under secured notes payable 7.20% 125
Lower average outstanding balances and/or rate decreases on borrowings under commercial paper program and unsecured senior line of credit (3,219)
Other increase in interest 607
Change in gross interest 8,261
Decrease in capitalized interest 1,775
Total change in interest expense $10,036

All values are in US Dollars.

(1)Represents the weighted-average interest rate as of the end of the applicable period, including amortization of loan fees, amortization of debt premiums (discounts), and

other bank fees.

Investment (loss) income

During the three months ended March 31, 2025, we recognized investment loss aggregating $50.0 million, which consisted of

$29.3 million of realized gains, $68.1 million of unrealized losses, and $11.2 million of impairment charges.

During the three months ended March 31, 2024, we recognized investment income aggregating $43.3 million, which consisted

of $28.8 million of realized gains, $29.2 million of unrealized gains, and $14.7 million of impairment charges.

For more information about our investments, refer to Note 7 – “Investments” to our unaudited consolidated financial

statements. For our impairments accounting policy, refer to “Investments” in Note 2 – “Summary of significant accounting policies” to

our unaudited consolidated financial statements in Item 1.

Other comprehensive income (loss)

Total other comprehensive income for the three months ended March 31, 2025 aggregated $50 thousand, compared to total

other comprehensive loss of $7.9 million for the three months ended March 31, 2024. The difference is primarily due to the unrealized

foreign currency translation gains related to our operations in Canada.

74

Summary of capital expenditures

Our construction spending for the three months ended March 31, 2025 and projected spending for the year ending December

31, 2025 consist of the following (in thousands):

Three Months Ended<br><br>March 31, 2025 Projected Guidance Midpoint for Year Ending December 31, 2025
Construction of Class A/A+ properties:
Active construction projects
Under construction(1) $ 307,490
Future pipeline pre-construction
Primarily Megacampus expansion pre-construction work (entitlement,<br><br>design, and site work) 92,955
Revenue- and non-revenue-enhancing capital expenditures 58,464 (2)
Construction spending (before contributions from noncontrolling interests<br><br>or tenants) 458,909
Contributions from noncontrolling interests (consolidated real estate<br><br>joint ventures) (63,247) (3)
Tenant-funded and -built landlord improvements (39,950)
Total construction spending $ 355,712
2025 guidance range for construction spending 1,450,000 – 2,050,000

All values are in US Dollars.

(1)Includes projects under construction aggregating 4.0 million RSF. Refer to “Investments in real estate” in Item 2 for additional details.

(2)Represents revenue-enhancing and non-revenue-enhancing capital expenditures before contributions from noncontrolling interests and tenant-funded and tenant-built

landlord improvements for the year ending December 31, 2025. Our share of the 2025 revenue-enhancing and non-revenue-enhancing capital expenditures is projected

to be $370 million at the midpoint of our guidance for 2025 construction spending.

(3)Represents contractual capital commitments from existing consolidated real estate joint venture partners to fund construction.

Projected capital contributions from partners in consolidated real estate joint ventures to fund construction

The following table summarizes projected capital contributions from partners in our existing consolidated joint ventures to fund

construction through 2027 and beyond (in thousands):

Projected timing Amount(1)
April 1, 2025 through December 31, 2026 $247,964
2027 and beyond 166,896
Total $414,860

(1)Amounts represent reductions to our consolidated construction spending.

Average real estate basis used for capitalization of interest

Our construction spending includes capitalized interest. The table below provides key categories of interest capitalized during

the three months ended March 31, 2025 (in thousands):

Three Months Ended March 31, 2025
Average Real Estate Basis Capitalized Percentage of Total<br><br>Average Real Estate<br><br>Basis Capitalized
Construction of Class A/A+ properties:
Active construction projects
Under construction 2,951,331 37%
Future pipeline pre-construction
Primarily Megacampus expansion pre-construction work (entitlement, design,<br><br>and site work) 4,149,799 51
Smaller redevelopments and repositioning capital projects 925,436 12
8,026,566 100%

All values are in US Dollars.

(1)Average real estate basis capitalized during the three months ended March 31, 2025, which related to our future pipeline pre-construction activities, includes 29% from

four key active and future Megacampus development projects.

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Projected results

We present updated guidance for EPS attributable to Alexandria’s common stockholders – diluted, funds from operations per

share attributable to Alexandria’s common stockholders – diluted, and funds from operations per share attributable to Alexandria’s

common stockholders – diluted, as adjusted, based on our current view of existing market conditions and other assumptions for the

year ending December 31, 2025, as set forth in the tables below. The tables below also provide a reconciliation of EPS attributable to

Alexandria’s common stockholders – diluted, the most directly comparable financial measure presented in accordance with GAAP, to

funds from operations per share and funds from operations per share, as adjusted, non-GAAP measures, and other key assumptions

included in our updated guidance for the year ending December 31, 2025. There can be no assurance that actual amounts will not be

materially higher or lower than these expectations. Refer to our discussion of “Forward-looking statements” included in the beginning of

this Item 2.

The midpoint of our guidance range for 2025 funds from operations per share – diluted, as adjusted, was reduced by seven

cents, or 75 bps. Key changes to our guidance assumptions include the following:

•Slower than anticipated re-leasing of expiring spaces and lease-up of vacancy in our operating portfolio and our development

and redevelopment pipeline, resulting in the following changes to the midpoints of our guidance ranges:

•70 bps reduction in occupancy percentage in North America as of December 31, 2025,

•70 bps and 20 bps reduction in 2025 same property net operating income performance and same property net operating

income performance (cash basis), respectively, and

•$15 million reduction in 2025 straight-line rent revenue.

•A $20 million reduction to the midpoint of our guidance range for 2025 capitalization of interest with a corresponding

$20 million increase to the midpoint of our guidance range for 2025 interest expense, primarily due to various current and

future pipeline projects that are anticipated to cease construction activities in the latter part of the year.

•A $17 million reduction to the midpoint of our guidance range for 2025 general and administrative expenses from additional

cost control initiatives, including personnel-related costs and streamlining of business processes.

Projected 2025 Earnings per Share and Funds From Operations per<br><br>Share Attributable to Alexandria’s Common Stockholders – Diluted As of 4/28/25 Key Changes<br><br>to Midpoint
Earnings per share(1) 1.36 to 1.56
Depreciation and amortization of real estate assets
Gain on sales of real estate (2)
Impairment of real estate – rental properties (3)
Allocation of unvested restricted stock awards
Funds from operations per share(4) 8.51 to 8.71
Unrealized losses on non-real estate investments
Impairment of non-real estate investments (4)
Impairment of real estate
Allocation to unvested restricted stock awards
Funds from operations per share, as adjusted(4) 9.16 to 9.36
Midpoint Reduction of 7-<br><br>cents, or 75 bps

All values are in US Dollars.

(1)Excludes unrealized gains or losses on non-real estate investments after March 31, 2025 that are required to be recognized in earnings and are excluded from funds

from operations per share, as adjusted.

(2)Refer to “Dispositions and sales of partial interests” in Item 2 for additional information.

(3)Refer to Note 16 – “Subsequent events” to our unaudited consolidated financial statements in Item 1 for additional information.

(4)Refer to “Funds from operations and funds from operations, as adjusted, attributable to Alexandria Real Estate Equities, Inc.’s common stockholders” under “Definitions

and reconciliations” in Item 2 for additional information.

76

Key Assumptions(1)<br><br>(Dollars in millions) As of 4/28/25 As of 1/27/25 Key Changes<br><br>to Midpoint
Low High Low High
Occupancy percentage in North America as of December 31, 2025 90.9% 92.5% 91.6% 93.2% 70 bps reduction
Lease renewals and re-leasing of space:
Rental rate changes 9.0% 17.0% 9.0% 17.0% No change
Rental rate changes (cash basis) 0.5% 8.5% 0.5% 8.5%
Same property performance:
Net operating income (3.7)% (1.7)% (3.0)% (1.0)% 70 bps reduction
Net operating income (cash basis) (1.2)% 0.8% (1.0)% 1.0% 20 bps reduction
Straight-line rent revenue $96 $116 $111 $131 $15 million reduction
General and administrative expenses $112 $127 $129 $144 $17 million reduction
Capitalization of interest $320 $350 $340 $370 $20 million reduction
Interest expense $185 $215 $165 $195 $20 million increase
Realized gains on non-real estate investments(2) $100 $130 $100 $130 No change

(1)Our assumptions presented in the table above are subject to a number of variables and uncertainties, including those discussed as “Forward-looking statements” under

Part I; “Item 1A. Risk factors”; and “Item 7. Management’s discussion and analysis of financial condition and results of operations” of our annual report on Form 10-K for

the year ended December 31, 2024, as well as in “Item 1A. Risk factors” within “Part II – Other information” of this quarterly report on Form 10-Q. To the extent our full-

year earnings guidance is updated during the year, we will provide additional disclosure supporting reasons for any significant changes to such guidance.

(2)Represents realized gains and losses included in funds from operations per share – diluted, as adjusted, and excludes significant impairments realized on non-real

estate investments, if any. Refer to Note 7 – “Investments” to our unaudited consolidated financial statements in Item 1 for additional details.

Key Credit Metric Targets(1) As of 4/28/25 As of 1/27/25 Key Changes
Net debt and preferred stock to Adjusted EBITDA –<br><br>fourth quarter of 2025 annualized Less than or equal to 5.2x Less than or equal to 5.2x No change
Fixed-charge coverage ratio – fourth quarter of 2025<br><br>annualized 4.0x to 4.5x 4.0x to 4.5x

(1)Refer to “Definitions and reconciliations” in Item 2 for additional information.

77

Consolidated and unconsolidated real estate joint ventures

We present components of balance sheet and operating results information for the noncontrolling interest share of our

consolidated real estate joint ventures and for our share of investments in unconsolidated real estate joint ventures to help investors

estimate balance sheet and operating results information related to our partially owned entities. These amounts are estimated by

computing, for each joint venture that we consolidate in our financial statements, the noncontrolling interest percentage of each financial

item to arrive at the cumulative noncontrolling interest share of each component presented. In addition, for our real estate joint ventures

that we do not control and do not consolidate, we apply our economic ownership percentage to the unconsolidated real estate joint

ventures to arrive at our proportionate share of each component presented. Refer to Note 4 – “Consolidated and unconsolidated real

estate joint ventures” to our unaudited consolidated financial statements in Item 1 for further discussion.

Consolidated Real Estate Joint Ventures
Property/Market/Submarket Noncontrolling<br><br>Interest Share Operating RSF<br><br>at 100%
50 and 60 Binney Street/Greater Boston/Cambridge/Inner Suburbs 66.0% 532,395
75/125 Binney Street/Greater Boston/Cambridge/Inner Suburbs 60.0% 388,270
100 and 225 Binney Street and 300 Third Street/Greater Boston/Cambridge/Inner Suburbs 70.0% 870,641
99 Coolidge Avenue/Greater Boston/Cambridge/Inner Suburbs 24.3% 116,414 (1)
15 Necco Street/Greater Boston/Seaport Innovation District 43.3% 345,996
285, 299, 307, and 345 Dorchester Avenue/Greater Boston/Seaport Innovation District 40.0% (1)
Alexandria Center® for Science and Technology – Mission Bay/San Francisco Bay Area/<br><br>Mission Bay(2) 75.0% 1,001,281
601, 611, 651(1), 681, 685, and 701 Gateway Boulevard/San Francisco Bay Area/<br><br>South San Francisco 50.0% 851,991
751 Gateway Boulevard/San Francisco Bay Area/South San Francisco 49.0% 230,592
211(1) and 213 East Grand Avenue/San Francisco Bay Area/South San Francisco 70.0% 300,930
500 Forbes Boulevard/San Francisco Bay Area/South San Francisco 90.0% 155,685
Alexandria Center® for Life Science – Millbrae/San Francisco Bay Area/South San Francisco 51.7% 285,346
3215 Merryfield Row/San Diego/Torrey Pines 70.0% 170,523
Campus Point by Alexandria/San Diego/University Town Center(3) 45.0% 1,227,133
5200 Illumina Way/San Diego/University Town Center 49.0% 792,687
9625 Towne Centre Drive/San Diego/University Town Center 70.0% 163,648
SD Tech by Alexandria/San Diego/Sorrento Mesa(4) 50.0% 816,519
Pacific Technology Park/San Diego/Sorrento Mesa 50.0% 544,352
Summers Ridge Science Park/San Diego/Sorrento Mesa(5) 70.0% 316,531
1201 and 1208 Eastlake Avenue East/Seattle/Lake Union 70.0% 206,134
199 East Blaine Street/Seattle/Lake Union 70.0% 115,084
400 Dexter Avenue North/Seattle/Lake Union 70.0% 290,754
800 Mercer Street/Seattle/Lake Union 40.0% (1)
Unconsolidated Real Estate Joint Ventures
Property/Market/Submarket Our Ownership<br><br>Share(6) Operating RSF<br><br>at 100%
1655 and 1725 Third Street/San Francisco Bay Area/Mission Bay 10.0% 586,208
1450 Research Boulevard/Maryland/Rockville 73.2% (7) 42,012
101 West Dickman Street/Maryland/Beltsville 58.4% (7) 135,949

Refer to “Joint venture financial information” under “Definitions and reconciliations” in Item 2 for additional details.

(1)Represents a property currently under construction or in our development and redevelopment pipeline. Refer to “New Class A/A+ development and redevelopment

properties” in Item 2 for additional details.

(2)Includes 409 and 499 Illinois Street, 1450, 1500, and 1700 Owens Street, and 455 Mission Bay Boulevard South.

(3)Includes 10210, 10260, 10290, and 10300 Campus Point Drive and 4110, 4135, 4155, 4161, 4165, 4224, and 4242 Campus Point Court.

(4)Includes 9605, 9645, 9675, 9725, 9735, 9805, 9808, 9855, and 9868 Scranton Road and 10055, 10065, and 10075 Barnes Canyon Road.

(5)Includes 9965, 9975, 9985, and 9995 Summers Ridge Road.

(6)In addition to the real estate joint ventures listed, we hold an interest in one insignificant unconsolidated real estate joint venture in North America.

(7)Represents a joint venture with a local real estate operator in which our joint venture partner manages the day-to-day activities that significantly affect the economic

performance of the joint venture.

78

The following table presents key terms related to our unconsolidated real estate joint ventures’ secured loans as of March 31,

2025 (dollars in thousands):

Maturity Date Stated Rate Interest<br><br>Rate(1) At 100%
Unconsolidated Joint Venture Aggregate<br><br>Commitment Debt Balance(2)
101 West Dickman Street 11/10/26 SOFR+1.95% (3) 6.35% $26,750 19,139
1450 Research Boulevard 12/10/26 SOFR+1.95% (3) 6.41% 13,000 8,998
1655 and 1725 Third Street(4) 2/10/35 6.37% 6.44% 500,000 496,658
$539,750 524,795

All values are in US Dollars.

(1)Includes interest expense and amortization of loan fees.

(2)Represents outstanding principal, net of unamortized deferred financing costs, as of March 31, 2025.

(3)This loan is subject to a fixed SOFR floor of 0.75%.

(4)During the three months ended March 31, 2025, the unconsolidated real estate joint venture refinanced $500 million of its $600 million existing fixed-rate debt with a new

secured note payable maturing in 2035. The remaining debt balance of approximately $100 million was repaid through contributions from the unconsolidated joint

venture partners, including our share of $10.8 million. As of March 31, 2025, our investment in this unconsolidated real estate joint venture was $21.2 million.

The following tables present information related to the operating results and financial positions of our consolidated and

unconsolidated real estate joint ventures as of and for the three months ended March 31, 2025 (in thousands):

Three Months Ended March 31, 2025
Noncontrolling Interest<br><br>Share of Consolidated<br><br>Real Estate Joint Ventures Our Share of<br><br>Unconsolidated<br><br>Real Estate Joint Ventures
Total revenues $116,637 $2,575
Rental operations (34,769) (1,048)
81,868 1,527
General and administrative (633) (19)
Interest (424) (961)
Depreciation and amortization of real estate assets (33,411) (1,054)
Fixed returns allocated to redeemable noncontrolling interests(1) 201
$47,601 $(507)
Straight-line rent and below-market lease revenue $3,652 $158
Funds from operations(1) $81,012 $547

Refer to “Joint venture financial information” under “Definitions and reconciliations” in Item 2 for additional details.

(1)Refer to “Funds from operations and funds from operations, as adjusted, attributable to Alexandria Real Estate Equities, Inc.’s common stockholders” under “Definitions

and reconciliations” in Item 2 for the definition and its reconciliation from the most directly comparable financial measure presented in accordance with GAAP.

As of March 31, 2025
Noncontrolling Interest<br><br>Share of Consolidated<br><br>Real Estate Joint Ventures Our Share of<br><br>Unconsolidated<br><br>Real Estate Joint Ventures
Investments in real estate $4,254,013 $109,352
Cash, cash equivalents, and restricted cash 131,409 3,635
Other assets 424,919 10,291
Secured notes payable (36,562) (67,431)
Other liabilities (238,868) (5,761)
Redeemable noncontrolling interests (9,612)
$4,525,299 $50,086

During the three months ended March 31, 2025 and 2024, our consolidated real estate joint ventures distributed an aggregate

of $66.0 million and $59.8 million, respectively, to our joint venture partners. Refer to our consolidated statements of cash flows and

Note 4 – “Consolidated and unconsolidated real estate joint ventures” to our unaudited consolidated financial statements in Item 1 for

additional information.

79

Investments

We hold investments in publicly traded companies and privately held entities primarily involved in the life science industry. The

tables below summarize components of our investment income (loss) and non-real estate investments (in thousands). Refer to Note 7 –

“Investments” to our unaudited consolidated financial statements in Item 1 for additional information.

Three Months Ended March 31, 2025 Year Ended December 31, 2024
Realized gains 18,153 59,124
Unrealized losses (68,145) (112,246)
Investment loss (49,992) (53,122)

All values are in US Dollars.

March 31, 2025
Investments Cost Unrealized<br><br>Gains Unrealized<br><br>Losses Carrying Amount
Publicly traded companies 182,797 $24,425 $(122,472) 84,750
Entities that report NAV 511,907 105,405 (42,327) 574,985
Entities that do not report NAV:
Entities with observable price changes 106,465 75,087 (8,255) 173,297
Entities without observable price changes 422,052 422,052
Investments accounted for under the equity<br><br>method N/A N/A N/A 224,604
March 31, 2025 1,223,221 $204,917 $(173,054) 1,479,688
December 31, 2024 1,207,146 $228,100 $(144,489) 1,476,985

All values are in US Dollars.

Public/Private Mix (Cost) Tenant/Non-Tenant Mix (Cost)

1

13

87%

Private

13%

Public

24%

Tenant

76%

Non-Tenant

(1)Consists of realized gains of $29.3 million, offset by impairment charges of $11.2 million during the three months ended March 31, 2025.

(2)Consists of realized gains of $117.2 million, offset by impairment charges of $58.1 million during the year ended December 31, 2024.

(3)Consists of unrealized losses of $40.0 million primarily resulting from the decrease in fair values of our investments in publicly traded entities and investments in privately

held entities that report NAV and $28.1 million resulting from accounting reclassifications of unrealized gains recognized in prior periods into realized gains upon our

realization of investments during the three months ended March 31, 2025.

(4)Primarily relates to the accounting reclassifications of unrealized gains recognized in prior periods into realized gains upon our realization of investments during the year

ended December 31, 2024.

(5)Represents 2.8% of gross assets as of March 31, 2025. Refer to “Gross assets” under “Definitions and reconciliations” in Item 2 for additional details.

80

Liquidity

Liquidity
5.3B
(In millions)
Availability under our unsecured senior line of credit, net of amounts outstanding under our commercial paper program
Cash, cash equivalents, and restricted cash
Availability under our secured construction loan
Investments in publicly traded companies
Liquidity as of March 31, 2025

All values are in US Dollars.

We expect to meet certain long-term liquidity requirements, such as requirements for development, redevelopment, other

construction projects, capital improvements, tenant improvements, property acquisitions, equity repurchases, leasing costs, non-

revenue-enhancing capital expenditures, scheduled debt maturities, distributions to noncontrolling interests, and payment of dividends

through net cash provided by operating activities, periodic asset sales, strategic real estate joint ventures, long-term secured and

unsecured indebtedness, borrowings under our unsecured senior line of credit, issuances under our commercial paper program, and

issuances of additional debt and/or equity securities.

We also expect to continue meeting our short-term liquidity and capital requirements, as further detailed in this section,

generally through our working capital and net cash provided by operating activities. We believe that the net cash provided by operating

activities will continue to be sufficient to enable us to make the distributions necessary to continue qualifying as a REIT.

For additional information on our liquidity requirements related to our contractual obligations and commitments, refer to

Note 5 – “Leases” and Note 10 – “Secured and unsecured senior debt” to our unaudited consolidated financial statements in Item 1.

Over the next several years, our balance sheet, capital structure, and liquidity objectives are as follows:

•Retain net cash flows from operating activities after payment of dividends and distributions to noncontrolling interests for

investment in development and redevelopment projects and/or acquisitions;

•Maintain significant balance sheet liquidity;

•Maintain credit profile and relative long-term cost of capital;

•Maintain diverse sources of capital, including sources from net cash provided by operating activities, unsecured debt,

secured debt, selective real estate asset sales, strategic real estate joint ventures, non-real estate investment sales, and

common stock;

•Maintain commitment to long-term capital to fund growth;

•Maintain prudent laddering of debt maturities;

•Maintain solid credit metrics;

•Prudently manage variable-rate debt exposure;

•Maintain a large, unencumbered asset pool to provide financial flexibility;

•Fund common stock dividends and distributions to noncontrolling interests from net cash provided by operating activities;

•Manage a disciplined level of development and redevelopment projects as a percentage of our gross real estate assets;

and

•Maintain high levels of pre-leasing and percentage leased in development and redevelopment projects.

81

The following table presents the availability under our unsecured senior line of credit, net of amounts outstanding under our

commercial paper program; cash, cash equivalents, and restricted cash; availability under our secured construction loan; and

investments in publicly traded companies as of March 31, 2025 (in thousands):

Description Stated Rate Aggregate<br><br>Commitments Outstanding<br><br>Balance(1) Remaining<br><br>Commitments/<br><br>Liquidity
Availability under our unsecured senior line of credit, net of<br><br>amounts outstanding under our commercial paper program SOFR+0.855% $5,000,000 $299,883 $4,700,000
Cash, cash equivalents, and restricted cash 483,754
Secured construction loan SOFR+2.70% $195,300 $150,219 44,882
Investments in publicly traded companies 84,750
Liquidity as of March 31, 2025 $5,313,386

(1)Represents outstanding principal, net of unamortized deferred financing costs, as of March 31, 2025.

Cash, cash equivalents, and restricted cash

As of March 31, 2025 and December 31, 2024, we had $483.8 million and $559.8 million, respectively, of cash, cash

equivalents, and restricted cash. We expect existing cash, cash equivalents, and restricted cash, net cash provided by operating

activities, proceeds from real estate asset sales, sales of partial interests, strategic real estate joint ventures, non-real estate investment

sales, borrowings under our unsecured senior line of credit, issuances under our commercial paper program, issuances of unsecured

senior notes payable, borrowings under our secured construction loans, and issuances of common stock to continue to be sufficient to

fund our operating activities and cash commitments for investing and financing activities, such as regular quarterly dividends,

distributions to noncontrolling interests, scheduled debt repayments, acquisitions, and certain capital expenditures, including

expenditures related to construction activities and any common stock repurchases.

Cash flows

We report and analyze our cash flows based on operating activities, investing activities, and financing activities. The following

table summarizes changes in our cash flows for the three months ended March 31, 2025 and 2024 (in thousands):

Three Months Ended March 31,
2025 2024 Change
Net cash provided by operating activities $207,949 $341,157 $(133,208)
Net cash used in investing activities $(654,779) $(894,854) $240,075
Net cash provided by financing activities $370,775 $624,429 $(253,654)

Operating activities

Cash flows provided by operating activities are primarily dependent upon the occupancy level of our asset base, the rental

rates of our leases, the collectibility of rent and recovery of operating expenses from our tenants, the timing of completion of

development and redevelopment projects, and the timing of acquisitions and dispositions of operating properties. Net cash provided by

operating activities for the three months ended March 31, 2025 decreased by $133.2 million to $207.9 million, compared to

$341.2 million for the three months ended March 31, 2024. The decrease was primarily due to the ground lease prepayment of

$135.0 million made in January 2025 for a 24-year lease term extension to our existing ground lease agreement at the Alexandria

Technology Square® Megacampus in our Cambridge submarket.

82

Investing activities

Cash used in investing activities for the three months ended March 31, 2025 and 2024 consisted of the following (in

thousands):

Three Months Ended March 31, Change
2025 2024
Sources of cash from investing activities:
Proceeds from sales of real estate $68,182 $16,670 $51,512
Sales of and distributions from non-real estate investments 12,691 40,550 (27,859)
80,873 57,220 23,653
Uses of cash for investing activities:
Purchases of real estate 194,002 (194,002)
Additions to real estate 645,841 693,268 (47,427)
Change in escrow deposits 9,506 1,008 8,498
Investments in unconsolidated real estate joint ventures 10,994 3,224 7,770
Additions to non-real estate investments 69,311 60,572 8,739
735,652 952,074 (216,422)
Net cash used in investing activities $654,779 $894,854 $(240,075)

The decrease in net cash used in investing activities for the three months ended March 31, 2025, compared to the three

months ended March 31, 2024, was primarily due to a decreased use of cash for purchases of real estate. Refer to Note 3 –

“Investments in real estate” to our unaudited consolidated financial statements in Item 1 for additional information.

Financing activities

Cash flows provided by financing activities for the three months ended March 31, 2025 and 2024 consisted of the following

(in thousands):

Three Months Ended March 31,
2025 2024 Change
Borrowings under secured notes payable $824 $10,216 $(9,392)
Proceeds from issuance of unsecured senior notes payable 548,532 998,806 (450,274)
Proceeds from issuances under commercial paper program 2,700,000 3,170,000 (470,000)
Repayments of borrowings under commercial paper program (2,400,000) (3,270,000) 870,000
Payments of loan fees (5,406) (10,118) 4,712
Changes related to debt 843,950 898,904 (54,954)
Contributions from and sales of noncontrolling interests 54,409 82,853 (28,444)
Distributions to and purchases of noncontrolling interests (83,852) (111,540) 27,688
Repurchase of common stock (208,187) (208,187)
Dividends on common stock (229,987) (221,824) (8,163)
Taxes paid related to net settlement of equity awards (5,558) (23,964) 18,406
Net cash provided by financing activities $370,775 $624,429 $(253,654)

83

Capital resources

We expect that our principal liquidity needs for the year ending December 31, 2025 will be satisfied by the following multiple

sources of capital, as shown in the table below. There can be no assurance that our sources and uses of capital will not be materially

higher or lower than these expectations.

Key Sources and Uses of Capital<br><br>(In millions) 2025 Guidance As of 1/27/25 Midpoint
Range Midpoint
Sources of capital:
Net reduction in debt $(290) $(290) (290) (190)
Net cash provided by operating activities after dividends(1) 425 525 475 475
Dispositions and sales of partial interests 1,450 2,450 1,950 1,700
Total sources of capital $1,585 $2,685 2,135 1,985
Uses of capital:
Construction $1,450 $2,050 1,750 1,750
Acquisitions and other opportunistic uses of capital 500 250 100
Ground lease prepayment 135 135 135 135
Total uses of capital $1,585 $2,685 2,135 1,985
Net reduction in debt (included above):
Issuance of unsecured senior notes payable $550 $550 550 600
Repayment of unsecured notes payable(5) (600) (600) (600) (600)
Unsecured senior line of credit, commercial paper<br><br>program, and other (240) (240) (240) (190)
Net reduction in debt $(290) $(290) (290) (190)

All values are in US Dollars.

(1)Excludes the final installment payment of $135.0 million made in January 2025 for our ground lease at the Alexandria Technology Square® Megacampus. This amount

has been separately presented as “Ground lease prepayment” under “Uses of capital” in the table above.

(2)As of the date of this report, completed dispositions aggregated $176.4 million and our share of pending transactions subject to non-refundable deposits, signed letters

of intent, or purchase and sale agreement negotiations aggregated $432.5 million. As part of a completed transaction, we provided seller financing of $91.0 million. Refer

to “Dispositions and sales of partial interests” in Item 2 for additional information on our real estate dispositions.

(3)The increase to the midpoint of our guidance range for 2025 dispositions and sales of partial interests is primarily due to an increase in the midpoint of our guidance

range for 2025 acquisitions and other opportunistic uses of capital by $150 million.

(4)Under our common stock repurchase program authorized in December 2024, we may repurchase up to $500.0 million of our common stock through December 31,

  1. During the three months ended March 31, 2025, we repurchased 2.2 million shares of common stock for an aggregate value of $208.1 million at an average price

per share of $96.71. As of the date of this report, the approximate value of shares authorized and remaining under this program was $241.8 million. Subject to market

conditions, we may consider repurchasing additional shares of our common stock.

(5)Upon maturity on April 30, 2025, we expect to repay $600.0 million of our 3.45% unsecured senior notes payable.

The key assumptions behind the sources and uses of capital in the table above include a favorable real estate transaction and

capital market environments, performance of our core operating properties, lease-up and delivery of current and future development

and redevelopment projects, and leasing activity. Our expected sources and uses of capital are subject to a number of variables and

uncertainties, including those discussed as “Forward-looking statements” under Part I; “Item 1A. Risk factors”; and “Item 7.

Management’s discussion and analysis of financial condition and results of operations” of our annual report on Form 10-K for the year

ended December 31, 2024; as well as in “Item 1A. Risk factors” within “Part II – Other information” of this quarterly report on Form 10-

Q. We expect to update our forecast for key sources and uses of capital on a quarterly basis.

84

Sources of capital

Net cash provided by operating activities after dividends

We expect to retain $425 million to $525 million of net cash flows from operating activities after payment of common stock

dividends, and distributions to noncontrolling interests for the year ending December 31, 2025, excluding the payment of our final

installment of $135.0 million made in January 2025 for the ground lease at the Alexandria Technology Square® Megacampus. For

purposes of this calculation, changes in operating assets and liabilities representing timing differences are excluded. For the year

ending December 31, 2025, we expect our recently delivered projects, our development and redevelopment projects expected to be

delivered, contributions from Same Properties, and recently acquired income-producing properties to contribute to income from rentals,

net operating income, and cash flows. We anticipate contractual near-term growth in annual net operating income (cash basis) of

$61 million related to the commencement of contractual rents on the projects recently placed into service that are near the end of their

initial free rent period. Refer to “Cash flows” in Item 2 for a discussion of cash flows provided by operating activities for the three months

ended March 31, 2025.

Debt

We expect to fund a portion of our capital needs for 2025 from issuances under our commercial paper program, issuances of

unsecured senior notes payable, borrowings under our unsecured senior line of credit, and/or borrowings under our secured

construction loan.

As of March 31, 2025, our unsecured senior line of credit, which matures in 2030, including extension options under our

control, had aggregate commitments of $5.0 billion and bore an interest rate of SOFR plus 0.855%. In addition to the cost of borrowing,

the unsecured senior line of credit is subject to an annual facility fee of 0.145% based on the aggregate commitments outstanding.

Based upon our ability to achieve certain annual sustainability targets, the interest rate and facility fee rate are also subject to upward or

downward adjustments of up to four basis points with respect to the interest rate and up to one basis point with respect to the facility fee

rate.

Based on certain sustainability metrics achieved in accordance with the terms of our unsecured senior line of credit

agreement, the borrowing rate was reduced for a one-year period by two basis points to SOFR plus 0.855%, from SOFR plus 0.875%,

and the facility fee was reduced by 0.5 basis point to 0.145% from 0.15%. As of March 31, 2025, we had no outstanding balance on our

unsecured line of credit.

Our commercial paper program provides us with the ability to issue up to $2.5 billion of commercial paper notes with a maturity

of generally 30 days or less and with a maximum maturity of 397 days from the date of issuance. Our commercial paper program is

backed by our unsecured senior line of credit, and at all times we expect to retain a minimum undrawn amount of borrowing capacity

under our unsecured senior line of credit equal to any outstanding balance under our commercial paper program. We use borrowings

under the program to fund short-term capital needs. The notes issued under our commercial paper program are sold under customary

terms in the commercial paper market. They are typically issued at a discount to par, representing a yield to maturity dictated by market

conditions at the time of issuance. In the event we are unable to issue commercial paper notes or refinance outstanding commercial

paper notes under terms equal to or more favorable than those under the unsecured senior line of credit, we expect to borrow under the

unsecured senior line of credit. The commercial paper notes sold during the three months ended March 31, 2025 were issued at a

weighted-average yield to maturity of 4.60%. As of March 31, 2025, we had $299.9 million of commercial paper notes outstanding.

In February 2025, we issued $550.0 million of unsecured senior notes payable, due 2035, with an interest rate of 5.50%.

The following table presents our average debt outstanding and weighted-average interest rates during the three months ended

March 31, 2025 (dollars in thousands):

Three Months Ended March 31, 2025
Average Debt<br><br>Outstanding Weighted-Average<br><br>Interest Rate
Long-term fixed-rate debt $12,434,676 3.83%
Short-term variable-rate unsecured senior line of credit and commercial paper program<br><br>debt 375,884 4.59
Blended average interest rate 12,810,560 3.85
Loan fee amortization and annual facility fee related to unsecured senior line of credit N/A 0.14
Total/weighted average $12,810,560 3.99%

85

Real estate dispositions and sales of partial interests

We expect to continue to focus on the disciplined execution of select sales of real estate. Future sales will provide an important

source of capital to fund our development and redevelopment projects and opportunistic share repurchases and also provide significant

capital for growth. We may also consider additional sales of partial interests in core Class A/A+ properties, development projects, and/or

land. For the year ending December 31, 2025, we expect real estate dispositions and sales of partial interests in real estate assets to

range from $1.45 billion to $2.45 billion. The amount of asset sales necessary to meet our forecasted sources of capital will vary

depending upon the amount of EBITDA associated with the assets sold.

Refer to Note 3 – “Investments in real estate,” Note 4 – “Consolidated and unconsolidated real estate joint ventures,” and

Note 13 – “Stockholders’ equity” to our unaudited consolidated financial statements in Item 1 and to “Dispositions and sales of partial

interests” in Item 2 for additional information on our real estate dispositions.

As a REIT, we are generally subject to a 100% tax on the net income from real estate asset sales that the IRS characterizes as

“prohibited transactions.” We do not expect our sales will be categorized as prohibited transactions. However, unless we meet certain

“safe harbor” requirements, whether a real estate asset sale is a “prohibited transaction” will be based on the facts and circumstances

of the sale. Our real estate asset sales may not always meet such “safe harbor” requirements. Refer to “Item 1A. Risk factors” of our

annual report on Form 10-K for the year ended December 31, 2024 for additional information about the “prohibited transaction” tax.

Common equity transactions

As of March 31, 2025, the remaining aggregate amount available under our ATM program for future sales of common stock

was $1.47 billion.

Other sources

As a well-known seasoned issuer, we may, from time to time, issue securities at our discretion based on our needs and market

conditions, including, as necessary, to balance our use of incremental debt capital.

Additionally, we, together with joint venture partners, hold interests in real estate joint ventures that we consolidate in our

financial statements. These existing joint ventures provide significant equity capital to fund a portion of our future construction spend,

and our joint venture partners may also contribute equity into these entities for financing-related activities. From April 1, 2025 through

December 31, 2027 and beyond, we expect to receive capital contributions aggregating $414.9 million from existing consolidated real

estate joint venture partners to fund construction. During the year ending December 31, 2025, contributions from noncontrolling

interests from existing joint venture partners are expected to aggregate $230.0 million.

86

Uses of capital

Summary of capital expenditures

One of our primary uses of capital relates to the development, redevelopment, pre-construction, and construction of properties.

We currently have projects in our development and redevelopment pipeline aggregating 4.0 million RSF of Class A/A+ properties

undergoing construction. We incur capitalized construction costs related to development, redevelopment, pre-construction, and other

construction activities. We also incur additional capitalized project costs, including interest, property taxes, insurance, and other costs

directly related and essential to the development, redevelopment, pre-construction, or construction of a project, during periods when

activities necessary to prepare an asset for its intended use are in progress. Refer to “New Class A/A+ development and redevelopment

properties: current projects” and “Summary of capital expenditures” in Item 2 for additional information on our capital expenditures.

We capitalize interest cost as a cost of the project only during the period in which activities necessary to prepare an asset for

its intended use are ongoing, provided that expenditures for the asset have been made and interest cost has been incurred. Capitalized

interest for the three months ended March 31, 2025 and 2024 of $80.1 million and $81.8 million, respectively, was classified in

investments in real estate in our consolidated balance sheets. The decrease in capitalized interest was related to a lower weighted-

average capitalized cost basis of $8.0 billion for the three months ended March 31, 2025, as compared to $8.2 billion for the three

months ended March 31, 2024, partially offset by an increase in weighted-average interest rate used to capitalize interest to 3.99% for

the three months ended March 31, 2025 from 3.92% for the three months ended March 31, 2024.

Property taxes, insurance on real estate, and indirect project costs, such as construction, administration, legal fees, and office

costs that clearly relate to projects under development or construction, are capitalized as incurred during the period an asset is

undergoing activities to prepare it for its intended use. We capitalized payroll and other indirect costs related to development,

redevelopment, pre-construction, and construction projects, aggregating $24.8 million and $26.3 million, and property taxes, insurance

on real estate, and indirect project costs aggregating $36.2 million and $32.7 million during the three months ended March 31, 2025 and

2024, respectively.

Pre-construction activities include entitlements, permitting, design, site work, and other activities preceding commencement of

construction of aboveground building improvements. The advancement of pre-construction efforts is focused on reducing the time

required to deliver projects to prospective tenants. These critical activities add significant value for future ground-up development and

are required for the vertical construction of buildings. Should we cease activities necessary to prepare an asset for its intended use, the

interest, taxes, insurance, and certain other direct and indirect project costs related to the asset would be expensed as incurred.

Expenditures for repairs and maintenance are expensed as incurred.

Fluctuations in our development, redevelopment, and construction activities could result in significant changes to total

expenses and net income. For example, had we experienced a 10% reduction in development, redevelopment, and construction

activities without a corresponding decrease in indirect project costs, including interest and payroll, total expenses would have increased

by approximately $14.1 million for the three months ended March 31, 2025.

We use third-party brokers to assist in our leasing activity, who are paid on a contingent basis upon successful leasing. We are

required to capitalize initial direct costs related to successful leasing transactions that result directly from and are essential to the lease

transaction and would not have been incurred had that lease transaction not been successfully executed. During the three months

ended March 31, 2025, we capitalized total initial direct leasing costs of $27.4 million. Costs that we incur to negotiate or arrange a

lease regardless of its outcome, such as fixed employee compensation, tax, or legal advice to negotiate lease terms, and other costs,

are expensed as incurred.

Real estate acquisitions and common stock repurchase program

Under our common stock repurchase program authorized in December 2024, we may repurchase up to $500.0 million of our

common stock in the open market, in privately negotiated transactions, or otherwise through December 31, 2025.

•During the three months ended March 31, 2025, we repurchased 2.2 million shares of common stock for an aggregate value of

$208.1 million at an average price per share of $96.71.

•As of the date of this report, the approximate value of shares authorized and remaining under this program was $241.8 million.

For the year ending December 31, 2025, we expect real estate acquisitions and common stock repurchases to range from

$0 to $500 million. We completed no acquisitions during the three months ended March 31, 2025.

87

Dividends

During the three months ended March 31, 2025 and 2024, we paid common stock dividends of $230.0 million and

$221.8 million, respectively. The increase of $8.2 million in dividends paid on our common stock during the three months ended

March 31, 2025, compared to the three months ended March 31, 2024, was primarily due to an increase in the related dividends to

$1.32 per common share paid during the three months ended March 31, 2025 from $1.27 per common share paid during the three

months ended March 31, 2024.

Secured notes payable

Secured notes payable as of March 31, 2025 consisted of three notes secured by two properties. Our secured notes payable

typically require monthly payments of principal and interest and had a weighted-average interest rate of approximately 7.20%. As of

March 31, 2025, the total book value of our investments in real estate securing debt was approximately $374.1 million. As of March 31,

2025, our secured notes payable, including unamortized discounts and deferred financing costs, comprised approximately

$588 thousand and $150.2 million of fixed-rate debt and unhedged variable-rate debt, respectively.

During the three months ended March 31, 2025, our unconsolidated real estate joint venture in which we hold a 10%

ownership interest, located at 1655 and 1725 Third Street in our Mission Bay submarket, refinanced $500 million of an existing fixed-

rate debt with a new secured note payable, which bears a weighted-average interest rate of 6.37% and matures in 2035. The remaining

debt balance of approximately $100 million of the previous $600 million debt was repaid through contributions from the unconsolidated

joint venture partners, including our share of $10.8 million.

Unsecured senior notes payable and unsecured senior line of credit

The requirements of, and our actual performance with respect to, the key financial covenants under our unsecured senior

notes payable as of March 31, 2025 were as follows:

Covenant Ratios(1) Requirement March 31, 2025
Total Debt to Total Assets Less than or equal to 60% 31%
Secured Debt to Total Assets Less than or equal to 40% 0.4%
Consolidated EBITDA(2) to Interest Expense Greater than or equal to 1.5x 10.2x
Unencumbered Total Asset Value to Unsecured Debt Greater than or equal to 150% 311%

(1)All covenant ratio titles utilize terms as defined in the respective debt agreements.

(2)The calculation of consolidated EBITDA is based on the definitions contained in our loan agreements and is not directly comparable to the computation of EBITDA as

described in Exchange Act Release No. 47226.

In addition, the terms of the indentures, among other things, limit the ability of the Company, Alexandria Real Estate Equities,

L.P., and the Company’s subsidiaries to (i) consummate a merger, or consolidate, or sell all or substantially all of the Company’s assets

and (ii) incur certain secured or unsecured indebtedness.

The requirements of, and our actual performance with respect to, the key financial covenants under our unsecured senior line

of credit as of March 31, 2025 were as follows:

Covenant Ratios(1) Requirement March 31, 2025
Leverage Ratio Less than or equal to 60.0% 31.7%
Secured Debt Ratio Less than or equal to 45.0% 0.3%
Fixed-Charge Coverage Ratio Greater than or equal to 1.50x 3.83x
Unsecured Interest Coverage Ratio Greater than or equal to 1.75x 9.76x

(1)All covenant ratio titles utilize terms as defined in the credit agreement.

Estimated interest payments

Estimated interest payments on our fixed-rate debt are calculated based upon contractual interest rates, including interest

payment dates and scheduled maturity dates. As of March 31, 2025, 96.6% of our debt was fixed-rate debt. For additional information

regarding our debt, refer to Note 10 – “Secured and unsecured senior debt” to our unaudited consolidated financial statements in

Item 1.

88

Ground lease obligations

Ground lease obligations as of March 31, 2025 included leases for 32 of our properties and accounted for approximately 8% of

our total number of properties. Among these 32 properties, 17 properties are subject to ground leases with a weighted-average

remaining lease term of 41 years, including extension options that we are reasonably certain to exercise. These leases are with a single

lessor in our Greater Stanford submarket with whom we have extended three ground leases over the past 10 years.

Our remaining 15 properties subject to ground leases are located across multiple submarkets and have remaining lease terms

ranging from approximately 46 to 82 years. The weighted-average remaining lease term of these ground leases is 73 years, including

extension options that we are reasonably certain to exercise.

In many cases, we seek to extend our ground leases well ahead of their scheduled contractual expirations. If we are

successful in extending ground leases, we could see significant up-front or increased recurring future payments to the ground lessor

and/or increased ground lease expense, which may require us to increase our capital funding needs.

Operating lease agreements

As of March 31, 2025, the remaining contractual payments under ground and office lease agreements in which we are the

lessee aggregated $785.4 million and $23.6 million, respectively. As of March 31, 2025, our operating lease liability, calculated as the

present value of the remaining payments aggregating $809.0 million under our operating lease agreements, including our extension

options that we are reasonably certain to exercise, was $371.4 million, which was classified in accounts payable, accrued expenses,

and other liabilities in our consolidated balance sheet. As of March 31, 2025, the weighted-average remaining lease term of operating

leases in which we are the lessee was approximately 54 years, including extension options that we are reasonably certain to exercise,

and the weighted-average discount rate was 4.7%. Our corresponding operating lease right-of-use assets, adjusted for initial direct

leasing costs and other consideration exchanged with the landlord prior to the commencement of the lease, aggregated $728.9 million.

We classify the right-of-use asset in other assets in our consolidated balance sheets. Refer to “Lease accounting” in Note 2 –

“Summary of significant accounting policies” to our unaudited consolidated financial statements in Item 1 for additional information.

In July 2024, we entered into an amendment to our existing ground lease agreement at the Alexandria Technology Square®

Megacampus aggregating 1.2 million RSF in our Cambridge submarket, which extended the lease term by 24 years from January 1,

2065 to December 31, 2088. The amendment required that we prepay our entire rent obligation for the extended lease term

aggregating $270.0 million in two equal installments in December 2024 and in January 2025. On January 14, 2025, we made the

second and final installment payment of $135.0 million.

Commitments

As of March 31, 2025, remaining aggregate costs under contract for the construction of properties undergoing development,

redevelopment, and improvements under the terms of leases approximated $1.0 billion. We expect payments for these obligations to

occur over one to three years, subject to capital planning adjustments from time to time. We may have the ability to cease the

construction of certain projects, which would result in the reduction of our commitments. In addition, we have letters of credit and

performance obligations aggregating $5.3 million.

We are committed to funding approximately $386.5 million related to our non-real estate investments. These funding

commitments are primarily associated with our investments in privately held entities that report NAV and expire at various dates over

the next 12 years, with a weighted-average expiration of 8.1 years as of March 31, 2025.

Our former joint venture partner in our Greater Boston market has an option, subject to certain conditions, to obtain a

$50 million secured loan from us, which, if the option is exercised, will bear interest at SOFR plus 6.50%, with a floor of 9.0% and a

term not to exceed five years. As of March 31, 2025, the option has not been exercised.

Exposure to environmental liabilities

In connection with the acquisition of all of our properties, we have obtained Phase I environmental assessments to ascertain

the existence of any environmental liabilities or other issues. The Phase I environmental assessments of our properties have not

revealed any environmental liabilities that we believe would have a material adverse effect on our financial condition or results of

operations taken as a whole, nor are we aware of any material environmental liabilities that have occurred since the Phase I

environmental assessments were completed. In addition, we carry a policy of pollution legal liability insurance covering exposure to

certain environmental losses at substantially all of our properties.

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Foreign currency translation gains and losses

The following table presents the change in accumulated other comprehensive loss attributable to Alexandria Real Estate

Equities, Inc.’s stockholders during the three months ended March 31, 2025 primarily due to the changes in the foreign exchange rates

for our real estate investments in Canada (in thousands). We reclassify unrealized foreign currency translation gains and losses into net

income as we dispose of these holdings.

Total
Balance as of December 31, 2024 $(46,252)
Other comprehensive income before reclassifications 50
Net other comprehensive income 50
Balance as of March 31, 2025 $(46,202)

Inflation

As of March 31, 2025, approximately 91% of our leases (on an annual rental revenue basis) were triple net leases, which

require tenants to pay substantially all real estate taxes, insurance, utilities, repairs and maintenance, common area expenses, and

other operating expenses (including increases thereto) in addition to base rent. Approximately 98% of our leases (on an annual rental

revenue basis) contained effective annual rent escalations approximating 3% that were either fixed or indexed based on a consumer

price index or other indices. Accordingly, we do not believe that our cash flows or earnings from real estate operations are subject to

significant risks from inflation. A period of inflation, however, could cause an increase in the cost of our variable-rate borrowings,

including borrowings under our unsecured senior line of credit and commercial paper program, issuances of unsecured senior notes

payable, and borrowings under our secured construction loans, and secured loans held by our unconsolidated real estate joint ventures.

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Issuer and guarantor subsidiary summarized financial information

Alexandria Real Estate Equities, Inc. (the “Issuer”) has sold certain debt securities registered under the Securities Act of 1933,

as amended, that are fully and unconditionally guaranteed by Alexandria Real Estate Equities, L.P. (the “LP” or the “Guarantor

Subsidiary”), an indirectly 100% owned subsidiary of the Issuer. The Issuer’s other subsidiaries, including, but not limited to, the

subsidiaries that own substantially all of its real estate (collectively, the “Combined Non-Guarantor Subsidiaries”), will not provide a

guarantee of such securities, including the subsidiaries that are partially or 100% owned by the LP. The following summarized financial

information presents, on a combined basis, balance sheet information as of March 31, 2025 and December 31, 2024, and results of

operations and comprehensive income for the three months ended March 31, 2025 and year ended December 31, 2024 for the Issuer

and the Guarantor Subsidiary. The information presented below excludes eliminations necessary to arrive at the information on a

consolidated basis. In presenting the summarized financial statements, the equity method of accounting has been applied to (i) the

Issuer’s interests in the Guarantor Subsidiary, (ii) the Guarantor Subsidiary’s interests in the Combined Non-Guarantor Subsidiaries,

and (iii) the Combined Non-Guarantor Subsidiaries’ interests in the Guarantor Subsidiary, where applicable, even though all such

subsidiaries meet the requirements to be consolidated under GAAP. All assets and liabilities have been allocated to the Issuer and the

Guarantor Subsidiary generally based on legal entity ownership.

The following tables present combined summarized financial information as of March 31, 2025 and December 31, 2024 and for

the three months ended March 31, 2025 and year ended December 31, 2024 for the Issuer and Guarantor Subsidiary. Amounts

provided do not represent our total consolidated amounts (in thousands):

March 31, 2025 December 31, 2024
Assets:
Cash, cash equivalents, and restricted cash $99,711 $103,993
Other assets 158,773 153,913
Total assets $258,484 $257,906
Liabilities:
Unsecured senior notes payable $12,640,144 $12,094,465
Unsecured senior line of credit and commercial paper 299,883
Other liabilities 519,595 542,322
Total liabilities $13,459,622 $12,636,787 Three Months Ended<br><br>March 31, 2025 Year Ended<br><br>December 31, 2024
--- --- ---
Total revenues $9,530 $59,023
Total expenses (81,459) (349,437)
Net loss (71,929) (290,414)
Net income attributable to unvested restricted stock awards (2,660) (13,394)
Net loss attributable to Alexandria Real Estate Equities, Inc.’s common stockholders $(74,589) $(303,808)

As of March 31, 2025, 371 of our 386 properties were held indirectly by the REIT’s wholly owned consolidated subsidiary,

Alexandria Real Estate Equities, L.P.

Critical accounting estimates

Refer to our annual report on Form 10-K for the year ended December 31, 2024 for a discussion of our critical accounting

estimates related to recognition of real estate acquired, impairment of long-lived assets, impairment of non-real estate investments, and

monitoring of tenant credit quality.

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Definitions and reconciliations

This section contains additional information on certain non-GAAP financial measures, including reconciliations to the most

directly comparable financial measure calculated and presented in accordance with GAAP and the reasons why we use these

supplemental measures of performance and believe they provide useful information to investors, as well as the definitions of other

terms used in this report.

Funds from operations and funds from operations, as adjusted, attributable to Alexandria Real Estate Equities, Inc.’s common

stockholders

GAAP-basis accounting for real estate assets utilizes historical cost accounting and assumes that real estate values diminish

over time. In an effort to overcome the difference between real estate values and historical cost accounting for real estate assets, the

Nareit Board of Governors established funds from operations as an improved measurement tool. Since its introduction, funds from

operations has become a widely used non-GAAP financial measure among equity REITs. We believe that funds from operations is

helpful to investors as an additional measure of the performance of an equity REIT. Moreover, we believe that funds from operations, as

adjusted, allows investors to compare our performance to the performance of other real estate companies on a consistent basis, without

having to account for differences recognized because of real estate acquisition and disposition decisions, financing decisions, capital

structure, capital market transactions, variances resulting from the volatility of market conditions outside of our control, or other

corporate activities that may not be representative of the operating performance of our properties.

The 2018 White Paper published by the Nareit Board of Governors (the “Nareit White Paper”) defines funds from operations as

net income (computed in accordance with GAAP), excluding gains or losses on sales of real estate, and impairments of real estate, plus

depreciation and amortization of operating real estate assets, and after adjustments for our share of consolidated and unconsolidated

partnerships and real estate joint ventures. Impairments represent the write-down of assets when fair value over the recoverability

period is less than the carrying value due to changes in general market conditions and do not necessarily reflect the operating

performance of the properties during the corresponding period.

We compute funds from operations, as adjusted, as funds from operations calculated in accordance with the Nareit White

Paper, excluding significant gains, losses, and impairments realized on non-real estate investments, unrealized gains or losses on non-

real estate investments, impairments of real estate primarily consisting of right-of-use-assets and pre-acquisition costs related to

projects that we decided to no longer pursue, gains or losses on early extinguishment of debt, changes in the provision for expected

credit losses on financial instruments, significant termination fees, acceleration of stock compensation expense due to the resignations

of executive officers, deal costs, the income tax effect related to such items, and the amount of such items that is allocable to our

unvested restricted stock awards. We compute the amount that is allocable to our unvested restricted stock awards using the two-class

method. Under the two-class method, we allocate net income (after amounts attributable to noncontrolling interests) to common

stockholders and to unvested restricted stock awards by applying the respective weighted-average shares outstanding during each

quarter-to-date and year-to-date period. This may result in a difference of the summation of the quarter-to-date and year-to-date

amounts. Neither funds from operations nor funds from operations, as adjusted, should be considered as alternatives to net income

(determined in accordance with GAAP) as indications of financial performance, or to cash flows from operating activities (determined in

accordance with GAAP) as measures of liquidity, nor are they indicative of the availability of funds for our cash needs, including our

ability to make distributions.

The following table reconciles net income (loss) to funds from operations for the share of consolidated real estate joint

ventures attributable to noncontrolling interests and our share of unconsolidated real estate joint ventures for the three months ended

March 31, 2025 (in thousands):

Three Months Ended March 31, 2025
Noncontrolling<br><br>Interest Share of<br><br>Consolidated Real<br><br>Estate Joint Ventures Our Share of<br><br>Unconsolidated<br><br>Real Estate Joint<br><br>Ventures
Net income (loss) $47,601 $(507)
Depreciation and amortization of real estate assets 33,411 1,054
Funds from operations $81,012 $547

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The following tables present a reconciliation of net income attributable to Alexandria Real Estate Equities, Inc.’s common

stockholders, the most directly comparable financial measure presented in accordance with GAAP, including our share of amounts from

consolidated and unconsolidated real estate joint ventures, to funds from operations attributable to Alexandria Real Estate Equities,

Inc.’s common stockholders – diluted, and funds from operations attributable to Alexandria Real Estate Equities, Inc.’s common

stockholders – diluted, as adjusted, and the related per share amounts for the three months ended March 31, 2025 and 2024 (in

thousands, except per share amounts). Per share amounts may not add due to rounding.

Three Months Ended March 31,
2025 2024
Net (loss) income attributable to Alexandria Real Estate Equities, Inc.’s common stockholders –<br><br>basic and diluted (11,599) $166,886
Depreciation and amortization of real estate assets 339,381 284,950
Noncontrolling share of depreciation and amortization from consolidated real estate JVs (33,411) (30,904)
Our share of depreciation and amortization from unconsolidated real estate JVs 1,054 1,034
Gain on sales of real estate (13,165) (392)
Allocation to unvested restricted stock awards (686) (3,469)
Funds from operations attributable to Alexandria Real Estate Equities, Inc.’s common stockholders<br><br>– diluted(1) 281,574 418,105
Unrealized losses (gains) on non-real estate investments 68,145 (29,158)
Impairment of non-real estate investments 11,180 14,698
Impairment of real estate 32,154
Increase in provision for expected credit losses on financial instruments 285
Allocation to unvested restricted stock awards (1,329) 247
Funds from operations attributable to Alexandria Real Estate Equities, Inc.’s common stockholders<br><br>– diluted, as adjusted 392,009 $403,892

All values are in US Dollars.

(1)Calculated in accordance with standards established by the Nareit Board of Governors.

(2)Primarily related to four non-real estate investments in privately held entities that do not report NAV.

(3)Refer to Note 5 – “Leases” to our unaudited consolidated financial statements for additional information.

Three Months Ended March 31,
(Per share) 2025 2024
Net (loss) income per share attributable to Alexandria Real Estate Equities, Inc.’s common<br><br>stockholders – diluted $(0.07) $0.97
Depreciation and amortization of real estate assets 1.80 1.48
Gain on sales of real estate (0.08)
Allocation to unvested restricted stock awards (0.02)
Funds from operations per share attributable to Alexandria Real Estate Equities, Inc.’s common<br><br>stockholders – diluted 1.65 2.43
Unrealized losses (gains) on non-real estate investments 0.40 (0.17)
Impairment of non-real estate investments 0.07 0.09
Impairment of real estate 0.19
Allocation to unvested restricted stock awards (0.01)
Funds from operations per share attributable to Alexandria Real Estate Equities, Inc.’s common<br><br>stockholders – diluted, as adjusted $2.30 $2.35
Weighted-average shares of common stock outstanding – diluted(1)
Earnings per share – diluted 170,522 171,949
Funds from operations – diluted, per share 170,599 171,949
Funds from operations – diluted, as adjusted, per share 170,599 171,949

(1)Refer to “Weighted-average shares of common stock outstanding – diluted” in this section for additional information.

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Adjusted EBITDA and Adjusted EBITDA margin

We use Adjusted EBITDA as a supplemental performance measure of our operations, for financial and operational decision-

making, and as a supplemental means of evaluating period-to-period comparisons on a consistent basis. Adjusted EBITDA is calculated

as earnings before interest, taxes, depreciation, and amortization (“EBITDA”), excluding stock compensation expense, gains or losses

on early extinguishment of debt, gains or losses on sales of real estate, impairments of real estate, changes in the provision for

expected credit losses on financial instruments, and significant termination fees. Adjusted EBITDA also excludes unrealized gains or

losses and significant realized gains or losses and impairments that result from our non-real estate investments. These non-real estate

investment amounts are classified in our consolidated statements of operations outside of total revenues.

We believe Adjusted EBITDA provides investors with relevant and useful information as it allows investors to evaluate the

operating performance of our business activities without having to account for differences recognized because of investing and

financing decisions related to our real estate and non-real estate investments, our capital structure, capital market transactions, and

variances resulting from the volatility of market conditions outside of our control. For example, we exclude gains or losses on the early

extinguishment of debt to allow investors to measure our performance independent of our indebtedness and capital structure. We

believe that adjusting for the effects of impairments and gains or losses on sales of real estate, significant impairments and realized

gains or losses on non-real estate investments, changes in the provision for expected credit losses on financial instruments, and

significant termination fees allows investors to evaluate performance from period to period on a consistent basis without having to

account for differences recognized because of investing and financing decisions related to our real estate and non-real estate

investments or other corporate activities that may not be representative of the operating performance of our properties.

In addition, we believe that excluding charges related to stock compensation and unrealized gains or losses facilitates for

investors a comparison of our business activities across periods without the volatility resulting from market forces outside of our control.

Adjusted EBITDA has limitations as a measure of our performance. Adjusted EBITDA does not reflect our historical expenditures or

future requirements for capital expenditures or contractual commitments. While Adjusted EBITDA is a relevant measure of performance,

it does not represent net income (loss) or cash flows from operations calculated and presented in accordance with GAAP, and it should

not be considered as an alternative to those indicators in evaluating performance or liquidity.

In order to calculate the Adjusted EBITDA margin, we divide Adjusted EBITDA by total revenues as presented in our

consolidated statements of operations. We believe that this supplemental performance measure provides investors with additional

useful information regarding the profitability of our operating activities.

We are not able to forecast the net income of future periods without unreasonable effort and therefore do not provide a

reconciliation for Adjusted EBITDA on a forward-looking basis. This is due to the inherent difficulty of forecasting the timing and/or

amount of items that depend on market conditions outside of our control, including the timing of dispositions, capital events, and

financing decisions, as well as quarterly components such as gain on sales of real estate, unrealized gains or losses on non-real estate

investments, impairment of real estate, impairment of non-real estate investments, and changes in the provision for expected credit

losses on financial instruments. Our attempt to predict these amounts may produce significant but inaccurate estimates, which would be

potentially misleading for our investors.

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The following table reconciles net income, the most directly comparable financial measure calculated and presented in

accordance with GAAP, to Adjusted EBITDA and calculates the Adjusted EBITDA margin for the three months ended March 31, 2025

and 2024 (dollars in thousands):

Three Months Ended March 31,
2025 2024
Net income $38,662 $219,176
Interest expense 50,876 40,840
Income taxes 1,145 1,764
Depreciation and amortization 342,062 287,554
Stock compensation expense 10,064 17,125
Gain on sales of real estate (13,165) (392)
Unrealized losses (gains) on non-real estate investments 68,145 (29,158)
Impairment of real estate 32,154
Impairment of non-real estate investments 11,180 14,698
Increase in provision for expected credit losses on financial instruments 285
Adjusted EBITDA $541,408 $551,607
Total revenues $758,158 $769,108
Adjusted EBITDA margin 71% 72%

Annual rental revenue

Annual rental revenue represents the annualized fixed base rental obligations, calculated in accordance with GAAP, including

the amortization of deferred revenue related to tenant-funded and tenant-built landlord improvements, for leases in effect as of the end

of the period, related to our operating RSF. Annual rental revenue is presented using 100% of the annual rental revenue from our

consolidated properties and our share of annual rental revenue for our unconsolidated real estate joint ventures. Annual rental revenue

per RSF is computed by dividing annual rental revenue by the sum of 100% of the RSF of our consolidated properties and our share of

the RSF of properties held in unconsolidated real estate joint ventures. As of March 31, 2025, approximately 91% of our leases (on an

annual rental revenue basis) were triple net leases, which require tenants to pay substantially all real estate taxes, insurance, utilities,

repairs and maintenance, common area expenses, and other operating expenses (including increases thereto) in addition to base rent.

Annual rental revenue excludes these operating expenses recovered from our tenants. Amounts recovered from our tenants related to

these operating expenses, along with base rent, are classified in income from rentals in our consolidated statements of operations.

Capitalization rates

Capitalization rates are calculated based on net operating income and net operating income (cash basis) annualized,

excluding lease termination fees, on stabilized operating assets for the quarter preceding the date on which the property is sold, or

near-term prospective net operating income.

Capitalized interest

We capitalize interest cost as a cost of a project during periods for which activities necessary to develop, redevelop, or

reposition a project for its intended use are ongoing, provided that expenditures for the asset have been made and interest cost has

been incurred. Activities necessary to develop, redevelop, or reposition a project include pre-construction activities such as

entitlements, permitting, design, site work, and other activities preceding commencement of construction of aboveground building

improvements. The advancement of pre-construction efforts is focused on reducing the time required to deliver projects to prospective

tenants. These critical activities add significant value for future ground-up development and are required for the vertical construction of

buildings. If we cease activities necessary to prepare a project for its intended use, interest costs related to such project are expensed

as incurred.

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Cash interest

Cash interest is equal to interest expense calculated in accordance with GAAP plus capitalized interest, less amortization of

loan fees and debt premiums (discounts). Refer to “Fixed-charge coverage ratio” in this section for a reconciliation of interest expense,

the most directly comparable financial measure calculated and presented in accordance with GAAP, to cash interest.

Class A/A+ properties and AAA locations

Class A/A+ properties are properties clustered in AAA locations that provide innovative tenants with highly dynamic and

collaborative environments that enhance their ability to successfully recruit and retain world-class talent and inspire productivity,

efficiency, creativity, and success. These properties are typically well-located, professionally managed, and well-maintained, offering a

wide range of amenities and featuring premium construction materials and finishes. Class A/A+ properties are generally newer or have

undergone substantial redevelopment and are generally expected to command higher annual rental rates compared to other classes of

similar properties. AAA locations are in close proximity to concentrations of specialized skills, knowledge, institutions, and related

businesses. It is important to note that our definition of property classification may not be directly comparable to other equity REITs.

Credit Rating

Represents the credit ratings assigned by S&P Global Ratings or Moody’s Ratings as of March 31, 2025. A credit rating is not

a recommendation to buy, sell, or hold securities and may be subject to revision or withdrawal at any time.

Development, redevelopment, and pre-construction

A key component of our business model is our disciplined allocation of capital to the development and redevelopment of new

Class A/A+ properties, as well as property enhancements identified during the underwriting of certain acquired properties. These efforts

are primarily concentrated in collaborative Megacampus™ ecosystems within AAA life science innovation clusters, as well as other

strategic locations that support innovation and growth. These projects are generally focused on providing high-quality, generic, and

reusable spaces that meet the real estate requirements of a wide range of tenants. Upon completion, each development or

redevelopment project is expected to generate increases in rental income, net operating income, and cash flows. Our development and

redevelopment projects are generally in locations that are highly desirable to high-quality entities, which we believe results in higher

occupancy levels, longer lease terms, higher rental income, higher returns, and greater long-term asset value.

Development projects generally consist of the ground-up development of generic and reusable laboratory facilities.

Redevelopment projects consist of the permanent change in use of acquired office, warehouse, or shell space into laboratory space.

We generally will not commence new development projects for aboveground construction of new Class A/A+ laboratory space without

first securing significant pre-leasing for such space, except when there is solid market demand for high-quality Class A/A+ properties.

Pre-construction activities include entitlements, permitting, design, site work, and other activities preceding commencement of

construction of aboveground building improvements. The advancement of pre-construction efforts is focused on reducing the time

required to deliver projects to prospective tenants. These critical activities add significant value for future ground-up development and

are required for the vertical construction of buildings. Ultimately, these projects will provide high-quality facilities and are expected to

generate significant revenue and cash flows.

Development, redevelopment, and pre-construction spending also includes the following costs: (i) amounts to bring certain

acquired properties up to market standard and/or other costs identified during the acquisition process (generally within two years of

acquisition) and (ii) permanent conversion of space for highly flexible, move-in-ready laboratory space to foster the growth of promising

early- and growth-stage life science companies.

Revenue-enhancing and repositioning capital expenditures represent spending to reposition or significantly change the use of

a property, including through improvement in the asset quality from Class B to Class A/A+.

Non-revenue-enhancing capital expenditures represent costs required to maintain the current revenues of a stabilized

property, including the associated costs for renewed and re-leased space.

Dividend payout ratio (common stock)

Dividend payout ratio (common stock) is the ratio of the absolute dollar amount of dividends on our common stock (shares of

common stock outstanding on the respective record dates multiplied by the related dividend per share) to funds from operations

attributable to Alexandria’s common stockholders – diluted, as adjusted.

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Dividend yield

Dividend yield for the quarter represents the annualized quarter dividend divided by the closing common stock price at the end

of the quarter.

Fixed-charge coverage ratio

Fixed-charge coverage ratio is a non-GAAP financial measure representing the ratio of Adjusted EBITDA to cash interest and

fixed charges. We believe that this ratio is useful to investors as a supplemental measure of our ability to satisfy fixed financing

obligations and preferred stock dividends. Cash interest is equal to interest expense calculated in accordance with GAAP plus

capitalized interest, less amortization of loan fees and debt premiums (discounts).

The following table reconciles interest expense, the most directly comparable financial measure calculated and presented in

accordance with GAAP, to cash interest and computes fixed-charge coverage ratio for the three months ended March 31, 2025 and

2024 (dollars in thousands):

Three Months Ended March 31,
2025 2024
Adjusted EBITDA $541,408 $551,607
Interest expense $50,876 $40,840
Capitalized interest 80,065 81,840
Amortization of loan fees (4,691) (4,142)
Amortization of debt discounts (349) (318)
Cash interest and fixed charges $125,901 $118,220
Fixed-charge coverage ratio:
– quarter annualized 4.3x 4.7x
– trailing 12 months 4.4x 4.7x

We are not able to forecast the net income of future periods without unreasonable effort and therefore do not provide a

reconciliation for fixed-charge coverage ratio on a forward-looking basis. This is due to the inherent difficulty of forecasting the timing

and/or amount of items that depend on market conditions outside of our control, including the timing of dispositions, capital events, and

financing decisions, as well as quarterly components such as gain on sales of real estate, unrealized gains or losses on non-real estate

investments, impairment of real estate, impairment of non-real estate investments, and changes in the provision for expected credit

losses on financial instruments. Our attempt to predict these amounts may produce significant but inaccurate estimates, which would be

potentially misleading for our investors.

Gross assets

Gross assets are calculated as total assets plus accumulated depreciation as of March 31, 2025 and December 31, 2024 (in

thousands):

March 31, 2025 December 31, 2024
Total assets $37,600,428 $37,527,449
Accumulated depreciation 5,886,561 5,625,179
Gross assets $43,486,989 $43,152,628

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Incremental annual net operating income on development and redevelopment projects

Incremental annual net operating income represents the amount of net operating income, on an annual basis, expected to be

realized upon a project being placed into service and achieving full occupancy. Incremental annual net operating income is calculated

as the initial stabilized yield multiplied by the project’s total cost at completion.

Initial stabilized yield (unlevered)

Initial stabilized yield is calculated as the estimated amounts of net operating income at stabilization divided by our investment

in the property. For this calculation, we exclude any tenant-funded and tenant-built landlord improvements from our investment in the

property. Our initial stabilized yield excludes the benefit of leverage. Our cash rents related to our development and redevelopment

projects are generally expected to increase over time due to contractual annual rent escalations. Our estimates for initial stabilized

yields, initial stabilized yields (cash basis), and total costs at completion represent our initial estimates at the commencement of the

project. We expect to update this information upon completion of the project, or sooner if there are significant changes to the expected

project yields or costs.

•Initial stabilized yield reflects rental income, including contractual rent escalations and any rent concessions over the

term(s) of the lease(s), calculated on a straight-line basis, and any amortization of deferred revenue related to tenant-

funded and tenant-built landlord improvements.

•Initial stabilized yield (cash basis) reflects cash rents at the stabilization date after initial rental concessions, if any, have

elapsed and our total cash investment in the property.

Investment-grade or publicly traded large cap tenants

Investment-grade or publicly traded large cap tenants represent tenants that are investment-grade rated or publicly traded

companies with an average daily market capitalization greater than $10 billion for the twelve months ended March 31, 2025, as reported

by Bloomberg Professional Services. Credit ratings from Moody’s Ratings and S&P Global Ratings reflect credit ratings of the tenant’s

parent entity, and there can be no assurance that a tenant’s parent entity will satisfy the tenant’s lease obligation upon such tenant’s

default. We monitor the credit quality and related material changes of our tenants. Material changes that cause a tenant’s market

capitalization to decrease below $10 billion, which are not immediately reflected in the twelve-month average, may result in their

exclusion from this measure.

Investments in real estate

The following table presents our new Class A/A+ development and redevelopment pipeline, excluding properties held for sale,

as a percentage of gross assets and as a percentage of annual rental revenue as of March 31, 2025 (dollars in thousands):

Percentage of
Book Value Gross Assets Annual Rental<br><br>Revenue
Under construction projects $3,688,301 8% —%
Income-producing/potential cash flows/covered land play(1) 3,154,318 7 1
Land 1,614,352 4
$8,456,971 19% 1%

(1)Includes projects with existing buildings that are generating or can generate operating cash flows. Also includes development rights associated with existing operating

campuses.

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The square footage presented in the table below is classified as operating as of March 31, 2025. These lease expirations or

vacant space at recently acquired properties represent future opportunities for which we have the intent, subject to market conditions

and leasing, to commence first-time conversion from non-laboratory space to laboratory space, or to commence future ground-up

development:

Dev/Redev RSF of Lease Expirations Targeted for<br><br>Development and Redevelopment
Property/Submarket 2025 2026 Thereafter(1) Total
Future projects:
311 Arsenal Street/Cambridge/Inner Suburbs Redev 25,312 25,312
446, 458, 500, and 550 Arsenal Street/Cambridge/Inner Suburbs Dev 375,898 375,898
Other/Greater Boston Redev 167,549 167,549
1122 and 1150 El Camino Real/South San Francisco Dev 375,232 375,232
3875 Fabian Way/Greater Stanford Dev 228,000 228,000
2100, 2200, and 2400 Geng Road/Greater Stanford Dev 78,501 78,501
960 Industrial Road/Greater Stanford Dev 112,590 112,590
Campus Point by Alexandria/University Town Center Dev 164,144 164,144
Sequence District by Alexandria/Sorrento Mesa Dev/Redev 686,290 686,290
410 West Harrison Street/Elliott Bay Dev 17,205 17,205
Other/Seattle Dev 68,401 68,401
100 Capitola Drive/Research Triangle Dev 34,527 34,527
1001 Trinity Street and 1020 Red River Street/Austin Dev/Redev 198,972 198,972
Canada Redev 247,743 247,743
224,284 2,556,080 2,780,364

(1)Includes vacant square footage as of March 31, 2025.

Joint venture financial information

We present components of balance sheet and operating results information related to our real estate joint ventures, which are

not presented, or intended to be presented, in accordance with GAAP. We present the proportionate share of certain financial line items

as follows: (i) for each real estate joint venture that we consolidate in our financial statements, which are controlled by us through

contractual rights or majority voting rights, but of which we own less than 100%, we apply the noncontrolling interest economic

ownership percentage to each financial item to arrive at the amount of such cumulative noncontrolling interest share of each component

presented; and (ii) for each real estate joint venture that we do not control and do not consolidate, and are instead controlled jointly or

by our joint venture partners through contractual rights or majority voting rights, we apply our economic ownership percentage to each

financial item to arrive at our proportionate share of each component presented.

The components of balance sheet and operating results information related to our real estate joint ventures do not represent

our legal claim to those items. For each entity that we do not wholly own, the joint venture agreement generally determines what equity

holders can receive upon capital events, such as sales or refinancing, or in the event of a liquidation. Equity holders are normally

entitled to their respective legal ownership of any residual cash from a joint venture only after all liabilities, priority distributions, and

claims have been repaid or satisfied.

We believe that this information can help investors estimate the balance sheet and operating results information related to our

partially owned entities. Presenting this information provides a perspective not immediately available from consolidated financial

statements and one that can supplement an understanding of the joint venture assets, liabilities, revenues, and expenses included in

our consolidated results.

The components of balance sheet and operating results information related to our real estate joint ventures are limited as an

analytical tool as the overall economic ownership interest does not represent our legal claim to each of our joint ventures’ assets,

liabilities, or results of operations. In addition, joint venture financial information may include financial information related to the

unconsolidated real estate joint ventures that we do not control. We believe that in order to facilitate for investors a clear understanding

of our operating results and our total assets and liabilities, joint venture financial information should be examined in conjunction with our

consolidated statements of operations and balance sheets. Joint venture financial information should not be considered an alternative

to our consolidated financial statements, which are presented and prepared in accordance with GAAP.

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Megacampus™

A Megacampus ecosystem is a cluster campus that consist of approximately 1 million RSF or greater, including operating,

active development/redevelopment, and land RSF less operating RSF expected to be demolished. The following table reconciles our

annual rental revenue and development and redevelopment pipeline RSF as of March 31, 2025 (dollars in thousands):

Annual Rental<br><br>Revenue Development and<br><br>Redevelopment<br><br>Pipeline RSF
Megacampus $1,567,014 20,364,808
Core and non-core 509,796 8,513,815
Total $2,076,810 28,878,623
Megacampus as a percentage of annual rental revenue and of total development and<br><br>redevelopment pipeline RSF 75% 71%

Net cash provided by operating activities after dividends

Net cash provided by operating activities after dividends includes the deduction for distributions to noncontrolling interests. For

purposes of this calculation, changes in operating assets and liabilities are excluded as they represent timing differences.

Net debt and preferred stock to Adjusted EBITDA

Net debt and preferred stock to Adjusted EBITDA is a non-GAAP financial measure that we believe is useful to investors as a

supplemental measure of evaluating our balance sheet leverage. Net debt and preferred stock is equal to the sum of total consolidated

debt less cash, cash equivalents, and restricted cash, plus preferred stock outstanding as of the end of the period. Refer to “Adjusted

EBITDA and Adjusted EBITDA margin” in this section for further information on the calculation of Adjusted EBITDA.

We are not able to forecast the net income of future periods without unreasonable effort and therefore do not provide a

reconciliation for net debt and preferred stock to Adjusted EBITDA on a forward-looking basis. This is due to the inherent difficulty of

forecasting the timing and/or amount of items that depend on market conditions outside of our control, including the timing of

dispositions, capital events, and financing decisions, as well as quarterly components such as gain on sales of real estate, unrealized

gains or losses on non-real estate investments, impairment of real estate, impairment of non-real estate investments, and provision for

expected credit losses on financial instruments. Our attempt to predict these amounts may produce significant but inaccurate estimates,

which would be potentially misleading for our investors.

The following table reconciles debt to net debt and preferred stock and computes the ratio to Adjusted EBITDA as of March 31,

2025 and December 31, 2024 (dollars in thousands):

March 31, 2025 December 31, 2024
Secured notes payable $150,807 $149,909
Unsecured senior notes payable 12,640,144 12,094,465
Unsecured senior line of credit and commercial paper 299,883
Unamortized deferred financing costs 80,776 77,649
Cash and cash equivalents (476,430) (552,146)
Restricted cash (7,324) (7,701)
Preferred stock
Net debt and preferred stock $12,687,856 $11,762,176
Adjusted EBITDA:
– quarter annualized $2,165,632 $2,273,480
– trailing 12 months $2,218,722 $2,228,921
Net debt and preferred stock to Adjusted EBITDA:
– quarter annualized 5.9x 5.2x
– trailing 12 months 5.7x 5.3x

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Net operating income, net operating income (cash basis), and operating margin

The following table reconciles net income to net operating income and net operating income (cash basis) and computes

operating margin for the three months ended March 31, 2025 and 2024 (dollars in thousands):

Three Months Ended March 31,
2025 2024
Net income $38,662 $219,176
Equity in losses (earnings) of unconsolidated real estate joint ventures 507 (155)
General and administrative expenses 30,675 47,055
Interest expense 50,876 40,840
Depreciation and amortization 342,062 287,554
Impairment of real estate 32,154
Gain on sales of real estate (13,165) (392)
Investment loss (income) 49,992 (43,284)
Net operating income 531,763 550,794
Straight-line rent revenue (22,023) (48,251)
Amortization of deferred revenue related to tenant-funded and -built landlord improvements (1,651)
Amortization of acquired below-market leases (15,222) (30,340)
Provision for expected credit losses on financial instruments 285
Net operating income (cash basis) $493,152 $472,203
Net operating income (cash basis) – annualized $1,972,608 $1,888,812
Net operating income (from above) $531,763 $550,794
Total revenues $758,158 $769,108
Operating margin 70% 72%

Net operating income is a non-GAAP financial measure calculated as net income (loss), the most directly comparable financial

measure calculated and presented in accordance with GAAP, excluding equity in the earnings of our unconsolidated real estate joint

ventures, general and administrative expenses, interest expense, depreciation and amortization, impairments of real estate, gains or

losses on early extinguishment of debt, gains or losses on sales of real estate, and investment income or loss. We believe net operating

income provides useful information to investors regarding our financial condition and results of operations because it primarily reflects

those income and expense items that are incurred at the property level. Therefore, we believe net operating income is a useful measure

for investors to evaluate the operating performance of our consolidated real estate assets. Net operating income on a cash basis is net

operating income adjusted to exclude the effect of straight-line rent, amortization of acquired above- and below-market lease revenue,

amortization of deferred revenue related to tenant-funded and tenant-built landlord improvements, and changes in the provision for

expected credit losses on financial instruments required by GAAP. We believe that net operating income on a cash basis is helpful to

investors as an additional measure of operating performance because it eliminates straight-line rent revenue and the amortization of

acquired above- and below-market leases and tenant-funded and tenant-built landlord improvements.

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Furthermore, we believe net operating income is useful to investors as a performance measure of our consolidated properties

because, when compared across periods, net operating income reflects trends in occupancy rates, rental rates, and operating costs,

which provide a perspective not immediately apparent from net income or loss. Net operating income can be used to measure the initial

stabilized yields of our properties by calculating net operating income generated by a property divided by our investment in the property.

Net operating income excludes certain components from net income in order to provide results that are more closely related to the

results of operations of our properties. For example, interest expense is not necessarily linked to the operating performance of a real

estate asset and is often incurred at the corporate level rather than at the property level. In addition, depreciation and amortization,

because of historical cost accounting and useful life estimates, may distort comparability of operating performance at the property level.

Impairments of real estate have been excluded in deriving net operating income because we do not consider impairments of real estate

to be property-level operating expenses. Impairments of real estate relate to changes in the values of our assets and do not reflect the

current operating performance with respect to related revenues or expenses. Our impairments of real estate represent the write-down in

the value of the assets to the estimated fair value less cost to sell. These impairments result from investing decisions or a deterioration

in market conditions. We also exclude realized and unrealized investment gain or loss, which results from investment decisions that

occur at the corporate level related to non-real estate investments in publicly traded companies and certain privately held entities.

Therefore, we do not consider these activities to be an indication of operating performance of our real estate assets at the property

level. Our calculation of net operating income also excludes charges incurred from changes in certain financing decisions, such as

losses on early extinguishment of debt and changes in the provision for expected credit losses on financial instruments, as these

charges often relate to corporate strategy. Property operating expenses included in determining net operating income primarily consist

of costs that are related to our operating properties, such as utilities, repairs, and maintenance; rental expense related to ground leases;

contracted services, such as janitorial, engineering, and landscaping; property taxes and insurance; and property-level salaries.

General and administrative expenses consist primarily of accounting and corporate compensation, corporate insurance, professional

fees, rent, and supplies that are incurred as part of corporate office management. We calculate operating margin as net operating

income divided by total revenues.

We believe that in order to facilitate for investors a clear understanding of our operating results, net operating income should

be examined in conjunction with net income or loss as presented in our consolidated statements of operations. Net operating income

should not be considered as an alternative to net income or loss as an indication of our performance, nor as an alternative to cash flows

as a measure of our liquidity or our ability to make distributions.

Operating statistics

We present certain operating statistics related to our properties, including number of properties, RSF, occupancy percentage,

leasing activity, and contractual lease expirations as of the end of the period. We believe these measures are useful to investors

because they facilitate an understanding of certain trends for our properties. We compute the number of properties, RSF, occupancy

percentage, leasing activity, and contractual lease expirations at 100%, excluding RSF at properties classified as held for sale, for all

properties in which we have an investment, including properties owned by our consolidated and unconsolidated real estate joint

ventures. For operating metrics based on annual rental revenue, refer to “Annual rental revenue” in this section.

Same property comparisons

As a result of changes within our total property portfolio during the comparative periods presented, including changes from

assets acquired or sold, properties placed into development or redevelopment, and development or redevelopment properties recently

placed into service, the consolidated total income from rentals, as well as rental operating expenses in our operating results, can show

significant changes from period to period. In order to supplement an evaluation of our results of operations over a given quarterly or

annual period, we analyze the operating performance for all consolidated properties that were fully operating for the entirety of the

comparative periods presented, referred to as same properties. We separately present quarterly and year-to-date same property results

to align with the interim financial information required by the SEC in our management’s discussion and analysis of our financial

condition and results of operations. These same properties are analyzed separately from properties acquired subsequent to the first day

in the earliest comparable quarterly or year-to-date period presented, properties that underwent development or redevelopment at any

time during the comparative periods, unconsolidated real estate joint ventures, properties classified as held for sale, and corporate

entities (legal entities performing general and administrative functions), which are excluded from same property results. Additionally,

termination fees, if any, are excluded from the results of same properties. Refer to “Same properties” in Item 2 for additional information.

Stabilized occupancy date

The stabilized occupancy date represents the estimated date on which the project is expected to reach occupancy of 95% or

greater.

Tenant recoveries

Tenant recoveries represent revenues comprising reimbursement of real estate taxes, insurance, utilities, repairs and

maintenance, common area expenses, and other operating expenses and earned in the period during which the applicable expenses

are incurred and the tenant’s obligation to reimburse us arises.

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We classify rental revenues and tenant recoveries generated through the leasing of real estate assets within revenues in

income from rentals in our consolidated statements of operations. We provide investors with a separate presentation of rental revenues

and tenant recoveries in “Results of operations” in Item 2 because we believe it promotes investors’ understanding of our operating

results. We believe that the presentation of tenant recoveries is useful to investors as a supplemental measure of our ability to recover

operating expenses under our triple net leases, including recoveries of utilities, repairs and maintenance, insurance, property taxes,

common area expenses, and other operating expenses, and of our ability to mitigate the effect to net income for any significant

variability to components of our operating expenses.

The following table reconciles income from rentals to tenant recoveries for the three months ended March 31, 2025 and 2024

(in thousands):

Three Months Ended March 31,
2025 2024
Income from rentals $743,175 $755,551
Rental revenues (552,112) (581,400)
Tenant recoveries $191,063 $174,151

Total equity capitalization

Total equity capitalization is equal to the outstanding shares of common stock multiplied by the closing price on the last trading

day at the end of each period presented.

Total market capitalization

Total market capitalization is equal to the sum of total equity capitalization and total debt.

Unencumbered net operating income as a percentage of total net operating income

Unencumbered net operating income as a percentage of total net operating income is a non-GAAP financial measure that we

believe is useful to investors as a performance measure of the results of operations of our unencumbered real estate assets as it

reflects those income and expense items that are incurred at the unencumbered property level. Unencumbered net operating income is

derived from assets classified in continuing operations, which are not subject to any mortgage, deed of trust, lien, or other security

interest, as of the period for which income is presented.

The following table summarizes unencumbered net operating income as a percentage of total net operating income for the

three months ended March 31, 2025 and 2024 (dollars in thousands):

Three Months Ended March 31,
2025 2024
Unencumbered net operating income $530,691 $546,830
Encumbered net operating income 1,072 3,964
Total net operating income $531,763 $550,794
Unencumbered net operating income as a percentage of total net operating income 99.8% 99.3%

Weighted-average shares of common stock outstanding – diluted

From time to time, we enter into capital market transactions, including forward equity sales agreements (“Forward

Agreements”), to fund acquisitions, to fund construction of our development and redevelopment projects, and for general working

capital purposes. While the Forward Agreements are outstanding, we are required to consider the potential dilutive effect of our Forward

Agreements under the treasury stock method. Under this method, we also include the dilutive effect of unvested restricted stock awards

(“RSAs”) with forfeitable dividends in the calculation of diluted shares. Refer to Note 12 – “Earnings per share” and Note 13 –

“Stockholders’ equity” to our unaudited consolidated financial statements in Item 1 for additional information.

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The weighted-average shares of common stock outstanding used in calculating EPS – diluted, funds from operations per

share – diluted, and funds from operations per share – diluted, as adjusted, for the three months ended March 31, 2025 and 2024 are

calculated as follows. Also shown are the weighted-average unvested RSAs with nonforfeitable dividends used in calculating the

amounts allocable to these awards pursuant to the two-class method for each of the respective periods presented below (in thousands):

Three Months Ended March 31,
2025 2024
Basic shares for earnings per share 170,522 171,949
Unvested RSAs with forfeitable dividends
Diluted shares for earnings per share 170,522 171,949
Basic shares for funds from operations per share and funds from operations per share, as<br><br>adjusted 170,522 171,949
Unvested RSAs with forfeitable dividends 77
Diluted shares for funds from operations per share and funds from operations per share, as<br><br>adjusted 170,599 171,949
Weighted-average unvested RSAs with nonforfeitable dividends used in the allocations of net<br><br>income, funds from operations, and funds from operations, as adjusted 2,053 2,987

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ITEM 3. QUANTITATIVE AND QUALITATIVE DISCLOSURES ABOUT MARKET RISK

Interest rate risk

The primary market risk to which we believe we may be exposed is interest rate risk, which may result from many factors,

including government monetary and tax policies, domestic and international economic and political considerations, and other factors

that are beyond our control.

In order to modify and manage the interest rate characteristics of our outstanding debt and to limit the effects of interest rate

risks on our operations, we may utilize a variety of financial instruments, including interest rate hedge agreements, caps, floors, and

other interest rate exchange contracts. The use of these types of instruments to hedge a portion of our exposure to changes in interest

rates may carry additional risks, such as counterparty credit risk and the legal enforceability of hedge agreements. As of March 31,

2025, we did not have any outstanding interest rate hedge agreements.

Our future earnings and fair values relating to our outstanding debt are primarily dependent upon prevalent market rates of

interest. The following tables illustrate the effect of a 1% change in interest rates, assuming a zero percent interest rate floor, on our

fixed- and variable-rate debt as of March 31, 2025 (in thousands):

Annualized effect on future earnings due to variable-rate debt:
Rate increase of 1% $(1,240)
Rate decrease of 1% $1,240
Effect on fair value of total consolidated debt:
Rate increase of 1% $(780,013)
Rate decrease of 1% $893,198

These amounts are determined by considering the effect of the hypothetical interest rates on our borrowings as of March 31,

  1. These analyses do not consider the effects of the reduced level of overall economic activity that could exist in such an

environment. Furthermore, in the event of a change of such magnitude, we would consider taking actions to further mitigate our

exposure to the change. Because of the uncertainty of the specific actions that would be taken and their possible effects, the sensitivity

analyses assume no changes in our capital structure.

Equity price risk

We have exposure to equity price market risk because we hold equity investments in publicly traded companies and privately

held entities. All of our investments in actively traded public companies are reflected in our consolidated balance sheets at fair value.

Our investments in privately held entities that report NAV per share are measured at fair value using NAV as a practical expedient to fair

value. Our equity investments in privately held entities that do not report NAV per share are measured at cost less impairments,

adjusted for observable price changes during the period. Changes in fair value of public investments, changes in NAV per share

reported by privately held entities, and observable price changes of privately held entities that do not report NAV per share are

classified as investment income (loss) in our consolidated statements of operations. There is no assurance that future declines in value

will not have a material adverse effect on our future results of operations. The following table illustrates the effect that a 10% change in

the value of our equity investments would have on earnings as of March 31, 2025 (in thousands):

Equity price risk:
Fair value increase of 10% $147,969
Fair value decrease of 10% $(147,969)

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Foreign currency exchange rate risk

We have exposure to foreign currency exchange rate risk related to our subsidiaries operating in Canada and Asia. The

functional currencies of our foreign subsidiaries are the local currencies in each respective country. Gains or losses resulting from the

translation of our foreign subsidiaries’ balance sheets and statements of operations are classified in accumulated other comprehensive

income (loss) as a separate component of total equity and are excluded from net income (loss). Gains or losses will be reflected in our

consolidated statements of operations when there is a sale or partial sale of our investment in these operations or upon a complete or

substantially complete liquidation of the investment. The following tables illustrate the effect that a 10% change in foreign currency rates

relative to the U.S. dollar would have on our potential future earnings and on the fair value of our net investment in foreign subsidiaries

based on our current operating assets outside the U.S. as of March 31, 2025 (in thousands):

Effect on potential future earnings due to foreign currency exchange rate:
Rate increase of 10% $24
Rate decrease of 10% $(24)
Effect on the fair value of net investment in foreign subsidiaries due to foreign currency exchange rate:
Rate increase of 10% $35,533
Rate decrease of 10% $(35,533)

The sensitivity analyses assume a parallel shift of all foreign currency exchange rates with respect to the U.S. dollar; however,

foreign currency exchange rates do not typically move in such a manner, and actual results may differ materially.

Our exposure to market risk elements for the three months ended March 31, 2025 was consistent with the risk elements

presented above, including the effects of changes in interest rates, equity prices, and foreign currency exchange rates.

ITEM 4. CONTROLS AND PROCEDURES

Evaluation of disclosure controls and procedures

As of March 31, 2025, we had performed an evaluation, under the supervision of our principal executive officers and principal

financial officer, of the effectiveness of the design and operation of our disclosure controls and procedures. These controls and

procedures have been designed to ensure that information required for disclosure is recorded, processed, summarized, and reported

within the requisite time periods. Based on our evaluation, the principal executive officers and principal financial officer concluded that

our disclosure controls and procedures were effective as of March 31, 2025.

Changes in internal control over financial reporting

There has not been any change in our internal control over financial reporting during the three months ended March 31, 2025

that has materially affected, or is reasonably likely to materially affect, our internal control over financial reporting.

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PART II – OTHER INFORMATION

ITEM 1. LEGAL PROCEEDINGS

In 2006, ARE-East River Science Park, LLC, a subsidiary of Alexandria Real Estate Equities, Inc., was granted an option to

incorporate a land parcel adjacent to and north of the Alexandria Center® for Life Science – New York City (“ACLS-NYC”) campus

(“Option Parcel”) into the existing ground lease of that campus. The Option Parcel will allow ARE-East River Science Park, LLC to

develop a future world-class life science building within the ACLS-NYC campus. ARE-East River Science Park, LLC’s investment in pre-

construction costs related to the development of the Option Parcel, including costs related to design, engineering, environmental,

survey/title, and permitting and legal costs, aggregated $171.1 million as of March 31, 2025.

On August 6, 2024, ARE-East River Science Park, LLC filed a lawsuit in the U.S. District Court for the Southern District of New

York against its landlord, New York City Health + Hospitals Corporation (“H+H”), and the New York City Economic Development

Corporation (“EDC”). On January 24, 2025, ARE-East River Science Park, LLC filed a first amended complaint. The lawsuit alleges two

principal claims against H+H and EDC: fraud in the inducement, and, in the alternative, breach of contract in violation of the implied

covenant of good faith and fair dealing. As alleged in the complaint, ARE-East River Science Park, LLC’s claims arise from H+H’s and

EDC’s misrepresentations and concealment of material facts in connection with a floodwall, which H+H and EDC are seeking to require

ARE-East River Science Park, LLC to integrate into the development of the Option Parcel. ARE-East River Science Park, LLC alleges

that H+H’s and EDC’s misconduct have prevented it from commencing the development of the Option Parcel. In light of the pending

litigation, the closing date for our option and thus the commencement date for construction of the third tower at the campus are

presently indeterminate. Among other things, ARE-East River Science Park, LLC is seeking significant damages and equitable relief

from the court to confirm our understanding that the option is in full force and effect.

This matter exposes us to potential losses ranging from zero to the full amount of the investment in the project aggregating

$171.1 million as of March 31, 2025, depending on any collection of damages and/or the ability to develop the project. We performed a

probability-weighted recoverability analysis based on initial estimates of various possible outcomes and determined no impairment was

present as of March 31, 2025.

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ITEM 1A. RISK FACTORS

In addition to the information set forth in this quarterly report on Form 10-Q, one should also carefully review and consider the

information contained in the other reports and periodic filings that we make with the SEC, including, without limitation, the information

contained under the caption “Item 1A. Risk factors” in our annual report on Form 10-K for the year ended December 31, 2024. Those risk

factors could materially affect our business, financial condition, and results of operations. The risks that we describe in our public filings

are not the only risks that we face. Additional risks and uncertainties not currently known to us, or that we presently deem to be

immaterial, also may materially adversely affect our business, financial condition, and results of operations.

There have been no material changes in our risk factors from those disclosed under the caption “Item 1A. Risk factors” in our

annual report on Form 10-K for the year ended December 31, 2024, except for the following updates:

Changes to regulatory, funding, staffing, trade, and other policies and actions by the current U.S. government

could adversely affect our business operations or those of our tenants and our venture investment portfolio

companies.

Domestic and international policy shifts may introduce considerable uncertainty to the macroeconomic and regulatory

landscape in which we, our tenants, and our venture investment portfolio companies operate. Our tenants and our venture

investment portfolio companies include entities in the pharmaceutical, biotechnology, medical device, life science and related

industries, academic institutions, government institutions, and private foundations that determine their research and

development budgets based on several factors, including the availability of government and other funding, and the operational

efficiency of public regulatory institutions.

Since January 2025, the current U.S. administration has enacted and proposed substantial policy changes that affect

federal health agencies, research funding, public health priorities, and international trade. These measures — ranging from

staffing and budget reductions at the U.S. Food and Drug Administration (“FDA”) and the National Institutes of Health (“NIH”) to

sweeping tariff actions, as described below — may significantly disrupt the life science ecosystem in which we, our tenants, and

our venture investment portfolio companies operate.

Reductions in FDA Workforce and Budget

On April 1, 2025, the FDA underwent layoffs of approximately 3,400 employees, representing greater than 15% of its

workforce. Such workforce reductions at the FDA have raised concerns regarding the agency’s capacity to perform timely

regulatory reviews and approvals of drugs and other medical products. Recent and/or potential further reductions in workforce

or other personnel changes at the FDA, including terminations, may disrupt the agency’s review and approval processes for our

tenants’ and our venture investment portfolio companies’ products. Such disruptions could lead to setbacks in research and

development timelines, negatively impacting life science companies’ ability to advance their pipelines, secure investor funding,

or achieve commercial viability, which could severely affect their operations and financial performance and, as a result,

adversely impact our operating and financial results.

NIH Grant Cuts and Impact on Research Institutions

The current U.S. administration has implemented significant policy changes affecting the NIH and leading to

substantial disruptions in biomedical research across the U.S. These actions included staff layoffs and funding cuts, as

described below, and resulted in the suspension of numerous research projects, posing risks to scientific advancement and

introducing significant uncertainty for some of our tenants and venture investment portfolio companies.

•NIH budget freeze. On January 27, 2025, the U.S. administration issued an executive order to suspend NIH grant

funding, freezing much of NIH’s approximately $47 billion budget for 2025. Though the suspension was eventually

blocked and reversed, the NIH subsequently laid off nearly 1,200 employees, with potential plans to eliminate up to

5,000 positions across its approximately 20,000-person workforce.

•Termination of NIH’s grants and funding commitments to major research institutions. On January 20, 2025, President

Trump issued an executive order directing every U.S. agency, including the NIH, to “terminate, to the maximum extent

allowed by law” all grants relating to diversity, equity, and inclusion. On January 29, 2025, the President issued an

executive order to make it “the policy of the United States to combat anti-Semitism vigorously, using all available and

appropriate legal tools, to prosecute, remove, or otherwise hold to account the perpetrators of unlawful anti-Semitic

harassment and violence.” As a result of one or both executive orders, the NIH, the world’s largest funder of

biomedical research, has withheld funding from certain U.S. research institutions.

•15% cap on indirect cost reimbursements of all NIH grants. On February 7 , 2025, the NIH introduced a policy limiting

indirect cost reimbursements to 15% for all NIH grants, representing a significant reduction from historic levels, which

were approximately double that rate on average, and in some cases significantly higher. This change threatens to

substantially impact the ability of research institutions to support their infrastructure and administrative costs, including

their ability to lease life science facilities.

108

A coalition of 22 state attorneys general, along with organizations like the Association of American Medical Colleges,

filed lawsuits challenging the NIH’s policy changes, particularly the 15% cap on indirect costs. On April 7, 2025, a federal court

issued a permanent injunction blocking the enforcement of this cap. However, the current U.S. administration has signaled its

intent to appeal and/or pursue similar funding restrictions through future legislative or administrative actions. If implemented,

any such funding cap could negatively impact our tenants that depend on grant funding for its operations. It could also reduce

the financial resources available to such tenants, forcing them to scale back operations, reduce leased space, or delay their

plans for lease expansion.

Tariff Escalation, Trade Disruption, and Financial Market Instability

Beginning in March 2025, the U.S. government implemented a series of trade actions that have reshaped global

economic relations and triggered market volatility, specifically:

•On February 1, 2025, President Trump signed executive orders imposing a 25% tariff on all goods from Mexico and

Canada and a 10% tariff on China.

•On March 3, 2025, the President increased tariffs on all products from China from 10% to 20%. He also implemented

new 25% tariffs on imports from Mexico and Canada.

•On April 2, 2025, the President declared a national emergency to address the U.S. trade deficit and imposed a 10%

universal import tariff on all goods, with higher rates for 57 trading partners. This announcement led to a significant

stock market decline, with the S&P 500 Index, Dow Jones Industrial Average, and the Nasdaq Composite dropping by

approximately 6.0%, 5.5%, and 5.8%, respectively.

•On April 9, 2025, facing a global market meltdown, the President announced a 90-day pause on tariffs for most

countries but raised the tax rate on Chinese imports to 125%. Following the announcement, the S&P 500 Index surged

9.5%. However, on April 10, 2025, U.S. stocks fell as the initial euphoria over the pause on tariffs faded, with investors

reassessing ongoing trade tensions and their potential impact.

•On April 14, 2025, the U.S. government launched an investigation into pharmaceuticals to justify tariffs that may be

implemented on pharmaceutical products. In 2024, over $200 billion in pharmaceutical products were imported to the

U.S., and it is estimated that U.S. tariffs could add $46 billion in costs to the pharmaceutical industry.

If financial markets continue to be disrupted, we may face the following risks:

•Restricted access to capital. Market instability may hinder our ability to raise capital, including through dispositions,

sales of partial interests, and new debt capital, and could potentially delay our current or future development and

redevelopment projects.

•Rising construction costs. Our general contractors may face difficulty procuring construction materials at reasonable

prices, particularly those subject to tariffs or disrupted supply, which may lead to project delays and/or increased costs.

Rising costs and procurement challenges could significantly impact the yields and delay net operating income

commencement from our current and future development and redevelopment pipeline.

•Risks to tenant operations. Many of our tenants rely on the import and export of materials, components, and/or

specialized equipment. As a result, their products may become prohibitively expensive to manufacture or sell. These

challenges may adversely affect our tenants’ ability to meet their lease obligations or to renew their leases with us.

•Macroeconomic impact. Widespread tariffs, restricted trade, increased market volatility, and reduced investor

confidence may trigger inflationary pressure and elevate the risk of a U.S. recession.

The cost increases that may result from tariffs, trade conflicts, and financial market volatility may significantly impact

our development and redevelopment projects. Elevated material costs may lead to higher overall project budgets and extended

construction timelines or require modifications to project scope to preserve economic feasibility. Any such adjustments may

adversely affect our ability to deliver space on time and within budget, delay occupancy and commencement of rental income,

and impact projected net operating income and yields.

We cannot provide assurance that our tenants or venture investment portfolio companies will be able to raise capital,

secure approvals, or sustain operations in this environment. Tenants that are unable to sufficiently mitigate the regulatory,

financial, or geopolitical risks described above may not be able to meet their lease obligations, which may force them to reduce

leased space, not renew, or terminate their leases with us. Such developments may reduce the performance of our real estate

and non-real estate portfolios and negatively affect our financial condition, results of operations, or cash flows, our ability to

make distributions to our stockholders, and our stock price. Any negative news relating to the life science industry, our tenants,

and our venture investment portfolio companies may also adversely impact our stock price.

109

ITEM 2. UNREGISTERED SALES OF EQUITY SECURITIES AND USE OF PROCEEDS

Issuer Purchases of Equity Securities

From January 1, 2025 to March 31, 2025, we repurchased 2,152,293 shares of our common stock aggregating $208.1 million

under the program. The repurchases were made on the open market pursuant to a trading plan established under Rule 10b5-1 of the

Securities Exchange Act of 1934, as amended. As of March 31, 2025, we had remaining authorization to repurchase shares with an

aggregate value up to $241.8 million.

The following table summarizes share repurchases executed under the program during the three months ended March 31,

2025:

Total Number<br><br>of Shares<br><br>Purchased(1) Average Price Paid<br><br>per Share Total Number of<br><br>Shares Purchased as<br><br>Part of Publicly<br><br>Announced Plans Approximate Value of<br><br>Shares That May Yet<br><br>Be Purchased<br><br>Under Plans
January 1, 2025 – January 31, 2025 1,541,974 $97.26 2,038,250 $299,934,205
February 1, 2025 – February 28, 2025 610,319 $95.32 2,648,569 $241,759,706

(1)On December 9, 2024, we announced that our Board of Directors authorized a share repurchase program allowing the repurchase of shares with an aggregate

value up to $500.0 million until December 31, 2025 in the open market, through privately negotiated transactions, or otherwise, in accordance with all applicable

securities laws and regulations, including Rule 10b-18 of the Exchange Act.

ITEM 5. OTHER INFORMATION

Disclosure of 10b5-1 plans

During the three months ended March 31, 2025, none of our officers or directors adopted or terminated any contract,

instruction, or written plan for the purchase or sale of our securities that was intended to satisfy the affirmative defense conditions of

Rule 10b5-1(c) or any “non-Rule 10b5-1 trading arrangement.”

110

ITEM 6. EXHIBITS

Exhibit<br><br>Number Exhibit Title Incorporated by<br><br>Reference to: Date Filed
3.1* Articles of Amendment and Restatement of the Company, dated May 21, 1997 Form 10-Q August 14, 1997
3.2* Certificate of Correction of the Company, dated June 20, 1997 Form 10-Q August 14, 1997
3.3* Articles of Amendment of the Company, effective as of May 10, 2017 Form 8-K May 12, 2017
3.4* Articles of Amendment of the Company, effective as of May 18, 2022 Form 8-K May 19, 2022
3.5* Articles Supplementary, dated June 9, 1999, relating to the 9.50% Series A<br><br>Cumulative Redeemable Preferred Stock Form 10-Q August 13, 1999
3.6* Articles Supplementary, dated February 10, 2000, relating to the election to be<br><br>subject to Subtitle 8 of Title 3 of the Maryland General Corporation Law Form 8-K February 10, 2000
3.7* Articles Supplementary, dated February 10, 2000, relating to the Series A<br><br>Junior Participating Preferred Stock Form 8-K February 10, 2000
3.8* Articles Supplementary, dated January 18, 2002, relating to the 9.10%<br><br>Series B Cumulative Redeemable Preferred Stock Form 8-A January 18, 2002
3.9* Articles Supplementary, dated June 22, 2004, relating to the 8.375% Series C<br><br>Cumulative Redeemable Preferred Stock Form 8-A June 28, 2004
3.10* Articles Supplementary, dated March 25, 2008, relating to the 7.00% Series D<br><br>Cumulative Convertible Preferred Stock Form 8-K March 25, 2008
3.11* Articles Supplementary, dated March 12, 2012, relating to the 6.45% Series E<br><br>Cumulative Redeemable Preferred Stock Form 8-K March 14, 2012
3.12* Articles Supplementary, effective as of May 10, 2017, relating to Reclassified<br><br>Preferred Stock Form 8-K May 12, 2017
3.13* Amended and Restated Bylaws of the Company (Amended December 6,<br><br>2024) Form 8-K December 9, 2024
4.1* Indenture, dated as of February 13, 2025, among Alexandria Real Estate<br><br>Equities, Inc., as Issuer, Alexandria Real Estate Equities, L.P., as Guarantor,<br><br>and U.S. Bank Trust Company, National Association, as Trustee Form 8-K February 13, 2025
4.2* Supplemental Indenture No. 1, dated as of February 13, 2025, by and among<br><br>Alexandria Real Estate Equities, Inc., as Issuer, Alexandria Real Estate<br><br>Equities, L.P., as Guarantor, and U.S. Bank Trust Company, National<br><br>Association, as Trustee Form 8-K February 13, 2025
4.3* Form of 5.50% Senior Note due 2035 (included in Exhibit 4.2 above) Form 8-K February 13, 2025
22.1 List of Guarantor Subsidiaries of the Company N/A Filed herewith
31.1 Certification of Principal Executive Officer Pursuant to Section 302 of the<br><br>Sarbanes-Oxley Act of 2002 N/A Filed herewith
31.2 Certification of Principal Executive Officer Pursuant to Section 302 of the<br><br>Sarbanes-Oxley Act of 2002 N/A Filed herewith
31.3 Certification of Principal Financial Officer Pursuant to Section 302 of the<br><br>Sarbanes-Oxley Act of 2002 N/A Filed herewith
32.0 Certification of Principal Executive Officers and Principal Financial Officer<br><br>Pursuant to 18 U.S.C. Section 1350, as Adopted Pursuant to Section 906 of<br><br>the Sarbanes-Oxley Act of 2002 N/A Filed herewith
101.1 The following materials from the Company’s quarterly report on Form 10-Q for<br><br>the quarterly period ended March 31, 2025, formatted in iXBRL (Inline<br><br>eXtensible Business Reporting Language): (i) Consolidated Balance Sheets<br><br>as of March 31, 2025 and December 31, 2024 (unaudited), (ii) Consolidated<br><br>Statements of Operations for the three months ended March 31, 2025 and<br><br>2024 (unaudited), (iii) Consolidated Statements of Comprehensive Income for<br><br>the three months ended March 31, 2025 and 2024 (unaudited), (iv)<br><br>Consolidated Statements of Changes in Stockholders’ Equity and<br><br>Noncontrolling Interests for the three months ended March 31, 2025 and 2024<br><br>(unaudited), (v) Consolidated Statements of Cash Flows for the three months<br><br>ended March 31, 2025 and 2024 (unaudited), and (vi) Notes to Consolidated<br><br>Financial Statements (unaudited) N/A Filed herewith
104 Cover Page Interactive Data File – the cover page from this Quarterly Report<br><br>on Form 10-Q for the quarter ended March 31, 2025 is formatted in Inline<br><br>XBRL and contained in Exhibit 101.1 N/A Filed herewith

(*) Incorporated by reference.

111

SIGNATURES

Pursuant to the requirements of the Securities Exchange Act of 1934, the registrant has duly caused this report to be signed

on its behalf by the undersigned, thereunto duly authorized, on April 28, 2025.

ALEXANDRIA REAL ESTATE EQUITIES, INC.
/s/ Joel S. Marcus
Joel S. Marcus<br><br>Executive Chairman<br><br>(Principal Executive Officer)
/s/ Peter M. Moglia
Peter M. Moglia<br><br>Chief Executive Officer and Chief Investment Officer<br><br>(Principal Executive Officer)
/s/ Marc E. Binda
Marc E. Binda<br><br>Chief Financial Officer and Treasurer<br><br>(Principal Financial Officer)

Document

EXHIBIT 22.1

List of Guarantor Subsidiaries of Alexandria Real Estate Equities, Inc.

The following subsidiary was, as of March 31, 2025, a guarantor of the registrant's 3.45% Senior Notes due 2025, 4.30% Senior Notes due 2026, 3.80% Senior Notes due 2026, 3.95% Senior Notes due 2027, 3.95% Senior Notes due 2028, 4.50% Senior Notes due 2029, 2.75% Senior Notes due 2029, 4.70% Senior Notes due 2030, 4.90% Senior Notes due 2030, 3.375% Senior Notes due 2031, 2.00% Senior Notes due 2032, 1.875% Senior Notes due 2033, 2.95% Senior Notes due 2034, 4.75% Senior Notes due 2035, 5.50% Senior Notes due 2035, 5.25% Senior Notes due 2036, 4.85% Senior Notes due 2049, 4.00% Senior Notes due 2050, 3.00% Senior Notes due 2051, 3.55% Senior Notes due 2052, 5.15% Senior Notes due 2053, and 5.625% Senior Notes due 2054.

Name of Subsidiary Jurisdiction of Organization
Alexandria Real Estate Equities, L.P. Delaware

Document

EXHIBIT 31.1

CERTIFICATION OF PRINCIPAL EXECUTIVE OFFICER

PURSUANT TO

SECTION 302 OF THE SARBANES-OXLEY ACT OF 2002

I, Joel S. Marcus, certify that:

1.    I have reviewed this Quarterly Report on Form 10-Q of Alexandria Real Estate Equities, Inc.;

2.    Based on my knowledge, this report does not contain any untrue statement of a material fact or omit to state a material fact necessary to make the statements made, in light of the circumstances under which such statements were made, not misleading with respect to the period covered by this report;

3.    Based on my knowledge, the financial statements, and other financial information included in this report, fairly present in all material respects the financial condition, results of operations, and cash flows of the registrant as of, and for, the periods presented in this report;

4.    The registrant’s other certifying officers and I are responsible for establishing and maintaining disclosure controls and procedures (as defined in Exchange Act Rules 13a-15(e) and 15d-15(e)) and internal control over financial reporting (as defined in Exchange Act Rules 13a-15(f) and 15d-15(f)) for the registrant and have:

a.    Designed such disclosure controls and procedures, or caused such disclosure controls and procedures to be designed under our supervision, to ensure that material information relating to the registrant, including its consolidated subsidiaries, is made known to us by others within those entities, particularly during the period in which this report is being prepared;

b.    Designed such internal control over financial reporting, or caused such internal control over financial reporting to be designed under our supervision, to provide reasonable assurance regarding the reliability of financial reporting and the preparation of financial statements for external purposes in accordance with generally accepted accounting principles;

c.    Evaluated the effectiveness of the registrant’s disclosure controls and procedures and presented in this report our conclusions about the effectiveness of the disclosure controls and procedures, as of the end of the period covered by this report based on such evaluation; and

d.    Disclosed in this report any change in the registrant’s internal control over financial reporting that occurred during the registrant’s most recent fiscal quarter (the registrant’s fourth fiscal quarter in the case of an annual report) that has materially affected, or is reasonably likely to materially affect, the registrant’s internal control over financial reporting; and

5.    The registrant’s other certifying officers and I have disclosed, based on our most recent evaluation of internal control over financial reporting, to the registrant’s auditors and the audit committee of the registrant’s board of directors (or persons performing the equivalent functions):

a.    All significant deficiencies and material weaknesses in the design or operation of internal control over financial reporting which are reasonably likely to adversely affect the registrant’s ability to record, process, summarize, and report financial information; and

b.    Any fraud, whether or not material, that involves management or other employees who have a significant role in the registrant’s internal control over financial reporting.

Date: April 28, 2025

/s/ Joel S. Marcus
Joel S. Marcus
Executive Chairman

Document

EXHIBIT 31.2

CERTIFICATION OF PRINCIPAL EXECUTIVE OFFICER

PURSUANT TO

SECTION 302 OF THE SARBANES-OXLEY ACT OF 2002

I, Peter M. Moglia, certify that:

1.    I have reviewed this Quarterly Report on Form 10-Q of Alexandria Real Estate Equities, Inc.;

2.    Based on my knowledge, this report does not contain any untrue statement of a material fact or omit to state a material fact necessary to make the statements made, in light of the circumstances under which such statements were made, not misleading with respect to the period covered by this report;

3.    Based on my knowledge, the financial statements, and other financial information included in this report, fairly present in all material respects the financial condition, results of operations, and cash flows of the registrant as of, and for, the periods presented in this report;

4.    The registrant’s other certifying officers and I are responsible for establishing and maintaining disclosure controls and procedures (as defined in Exchange Act Rules 13a-15(e) and 15d-15(e)) and internal control over financial reporting (as defined in Exchange Act Rules 13a-15(f) and 15d-15(f)) for the registrant and have:

a.    Designed such disclosure controls and procedures, or caused such disclosure controls and procedures to be designed under our supervision, to ensure that material information relating to the registrant, including its consolidated subsidiaries, is made known to us by others within those entities, particularly during the period in which this report is being prepared;

b.    Designed such internal control over financial reporting, or caused such internal control over financial reporting to be designed under our supervision, to provide reasonable assurance regarding the reliability of financial reporting and the preparation of financial statements for external purposes in accordance with generally accepted accounting principles;

c.    Evaluated the effectiveness of the registrant’s disclosure controls and procedures and presented in this report our conclusions about the effectiveness of the disclosure controls and procedures, as of the end of the period covered by this report based on such evaluation; and

d.    Disclosed in this report any change in the registrant’s internal control over financial reporting that occurred during the registrant’s most recent fiscal quarter (the registrant’s fourth fiscal quarter in the case of an annual report) that has materially affected, or is reasonably likely to materially affect, the registrant’s internal control over financial reporting; and

5.    The registrant’s other certifying officers and I have disclosed, based on our most recent evaluation of internal control over financial reporting, to the registrant’s auditors and the audit committee of the registrant’s board of directors (or persons performing the equivalent functions):

a.    All significant deficiencies and material weaknesses in the design or operation of internal control over financial reporting which are reasonably likely to adversely affect the registrant’s ability to record, process, summarize, and report financial information; and

b.    Any fraud, whether or not material, that involves management or other employees who have a significant role in the registrant’s internal control over financial reporting.

Date: April 28, 2025

/s/ Peter M. Moglia
Peter M. Moglia
Chief Executive Officer and Chief Investment Officer

Document

EXHIBIT 31.3

CERTIFICATION OF PRINCIPAL FINANCIAL OFFICER

PURSUANT TO

SECTION 302 OF THE SARBANES-OXLEY ACT OF 2002

I, Marc E. Binda, certify that:

1.    I have reviewed this Quarterly Report on Form 10-Q of Alexandria Real Estate Equities, Inc.;

2.    Based on my knowledge, this report does not contain any untrue statement of a material fact or omit to state a material fact necessary to make the statements made, in light of the circumstances under which such statements were made, not misleading with respect to the period covered by this report;

3.    Based on my knowledge, the financial statements, and other financial information included in this report, fairly present in all material respects the financial condition, results of operations, and cash flows of the registrant as of, and for, the periods presented in this report;

4.    The registrant’s other certifying officers and I are responsible for establishing and maintaining disclosure controls and procedures (as defined in Exchange Act Rules 13a-15(e) and 15d-15(e)) and internal control over financial reporting (as defined in Exchange Act Rules 13a-15(f) and 15d-15(f)) for the registrant and have:

a.    Designed such disclosure controls and procedures, or caused such disclosure controls and procedures to be designed under our supervision, to ensure that material information relating to the registrant, including its consolidated subsidiaries, is made known to us by others within those entities, particularly during the period in which this report is being prepared;

b.    Designed such internal control over financial reporting, or caused such internal control over financial reporting to be designed under our supervision, to provide reasonable assurance regarding the reliability of financial reporting and the preparation of financial statements for external purposes in accordance with generally accepted accounting principles;

c.    Evaluated the effectiveness of the registrant’s disclosure controls and procedures and presented in this report our conclusions about the effectiveness of the disclosure controls and procedures, as of the end of the period covered by this report based on such evaluation; and

d.    Disclosed in this report any change in the registrant’s internal control over financial reporting that occurred during the registrant’s most recent fiscal quarter (the registrant’s fourth fiscal quarter in the case of an annual report) that has materially affected, or is reasonably likely to materially affect, the registrant’s internal control over financial reporting; and

5.    The registrant’s other certifying officers and I have disclosed, based on our most recent evaluation of internal control over financial reporting, to the registrant’s auditors and the audit committee of the registrant’s board of directors (or persons performing the equivalent functions):

a.    All significant deficiencies and material weaknesses in the design or operation of internal control over financial reporting which are reasonably likely to adversely affect the registrant’s ability to record, process, summarize, and report financial information; and

b.    Any fraud, whether or not material, that involves management or other employees who have a significant role in the registrant’s internal control over financial reporting.

Date: April 28, 2025

/s/ Marc E. Binda
Marc E. Binda
Chief Financial Officer and Treasurer

Document

EXHIBIT 32.0

CERTIFICATION OF PRINCIPAL EXECUTIVE OFFICERS AND PRINCIPAL FINANCIAL OFFICER

PURSUANT TO

18 U.S.C. SECTION 1350.

AS ADOPTED PURSUANT TO

SECTION 906 OF THE SARBANES-OXLEY ACT OF 2002

I, Joel S. Marcus, certify, pursuant to 18 U.S.C. Section 1350, as adopted pursuant to Section 906 of the Sarbanes-Oxley Act of 2002, that the Quarterly Report on Form 10-Q of Alexandria Real Estate Equities, Inc. for the quarter ended March 31, 2025, fully complies with the requirements of Section 13(a) or 15(d) of the Securities Exchange Act of 1934, as amended, and that the information contained in such report fairly presents, in all material respects, the financial condition and results of operations of Alexandria Real Estate Equities, Inc.

Date: April 28, 2025

/s/ Joel S. Marcus
Joel S. Marcus
Executive Chairman

I, Peter M. Moglia, certify, pursuant to 18 U.S.C. Section 1350, as adopted pursuant to Section 906 of the Sarbanes-Oxley Act of 2002, that the Quarterly Report on Form 10-Q of Alexandria Real Estate Equities, Inc. for the quarter ended March 31, 2025, fully complies with the requirements of Section 13(a) or 15(d) of the Securities Exchange Act of 1934, as amended, and that the information contained in such report fairly presents, in all material respects, the financial condition and results of operations of Alexandria Real Estate Equities, Inc.

Date: April 28, 2025

/s/ Peter M. Moglia
Peter M. Moglia
Chief Executive Officer and Chief Investment Officer

I, Marc E. Binda, certify, pursuant to 18 U.S.C. Section 1350, as adopted pursuant to Section 906 of the Sarbanes-Oxley Act of 2002, that the Quarterly Report on Form 10-Q of Alexandria Real Estate Equities, Inc. for the quarter ended March 31, 2025, fully complies with the requirements of Section 13(a) or 15(d) of the Securities Exchange Act of 1934, as amended, and that the information contained in such report fairly presents, in all material respects, the financial condition and results of operations of Alexandria Real Estate Equities, Inc.

Date: April 28, 2025

/s/ Marc E. Binda
Marc E. Binda
Chief Financial Officer and Treasurer