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10-Q

Alexandria Real Estate Equities, Inc. (ARE)

10-Q 2026-04-27 For: 2026-03-31
View Original
Added on April 27, 2026

UNITED STATES

SECURITIES AND EXCHANGE COMMISSION

Washington, D.C. 20549

FORM 10-Q

(Mark One)

☒QUARTERLY REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES

EXCHANGE ACT OF 1934

For the quarterly period ended March 31, 2026

OR

☐TRANSITION REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES

EXCHANGE ACT OF 1934

For the transition period from ____________ to ____________

Commission file number 1-12993

ALEXANDRIA REAL ESTATE EQUITIES, INC.

(Exact name of registrant as specified in its charter)

Maryland 95-4502084
(State or other jurisdiction of<br><br>incorporation or organization) (I.R.S. Employer Identification Number)

26 North Euclid Avenue, Pasadena, California 91101

(Address of principal executive offices) (Zip code)

(626) 578-0777

(Registrant’s telephone number, including area code)

N/A

(Former name, former address and former fiscal year, if changed since last report)

Securities registered pursuant to Section 12(b) of the Exchange Act:

Title of each class Trading Symbol(s) Name of each exchange on which registered
Common Stock, $0.01 par value per share ARE New York Stock Exchange

Indicate by check mark whether the registrant (1) has filed all reports required to be filed by Section 13 or 15(d) of the Securities

Exchange Act of 1934 during the preceding 12 months (or for such shorter period that the registrant was required to file such reports),

and (2) has been subject to such filing requirements for the past 90 days. Yes ☒  No ☐

Indicate by check mark whether the registrant has submitted electronically every Interactive Data File required to be submitted and

posted pursuant to Rule 405 of Regulation S-T (§ 232.405 of this chapter) during the preceding 12 months (or for such shorter period

that the registrant was required to submit and post such files). Yes ☒  No  ☐

Indicate by check mark whether the registrant is a large accelerated filer, an accelerated filer, a non-accelerated filer, a smaller reporting

company, or an emerging growth company. See the definitions of “large accelerated filer,” “accelerated filer,” “smaller reporting

company,” and “emerging growth company” in Rule 12b-2 of the Exchange Act.

Large accelerated filer Smaller reporting company
Accelerated filer Emerging growth company
Non-accelerated filer

If an emerging growth company, indicate by check mark if the registrant has elected not to use the extended transition period for

complying with any new or revised financial accounting standards provided pursuant to Section 13(a) of the Exchange Act. ☐

Indicate by check mark whether the registrant is a shell company (as defined in Rule 12b-2 of the Exchange Act). Yes ☐  No ☒

As of April 15, 2026, 174,269,480 shares of common stock, par value $0.01 per share, were outstanding.

i

TABLE OF CONTENTS

Page
PART I – FINANCIAL INFORMATION
Item 1. FINANCIAL STATEMENTS (UNAUDITED)
Consolidated Balance Sheets as of March 31, 2026 and December 31, 2025 .......................................................... 1
Consolidated Financial Statements for the Three Months Ended March 31, 2026 and 2025:
Consolidated Statements of Operations ................................................................................................................... 2
Consolidated Statements of Comprehensive Income ............................................................................................ 3
Consolidated Statements of Changes in Stockholders’ Equity and Noncontrolling Interests .......................... 4
Consolidated Statements of Cash Flows ................................................................................................................ 6
Notes to Consolidated Financial Statements .................................................................................................................... 8
Item 2. MANAGEMENT’S DISCUSSION AND ANALYSIS OF FINANCIAL CONDITION AND RESULTS OF<br><br>OPERATIONS ........................................................................................................................................................................ 45
Item 3. QUANTITATIVE AND QUALITATIVE DISCLOSURES ABOUT MARKET RISK ......................................................... 116
Item 4. CONTROLS AND PROCEDURES ..................................................................................................................................... 117
PART II – OTHER INFORMATION
Item 1. LEGAL PROCEEDINGS ...................................................................................................................................................... 118
Item 1A. RISK FACTORS .................................................................................................................................................................... 118
Item 2. UNREGISTERED SALES OF EQUITY SECURITIES AND USE OF PROCEEDS ................................................... 118
Item 5. OTHER INFORMATION ....................................................................................................................................................... 118
Item 6. EXHIBITS ............................................................................................................................................................................... 119
SIGNATURES ................................................................................................................................................................................................. 120

ii

GLOSSARY

The following abbreviations or acronyms that may be used in this document

shall have the adjacent meanings set forth below:

ASU Accounting Standards Update
ATM At the Market
CAD Canadian Dollar
CIP Construction in Progress
EPS Earnings per Share
FASB Financial Accounting Standards Board
FFO Funds From Operations
GAAP U.S. Generally Accepted Accounting Principles
IRS Internal Revenue Service
JV Joint Venture
Nareit National Association of Real Estate Investment Trusts
NAV Net Asset Value
NYSE New York Stock Exchange
REIT Real Estate Investment Trust
RSF Rentable Square Feet/Foot
SEC Securities and Exchange Commission
SF Square Feet/Foot
SoDo South of Downtown submarket of Seattle
SOFR Secured Overnight Financing Rate
U.S. United States
USD U.S. Dollar
VIE Variable Interest Entity

1

PART I – FINANCIAL INFORMATION

ITEM 1. FINANCIAL STATEMENTS (UNAUDITED)

Alexandria Real Estate Equities, Inc.

Consolidated Balance Sheets

(In thousands)

March 31, 2026 December 31, 2025
(Unaudited)
Assets
Investments in real estate $28,830,116 $28,689,996
Investments in unconsolidated real estate joint ventures 30,520 30,677
Cash and cash equivalents 418,720 549,062
Restricted cash 4,665 4,693
Tenant receivables 7,362 6,672
Deferred rent 1,200,047 1,179,403
Deferred leasing costs 456,405 458,311
Investments 1,536,419 1,501,249
Other assets 1,683,143 1,661,772
Total assets $34,167,397 $34,081,835
Liabilities, Noncontrolling Interests, and Equity
Unsecured senior notes payable $11,166,009 $12,047,394
Unsecured senior line of credit and commercial paper 1,353,986 353,161
Accounts payable, accrued expenses, and other liabilities 2,154,782 2,397,073
Dividends payable 128,880 127,771
Total liabilities 14,803,657 14,925,399
Commitments and contingencies
Redeemable noncontrolling interests 9,234 58,788
Alexandria Real Estate Equities, Inc.’s stockholders’ equity:
Common stock 1,707 1,705
Additional paid-in capital 15,763,321 15,497,760
Accumulated other comprehensive loss (30,936) (29,395)
Alexandria Real Estate Equities, Inc.’s stockholders’ equity 15,734,092 15,470,070
Noncontrolling interests 3,620,414 3,627,578
Total equity 19,354,506 19,097,648
Total liabilities, noncontrolling interests, and equity $34,167,397 $34,081,835

The accompanying notes are an integral part of these consolidated financial statements.

2

Alexandria Real Estate Equities, Inc.

Consolidated Statements of Operations

(In thousands, except per share amounts)

(Unaudited)

Three Months Ended March 31,
2026 2025
Revenues:
Income from rentals $653,013 $743,175
Other income 18,009 14,983
Total revenues 671,022 758,158
Expenses:
Rental operations 224,142 226,395
General and administrative 34,685 30,675
Interest 64,584 50,876
Depreciation and amortization 305,441 342,062
Impairment of real estate 5,499 32,154
Total expenses 634,351 682,162
Equity in losses of unconsolidated real estate joint ventures (147) (507)
Investment loss (4,582) (49,992)
Gain on early extinguishment of debt 366,435
Gain on sales of real estate 13,165
Net income 398,377 38,662
Net income attributable to noncontrolling interests (36,724) (47,601)
Net income (loss) attributable to Alexandria Real Estate Equities, Inc.’s stockholders 361,653 (8,939)
Net income attributable to unvested restricted stock awards (2,779) (2,660)
Net income (loss) attributable to Alexandria Real Estate Equities, Inc.’s common stockholders $358,874 $(11,599)
Net income (loss) per share attributable to Alexandria Real Estate Equities, Inc.’s common<br><br>stockholders:
Basic $2.10 $(0.07)
Diluted $2.10 $(0.07)

The accompanying notes are an integral part of these consolidated financial statements.

3

Alexandria Real Estate Equities, Inc.

Consolidated Statements of Comprehensive Income

(In thousands)

(Unaudited)

Three Months Ended March 31,
2026 2025
Net income $398,377 $38,662
Other comprehensive (loss) income
Change in foreign currency translation adjustments:
Unrealized foreign currency translation (losses) gains arising during the period (1,518) 50
Reclassification of gains (23)
Unrealized (losses) gains on foreign currency translation, net (1,541) 50
Total other comprehensive (loss) income (1,541) 50
Comprehensive income 396,836 38,712
Less: comprehensive income attributable to noncontrolling interests (36,724) (47,601)
Comprehensive income (loss) attributable to Alexandria Real Estate Equities, Inc.’s stockholders $360,112 $(8,889)

The accompanying notes are an integral part of these consolidated financial statements.

4

Alexandria Real Estate Equities, Inc.

Consolidated Statement of Changes in Stockholders’ Equity and Noncontrolling Interests

(Dollars in thousands)

(Unaudited)

Alexandria Real Estate Equities, Inc.’s Stockholders’ Equity
Number of<br><br>Common<br><br>Shares Common<br><br>Stock Additional<br><br>Paid-In<br><br>Capital Retained<br><br>Earnings Accumulated<br><br>Other<br><br>Comprehensive<br><br>Loss Noncontrolling<br><br>Interests Total<br><br>Equity Redeemable<br><br>Noncontrolling<br><br>Interests
Balance as of December 31, 2025 170,537,867 $1,705 $15,497,760 $— $(29,395) $3,627,578 $19,097,648 $58,788
Net income 361,653 36,377 398,030 347
Total other comprehensive loss (1,541) (1,541)
Contributions from and sales of noncontrolling interests 7,079 16,377 23,456
Distributions to and redemption of noncontrolling interests (59,918) (59,918) (49,901)
Issuance pursuant to stock plan 289,485 3 28,262 28,265
Taxes related to net settlement of equity awards (115,062) (1) (5,960) (5,961)
Dividends declared on common stock ($0.72 per share) (125,473) (125,473)
Reclassification of earnings in excess of distributions 236,180 (236,180)
Balance as of March 31, 2026 170,712,290 $1,707 $15,763,321 $— $(30,936) $3,620,414 $19,354,506 $9,234

The accompanying notes are an integral part of these consolidated financial statements.

5

Alexandria Real Estate Equities, Inc.

Consolidated Statement of Changes in Stockholders’ Equity and Noncontrolling Interests

(Dollars in thousands)

(Unaudited)

Alexandria Real Estate Equities, Inc.’s Stockholders’ Equity
Number of<br><br>Common<br><br>Shares Common<br><br>Stock Additional<br><br>Paid-In<br><br>Capital Retained<br><br>Earnings Accumulated<br><br>Other<br><br>Comprehensive<br><br>Loss Noncontrolling<br><br>Interests Total<br><br>Equity Redeemable<br><br>Noncontrolling<br><br>Interests
Balance as of December 31, 2024 172,203,443 $1,722 $17,933,572 $— $(46,252) $4,489,447 $22,378,489 $19,972
Net (loss) income (8,939) 47,331 38,392 270
Total other comprehensive income 50 50
Contributions from and sales of noncontrolling interests 54 54,354 54,408
Distributions to and redemption of noncontrolling interests (7,048) (65,833) (72,881) (10,630)
Issuance pursuant to stock plan 125,280 1 32,755 32,756
Taxes related to net settlement of equity awards (46,547) (4,735) (4,735)
Repurchase of common stock (2,152,293) (22) (208,165) (208,187)
Dividends declared on common stock ($1.32 per share) (228,346) (228,346)
Reclassification of distributions and net loss (237,285) 237,285
Balance as of March 31, 2025 170,129,883 $1,701 $17,509,148 $— $(46,202) $4,525,299 $21,989,946 $9,612

The accompanying notes are an integral part of these consolidated financial statements.

6

Alexandria Real Estate Equities, Inc.<br><br>Consolidated Statements of Cash Flows<br><br>(In thousands)<br><br>(Unaudited)
Three Months Ended March 31,
2026 2025
Operating Activities:
Net income $398,377 $38,662
Adjustments to reconcile net income to net cash provided by operating activities:
Depreciation and amortization 305,441 342,062
Impairment of real estate 5,499 32,154
Gain on sales of real estate (13,165)
Gain on early extinguishment of debt (366,435)
Equity in losses of unconsolidated real estate joint ventures 147 507
Distributions of earnings from unconsolidated real estate joint ventures 233 172
Amortization of loan fees 4,428 4,691
Amortization of debt discounts 320 349
Amortization of acquired above- and below-market leases (5,615) (15,222)
Deferred rent (17,862) (22,023)
Stock compensation expense 11,032 10,064
Investment loss 4,582 49,992
Changes in operating assets and liabilities:
Tenant receivables (693) (467)
Deferred leasing costs (20,121) (26,645)
Other assets (35,274) (37,034)
Accounts payable, accrued expenses, and other liabilities (87,435) (156,148)
Net cash provided by operating activities 196,624 207,949
Investing Activities:
Proceeds from sales of real estate 68,182
Additions to real estate (545,999) (645,841)
Change in escrow deposits (9,506)
Investments in unconsolidated real estate joint ventures (297) (10,994)
Return of capital from unconsolidated real estate joint ventures 113
Additions to non-real estate investments (73,627) (69,311)
Sales of and distributions from non-real estate investments 35,000 12,691
Net cash used in investing activities $(584,810) $(654,779)

7

Alexandria Real Estate Equities, Inc.<br><br>Consolidated Statements of Cash Flows<br><br>(In thousands)<br><br>(Unaudited)
Three Months Ended March 31,
2026 2025
Financing Activities:
Borrowings under secured note payable $— $824
Repayments of borrowings under secured notes payable (8,892)
Proceeds from issuance of unsecured senior notes payable 747,592 548,532
Repayment of unsecured senior notes payable (1,252,203)
Proceeds from issuances under commercial paper program 12,319,811 2,700,000
Repayments of borrowings under commercial paper program (11,318,040) (2,400,000)
Payments of loan fees (8,814) (5,406)
Taxes paid related to net settlement of equity awards (5,946) (5,558)
Repurchase of common stock (208,187)
Dividends on common stock (123,752) (229,987)
Contributions from and sales of noncontrolling interests 18,065 54,409
Distributions to noncontrolling interests (60,111) (66,034)
Purchases and redemptions of noncontrolling interests (49,707) (17,818)
Net cash provided by financing activities 258,003 370,775
Effect of foreign exchange rate changes on cash and cash equivalents (187) (38)
Net decrease in cash, cash equivalents, and restricted cash (130,370) (76,093)
Cash, cash equivalents, and restricted cash as of the beginning of period 553,755 559,847
Cash, cash equivalents, and restricted cash as of the end of period $423,385 $483,754
Supplemental Disclosure and Non-Cash Investing and Financing Activities:
Cash paid during the period for interest, net of interest capitalized $56,403 $33,776
Accrued construction for current-period additions to real estate $137,633 $147,045
Transfer of real estate assets and/or equipment from tenants $2,694 $39,950
Notes receivable issued in connection with sales of real estate $— $91,000
Derecognition of net investment in real estate from sales-type lease $— $4,677
Acquisition of real estate and other assets in connection with assumption of related secured<br><br>notes payable of unconsolidated joint venture $8,892 $—

The accompanying notes are an integral part of these consolidated financial statements.

8

Alexandria Real Estate Equities, Inc.

Notes to Consolidated Financial Statements

(Unaudited)

1.ORGANIZATION AND BASIS OF PRESENTATION

Alexandria Real Estate Equities, Inc. (NYSE: ARE), an S&P 500® life science REIT, is the pioneer of the life science real estate

niche since its founding in 1994. Alexandria is the preeminent and longest-tenured owner, operator, and developer of collaborative

Megacampus™ ecosystems in AAA life science innovation cluster locations, including Greater Boston, the San Francisco Bay Area,

San Diego, Seattle, Maryland, Research Triangle, and New York City. As of March 31, 2026, Alexandria has a total market capitalization

of $20.44 billion and an asset base that includes 35.8 million RSF of operating properties and 3.4 million RSF of Class A/A+ properties

undergoing construction. As used in this quarterly report on Form 10-Q, references to the “Company,” “Alexandria,” “ARE,” “we,” “us,”

and “our” refer to Alexandria Real Estate Equities, Inc. and its consolidated subsidiaries. The accompanying unaudited consolidated

financial statements include the accounts of Alexandria Real Estate Equities, Inc. and its consolidated subsidiaries. All significant

intercompany balances and transactions have been eliminated.

We have prepared the accompanying interim consolidated financial statements in accordance with GAAP and in conformity

with the rules and regulations of the SEC. In our opinion, these interim consolidated financial statements presented herein reflect all

adjustments, of a normal recurring nature, that are necessary to fairly present the interim consolidated financial statements. The results

of operations for the interim period are not necessarily indicative of the results that may be expected for the year ending December 31,

  1. These unaudited consolidated financial statements should be read in conjunction with the audited consolidated financial

statements and the notes thereto included in our annual report on Form 10-K for the year ended December 31, 2025. Any references to

our total market capitalization, number or quality of buildings or tenants, quality of location, square footage, number of leases, or

occupancy percentage, and any amounts derived from these values in these notes to consolidated financial statements are outside the

scope of our independent registered public accounting firm’s procedures.

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES

Consolidation

On an ongoing basis, as circumstances indicate the need for reconsideration, we evaluate each legal entity that is not wholly

owned by us in accordance with the consolidation accounting guidance. Our evaluation considers all of our variable interests, including

equity ownership, as well as fees paid to us for our involvement in the management of each partially owned entity. To fall within the

scope of the consolidation guidance, an entity must meet both of the following criteria:

•The entity has a legal structure that has been established to conduct business activities and to hold assets; such entity

can be in the form of a partnership, limited liability company, or corporation, among others; and

•We have a variable interest in the legal entity — i.e., variable interests that are contractual, such as equity ownership, or

other financial interests that change with changes in the fair value of the entity’s net assets.

If an entity does not meet both criteria above, we apply other accounting literature, such as the equity method of accounting. If

an entity does meet both criteria above, we evaluate such entity for consolidation under either the variable interest model if the legal

entity meets any of the characteristics below to qualify as a VIE, or under the voting model for all other legal entities that are not VIEs.

A legal entity is determined to be a VIE if it has any of the following three characteristics:

1)The entity does not have sufficient equity to finance its activities without additional subordinated financial support;

2)The entity is established with non-substantive voting rights (i.e., the entity deprives the majority economic interest

holder(s) of voting rights); or

3)The equity holders, as a group, lack the characteristics of a controlling financial interest. Equity holders meet this criterion

if they lack any of the following:

•The power, through voting rights or similar rights, to direct the activities of the entity that most significantly influence

the entity’s economic performance, as evidenced by:

•Substantive participating rights in day-to-day management of the entity’s activities; or

•Substantive kick-out rights over the party responsible for significant decisions;

•The obligation to absorb the entity’s expected losses; or

•The right to receive the entity’s expected residual returns.

9

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

For an entity, including our real estate joint ventures, structured as a limited partnership or a limited liability company, our

evaluation of whether the equity holders (equity partners other than the general partner or the managing member of a joint venture) lack

the characteristics of a controlling financial interest includes the evaluation of whether the limited partners or non-managing members

(the noncontrolling equity holders) lack both substantive participating rights and substantive kick-out rights, defined as follows:

•Participating rights provide the noncontrolling equity holders the ability to direct significant financial and operating

decisions made in the ordinary course of business that most significantly influence the entity’s economic performance.

•Kick-out rights allow the noncontrolling equity holders to remove the general partner or managing member without cause.

If we conclude that any of the three characteristics of a VIE are met, including that the equity holders lack the characteristics of

a controlling financial interest because they lack both substantive participating rights and substantive kick-out rights, we conclude that

the entity is a VIE and evaluate it for consolidation under the variable interest model.

Variable interest model

If an entity is determined to be a VIE, we evaluate whether we are the primary beneficiary. The primary beneficiary analysis is

a qualitative analysis based on power and benefits. We consolidate a VIE if we have both power and benefits — that is, (i) we have the

power to direct the activities of a VIE that most significantly influence the VIE’s economic performance (power) and (ii) we have the

obligation to absorb losses of or the right to receive benefits from the VIE that could potentially be significant to the VIE (benefits). We

consolidate VIEs whenever we determine that we are the primary beneficiary. Refer to Note 4 – “Consolidated and unconsolidated real

estate joint ventures” and Note 7 – “Investments” to our unaudited consolidated financial statements for information on specific entities

that qualify as VIEs. If we have a variable interest in a VIE but are not the primary beneficiary, we account for our investment using the

equity method.

Voting model

If a legal entity fails to meet any of the three characteristics of a VIE (i.e., insufficiency of equity, existence of non-substantive

voting rights, or lack of a controlling financial interest), we then evaluate such entity under the voting model. Under the voting model, we

consolidate the entity if we determine that we, directly or indirectly, have greater than 50% of the voting shares (or own a majority of the

limited partnership’s kick-out rights through voting interests), and that other equity holders do not have substantive participating rights.

Refer to Note 4 – “Consolidated and unconsolidated real estate joint ventures” to our unaudited consolidated financial statements for

information on specific joint ventures that qualify for evaluation under the voting model.

Noncontrolling interests in consolidated real estate joint ventures

Noncontrolling interests represent the third-party interests in consolidated real estate joint ventures in which we have a

controlling interest. Certain of our partners’ noncontrolling interests have the right to require us to redeem their ownership interests in

the respective entities. We classify the ownership interests in these entities as redeemable noncontrolling interests outside of total

equity in our consolidated balance sheets. Redeemable noncontrolling interests are adjusted for additional contributions and

distributions, the proportionate share of net earnings or losses, and other comprehensive income or loss. If the amount of a redeemable

noncontrolling interest is less than the maximum redemption value at the balance sheet date, such amount is adjusted to the maximum

redemption value. Subsequent declines in the redemption value are recognized only to the extent that previous increases have been

recognized.

Use of estimates

The preparation of consolidated financial statements in conformity with GAAP requires us to make estimates and assumptions

that affect the reported amounts of assets, liabilities, and equity; the disclosure of contingent assets and liabilities as of the date of the

consolidated financial statements; and the amounts of revenues and expenses during the reporting period. Actual results could

materially differ from those estimates.

10

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

Investments in real estate

Evaluation of business combination or asset acquisition

We evaluate each acquisition of real estate or in-substance real estate (including equity interests in entities that predominantly

hold real estate assets) to determine whether the integrated set of assets and activities acquired meets the definition of a business and

needs to be accounted for as a business combination. An acquisition of an integrated set of assets and activities that does not meet the

definition of a business is accounted for as an asset acquisition. If either of the following criteria is met, the integrated set of assets and

activities acquired would not qualify as a business:

•Substantially all of the fair value of the gross assets acquired is concentrated in either a single identifiable asset or a group

of similar identifiable assets; or

•The integrated set of assets and activities is lacking, at a minimum, an input and a substantive process that together

significantly contribute to the ability to create outputs (i.e., revenue generated before and after the transaction).

An acquired process is considered substantive if:

•The process includes an organized workforce (or includes an acquired contract that provides access to an organized

workforce) that is skilled, knowledgeable, and experienced in performing the process;

•The process cannot be replaced without significant cost, effort, or delay; or

•The process is considered unique or scarce.

Generally, our acquisitions of real estate or in-substance real estate do not meet the definition of a business because

substantially all of the fair value is concentrated in a single identifiable asset or group of similar identifiable assets (i.e., land, buildings,

and related intangible assets) or because the acquisition does not include a substantive process in the form of an acquired workforce or

an acquired contract that cannot be replaced without significant cost, effort, or delay. When evaluating acquired service or management

contracts, we consider the nature of the services performed, the terms of the contract relative to similar arm’s-length contracts, and the

availability of comparable vendors in evaluating whether the acquired contract constitutes a substantive process.

Recognition of real estate acquired

We evaluate each acquisition of real estate or in-substance real estate (including equity interests in entities that predominantly

hold real estate assets) to determine whether the integrated set of assets and activities acquired meets the definition of a business and

needs to be accounted for as a business combination. An acquisition of an integrated set of assets and activities that does not meet the

definition of a business is accounted for as an asset acquisition.

For acquisitions of real estate or in-substance real estate that are accounted for as business combinations, we allocate the

acquisition consideration (excluding acquisition costs) to the assets acquired, liabilities assumed, noncontrolling interests, and

previously existing ownership interests at fair value as of the acquisition date. Assets include intangible assets such as tenant

relationships, acquired in-place leases, and favorable intangibles associated with in-place leases in which we are the lessor. Liabilities

include unfavorable intangibles associated with in-place leases in which we are the lessor. In addition, for acquired in-place finance or

operating leases in which we are the lessee, acquisition consideration is allocated to lease liabilities and related right-of-use assets,

adjusted to reflect favorable or unfavorable terms of the lease when compared with market terms. Any excess (deficit) of the

consideration transferred relative to the fair value of the net assets acquired is accounted for as goodwill (bargain purchase gain).

Acquisition costs related to business combinations are expensed as incurred.

Generally, we expect that acquisitions of real estate or in-substance real estate will not meet the definition of a business

because substantially all of the fair value is concentrated in a single identifiable asset or group of similar identifiable assets (i.e., land,

buildings, and related intangible assets). The accounting model for asset acquisitions is similar to the accounting model for business

combinations, except that the acquisition consideration (including acquisition costs) is allocated to the individual assets acquired and

liabilities assumed on a relative fair value basis. Any excess (deficit) of the consideration transferred relative to the sum of the fair value

of the assets acquired and liabilities assumed is allocated to the individual assets and liabilities based on their relative fair values. As a

result, asset acquisitions do not result in the recognition of goodwill or a bargain purchase gain. Incremental and external direct

acquisition costs related to acquisitions of real estate or in-substance real estate (such as legal and other third-party services) are

capitalized.

We exercise judgment to determine the key assumptions used to allocate the purchase price of real estate acquired among its

components. The allocation of the consideration to the various components of properties acquired during the year can have an effect on

our net income due to the useful depreciable and amortizable lives applicable to each component and the recognition of the related

depreciation and amortization expense in our consolidated statements of operations. We apply judgment in utilizing available

comparable market information to assess relative fair value. We assess the relative fair values of tangible and intangible assets and

liabilities based on available comparable market information, including estimated replacement costs, rental rates, and recent market

transactions. In addition, we may use estimated cash flow projections that utilize appropriate discount and capitalization rates.

11

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

Estimates of future cash flows are based on a number of factors, including the historical operating results, known and anticipated

trends, and market/economic conditions that may affect the property.

The value of tangible assets acquired is based upon our estimation of fair value on an “as if vacant” basis. The value of

acquired in-place leases includes the estimated costs during the hypothetical lease-up period and other costs that would have been

incurred in the execution of similar leases under the market conditions at the acquisition date of the acquired in-place lease. If there is a

bargain fixed-rate renewal option for the period beyond the noncancelable lease term of an in-place lease, we evaluate intangible

factors, such as the business conditions in the industry in which the lessee operates, the economic conditions in the area in which the

property is located, and the ability of the lessee to sublease the property during the renewal term, in order to determine the likelihood

that the lessee will renew. When we determine that there is reasonable assurance that such bargain purchase option will be exercised,

we consider the option in determining the intangible value of such lease and its related amortization period. We also recognize the

relative fair values of assets acquired, the liabilities assumed, and any noncontrolling interest in acquisitions of less than a 100%

interest when the acquisition constitutes a change in control of the acquired entity.

Depreciation and amortization

The values allocated to buildings and building improvements, land improvements, tenant improvements, and equipment are

depreciated on a straight-line basis. For buildings and building improvements, we depreciate using the shorter of the respective ground

lease terms or their estimated useful lives, not to exceed 40 years. Land improvements are depreciated over their estimated useful

lives, not to exceed 20 years. Tenant improvements are depreciated over their respective lease terms or estimated useful lives, and

equipment is depreciated over the shorter of the lease term or its estimated useful life. The values of the right-of-use assets are

amortized on a straight-line basis over the remaining terms of each related lease. The values of acquired in-place leases and

associated favorable intangibles (i.e., acquired above-market leases) are classified in other assets in our consolidated balance sheets

and are amortized over the remaining terms of the related leases as a reduction of income from rentals in our consolidated statements

of operations. The values of unfavorable intangibles (i.e., acquired below-market leases) associated with acquired in-place leases are

classified in accounts payable, accrued expenses, and other liabilities in our consolidated balance sheets and are amortized over the

remaining terms of the related leases as an increase in income from rentals in our consolidated statements of operations.

Capitalized project costs

We capitalize project costs, including pre-construction costs, interest, property taxes, insurance, and other costs directly

related and essential to the development, redevelopment, pre-construction, or construction of a project. Capitalization of development,

redevelopment, pre-construction, and construction costs is required while activities are ongoing to prepare an asset for its intended use.

Fluctuations in our development, redevelopment, pre-construction, and construction activities could result in significant changes to total

expenses and net income. Costs incurred after a project is substantially complete and ready for its intended use are expensed as

incurred. Should development, redevelopment, pre-construction, or construction activity cease, interest, property taxes, insurance, and

certain other costs would no longer be eligible for capitalization and would be expensed as incurred. Expenditures for repairs and

maintenance are expensed as incurred.

Real estate sales

A property is classified as held for sale when all of the following criteria for a plan of sale have been met: (i) management,

having the authority to approve the action, commits to a plan to sell the property; (ii) the property is available for immediate sale in its

present condition, subject only to terms that are usual and customary; (iii) an active program to locate a buyer and other actions

required to complete the plan to sell have been initiated; (iv) the sale of the property is probable and is expected to be completed within

one year; (v) the property is being actively marketed for sale at a price that is reasonable in relation to its current fair value; and

(vi) actions necessary to complete the plan of sale indicate that it is unlikely that significant changes to the plan will be made or that the

plan will be withdrawn. Depreciation of assets ceases upon designation of a property as held for sale.

If the disposal of a property represents a strategic shift that has (or will have) a major effect on our operations or financial

results, such as (i) a major line of business, (ii) a major geographic area, (iii) a major equity method investment, or (iv) other major parts

of an entity, then the operations of the property, including any interest expense directly attributable to it, are classified as discontinued

operations in our consolidated statements of operations, and amounts for all prior periods presented are reclassified from continuing

operations to discontinued operations. The disposal of an individual property generally will not represent a strategic shift and therefore

will typically not meet the criteria for classification as a discontinued operation.

We recognize gains or losses on real estate sales in accordance with the accounting standard on the derecognition of

nonfinancial assets arising from contracts with noncustomers. Our ordinary output activities consist of the leasing of space to our

tenants in our operating properties, not the sales of real estate. Therefore, sales of real estate (in which we are the seller) qualify as

contracts with noncustomers. In our transactions with noncustomers, we apply certain recognition and measurement principles

consistent with our method of recognizing revenue arising from contracts with customers. Derecognition of the asset is based on the

transfer of control. If a real estate sales contract includes our ongoing involvement with the property, then we evaluate each promised

good or service under the contract to determine whether it represents a separate performance obligation, constitutes a guarantee, or

12

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

prevents the transfer of control. If a good or service is considered a separate performance obligation, an allocated portion of the

transaction price is recognized as revenue as we transfer the related good or service to the buyer.

The recognition of gain or loss on the sale of a partial interest also depends on whether we retain a controlling or

noncontrolling interest in the property. If we retain a controlling interest in the property upon completion of the sale, we continue to

reflect the asset at its book value, record a noncontrolling interest for the book value of the partial interest sold, and recognize additional

paid-in capital for the difference between the consideration received and the partial interest at book value. Conversely, if we retain a

noncontrolling interest upon completion of the sale of a partial interest of real estate, we recognize a gain or loss as if 100% of the asset

were sold.

Impairment of long-lived assets

Prior to and subsequent to the end of each quarter, we review current activities and changes in the business conditions of all of

our long-lived assets to determine the existence of any triggering events or impairment indicators requiring an impairment analysis. If

triggering events or impairment indicators are identified, we review an estimate of the future undiscounted cash flows, including, if

necessary, a probability-weighted approach if multiple outcomes are under consideration.

Long-lived assets to be held and used, including our rental properties, CIP, land held for development, right-of-use assets

related to operating leases in which we are the lessee, and intangibles, are individually evaluated for impairment when conditions exist

that may indicate that the carrying amount of a long-lived asset may not be recoverable. The carrying amount of a long-lived asset to be

held and used is not recoverable if it exceeds the sum of the undiscounted cash flows expected to result from the use and eventual

disposition of the asset. Triggering events or impairment indicators for long-lived assets to be held and used are assessed by project

and include significant fluctuations in estimated net operating income, occupancy changes, significant near-term lease expirations,

current and historical operating and/or cash flow losses, construction costs, estimated completion dates, rental rates, and other market

factors. We assess the expected undiscounted cash flows based upon numerous factors, including, but not limited to, projected rental

rates, estimated exit capitalization rates, and anticipated construction costs for projects under construction, which are based on

available market information, current and historical operating results, known trends, current market/economic conditions that may affect

the asset, and our assumptions about the use of the asset, including, if necessary, a probability-weighted approach if multiple outcomes

are under consideration.

Upon determination that an impairment has occurred, a write-down is recognized to reduce the carrying amount of the asset to

its estimated fair value. If an impairment charge is not required to be recognized, the recognition of depreciation or amortization is

adjusted prospectively, as necessary, to reduce the carrying amount of the asset to its estimated disposition value over the remaining

period that the asset is expected to be held and used. We may adjust depreciation of properties that are expected to be disposed of or

redeveloped prior to the end of their useful lives.

We use the held for sale impairment model for our properties classified as held for sale, which is different from the held and

used impairment model. Under the held for sale impairment model, an impairment charge is recognized if the carrying amount of the

long-lived asset classified as held for sale exceeds its fair value less cost to sell. Because of these two different models, it is possible for

a long-lived asset previously classified as held and used to require the recognition of an impairment charge upon classification as held

for sale.

International operations

As of March 31, 2026, in addition to operating properties in the U.S., we had 11 properties in Canada. The functional currency

for our subsidiaries operating in the U.S. is the U.S. dollar. The local currency of a foreign subsidiary serves as its functional currency.

The assets and liabilities of our foreign subsidiaries are translated into U.S. dollars at the exchange rate in effect as of the financial

statement date. Revenue and expense accounts of our foreign subsidiaries are translated using the weighted-average exchange rate

for the periods presented. Gains or losses resulting from the translation are classified in accumulated other comprehensive income

(loss) as a separate component of total equity and are excluded from net income (loss).

Whenever a foreign investment meets the criteria for classification as held for sale, we evaluate the recoverability of the

investment under the held for sale impairment model. We may recognize an impairment charge if the carrying amount of the investment

exceeds its fair value less cost to sell. In determining an investment’s carrying amount, we consider its net book value and any

cumulative unrealized foreign currency translation adjustment related to the investment. The appropriate amounts of foreign exchange

rate gains or losses classified in accumulated other comprehensive income (loss) are reclassified to net income (loss) when realized

upon the sale of our investment or upon the complete or substantially complete liquidation of our investment.

13

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

Investments

We hold investments in publicly traded companies and privately held entities primarily involved in the life science industry. As a

REIT, we generally limit our ownership of each individual entity’s voting stock to less than 10%. We evaluate each investment to

determine whether we have the ability to exercise significant influence, but not control, over an investee. We evaluate investments in

which our ownership is equal to or greater than 20%, but less than or equal to 50%, of an investee’s voting stock with a presumption

that we have this ability. For our investments in limited partnerships that maintain specific ownership accounts, we presume that such

ability exists when our ownership interest exceeds 3% to 5%. In addition to our ownership interest, we consider whether we have a

board seat or whether we participate in the investee’s policymaking process, among other criteria, to determine if we have the ability to

exert significant influence, but not control, over an investee. If we determine that we have such ability, we account for the investment

under the equity method, as described below. From time to time, we may hold equity investments in publicly traded companies that are

subject to temporary contractual sale restrictions. We do not recognize a discount related to a contractual sale restriction.

Investments accounted for under the equity method

Under the equity method of accounting, we initially recognize our investment at cost and subsequently adjust the carrying

amount of the investment for our share of earnings or losses reported by the investee, distributions received, and other-than-temporary

impairments. For additional information about our investments accounted for under the equity method, refer to Note 7 – “Investments” to

our unaudited consolidated financial statements.

Investments that do not qualify for the equity method of accounting

For investees over which we determine that we do not have the ability to exercise significant influence or control, we account

for each investment depending on whether it is an investment in a (i) publicly traded company, (ii) privately held entity that reports NAV

per share, or (iii) privately held entity that does not report NAV per share, as described below.

Investments in publicly traded companies

Our investments in publicly traded companies are classified as investments with readily determinable fair values and are

presented at fair value in our consolidated balance sheets, with changes in fair value classified in investment income (loss) in our

consolidated statements of operations. The fair values for our investments in publicly traded companies are determined based on sales

prices or quotes available on securities exchanges.

Investments in privately held companies

Our investments in privately held entities without readily determinable fair values consist of (i) investments in privately held

entities that report NAV per share and (ii) investments in privately held entities that do not report NAV per share. These investments are

accounted for as follows:

Investments in privately held entities that report NAV per share

Investments in privately held entities that report NAV per share, such as our privately held investments in limited partnerships,

are presented at fair value using NAV as a practical expedient, with changes in fair value classified in investment income (loss) in our

consolidated statements of operations. We use NAV per share reported by limited partnerships generally without adjustment, unless we

are aware of information indicating that the NAV reported by a limited partnership does not accurately reflect the fair value of the

investment at our reporting date.

Investments in privately held entities that do not report NAV per share

Investments in privately held entities that do not report NAV per share are accounted for using a measurement alternative

under which these investments are measured at cost, adjusted for observable price changes and impairments, with changes classified

in investment income (loss) in our consolidated statements of operations.

An observable price arises from an orderly transaction for an identical or similar investment of the same issuer, which is

observed by an investor without expending undue cost and effort. Observable price changes result from, among other things, equity

transactions of the same issuer executed during the reporting period, including subsequent equity offerings or other reported equity

transactions related to the same issuer. To determine whether these transactions are indicative of an observable price change, we

evaluate, among other factors, whether these transactions have similar rights and obligations, including voting rights, distribution

preferences, and conversion rights to the investments we hold.

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2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

Impairment evaluation of equity method investments and investments in privately held entities that do not report NAV per share

We monitor equity method investments and investments in privately held entities that do not report NAV per share for new

developments, including operating results, prospects and results of clinical trials, new product initiatives, new collaborative agreements,

capital-raising events, and merger and acquisition activities. These investments are evaluated on the basis of a qualitative assessment

for indicators of impairment by monitoring the presence of the following triggering events or impairment indicators:

(i)a significant deterioration in the earnings performance, credit rating, asset quality, or business prospects of the investee;

(ii)a significant adverse change in the regulatory, economic, or technological environment of the investee;

(iii)a significant adverse change in the general market condition, including the research and development of technology and

products that the investee is bringing or attempting to bring to the market;

(iv)significant concerns about the investee’s ability to continue as a going concern; and/or

(v)a decision by investors to cease providing support or reduce their financial commitment to the investee.

If such indicators are present, we are required to estimate the investment’s fair value and immediately recognize an

impairment charge in an amount equal to the investment’s carrying value in excess of its estimated fair value.

Investment income/loss recognition and classification

We recognize both realized and unrealized gains and losses in our consolidated statements of operations, classified in

investment income (loss) in our consolidated statements of operations. Unrealized gains and losses represent:

(i)changes in fair value for investments in publicly traded companies;

(ii)changes in NAV for investments in privately held entities that report NAV per share;

(iii)observable price changes for investments in privately held entities that do not report NAV per share; and

(iv)our share of unrealized gains or losses reported by our equity method investees.

Realized gains and losses on our investments represent the difference between proceeds received upon disposition of

investments and their historical or adjusted cost basis. For our equity method investments, realized gains and losses represent our

share of realized gains or losses reported by the investee. Impairments are realized losses, which result in an adjusted cost basis, and

represent charges to reduce the carrying values of investments in privately held entities that do not report NAV per share and equity

method investments, if impairments are deemed other than temporary, to their estimated fair value.

Revenues

The table below provides details of our consolidated total revenues for the three months ended March 31, 2026 and 2025

(in thousands):

Three Months Ended March 31,
2026 2025
Income from rentals:
Revenues subject to the lease accounting standard:
Operating leases $640,659 $731,421
Direct financing and sales-type leases 964 810
Revenues subject to the lease accounting standard 641,623 732,231
Revenues subject to the revenue recognition accounting standard 11,390 10,944
Income from rentals 653,013 743,175
Other income 18,009 14,983
Total revenues $671,022 $758,158

During the three months ended March 31, 2026 and 2025, revenues that were subject to the lease accounting standard

aggregated $641.6 million, or 95.6%, and $732.2 million, or 96.6%, of our total revenue, respectively. Our other income consisted

primarily of management fees and interest income earned during each period presented. For a detailed discussion related to our

revenue streams, refer to “Lease accounting” and “Recognition of revenue arising from contracts with customers” in Note 2 – “Summary

of significant accounting policies” to our unaudited consolidated financial statements.

15

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

Lease accounting

Definition and classification of a lease

When we enter into a contract or amend an existing contract, we evaluate whether the contract meets the definition of a lease.

To meet the definition of a lease, the contract must meet all three criteria:

(i)One party (lessor) must hold an identified asset;

(ii)The counterparty (lessee) must have the right to obtain substantially all of the economic benefits from the use of the asset

throughout the period of the contract; and

(iii)The counterparty (lessee) must have the right to direct the use of the identified asset throughout the period of the contract.

We classify our leases as either finance leases or operating leases if we are the lessee, or sales-type, direct financing, or

operating leases if we are the lessor. We use the following criteria to determine if a lease is a finance lease (as a lessee) or sales-type

or direct financing lease (as a lessor):

(i)Ownership is transferred from lessor to lessee by the end of the lease term;

(ii)An option to purchase is reasonably certain to be exercised;

(iii)The lease term is for the major part of the underlying asset’s remaining economic life;

(iv)The present value of lease payments equals or exceeds substantially all of the fair value of the underlying asset; or

(v)The underlying asset is specialized and is expected to have no alternative use at the end of the lease term.

If we meet any of the above criteria, we account for the lease as a finance, a sales-type, or a direct financing lease. If we do

not meet any of the criteria, we account for the lease as an operating lease.

A lease is accounted for as a sales-type lease if it is considered to transfer control of the underlying asset to the lessee. A

lease is accounted for as a direct financing lease if risks and rewards are conveyed without the transfer of control, which is normally

indicated by the existence of a residual value guarantee from an unrelated third party other than the lessee.

This classification will determine the method of recognition of the lease:

•For an operating lease, we recognize income from rentals if we are the lessor, or rental operations expense if we are the

lessee, over the term of the lease on a straight-line basis.

•For a sales-type lease or a direct financing lease, we recognize the income from rentals, or for a finance lease, we

recognize rental operations expense, over the term of the lease using the effective interest method.

•At inception of a sales-type lease or a direct financing lease, if we determine the fair value of the leased property is lower

than its carrying amount, we recognize a selling loss immediately at lease commencement. If fair value exceeds the

carrying amount of a lease, a gain is recognized at lease commencement on a sales-type lease. For a direct financing

lease, a gain is deferred at lease commencement and amortized over the lease term.

Lessor accounting

Costs to execute leases

We capitalize initial direct costs, which represent only incremental costs to execute a lease that would not have been incurred

if the lease had not been obtained. Costs that we incur to negotiate or arrange a lease, regardless of its outcome, such as for fixed

employee compensation, tax or legal advice to negotiate lease terms, and other costs, are expensed as incurred.

Operating leases

We account for the revenue from our lease contracts by utilizing the single component accounting policy. This policy requires

us to account for, by class of underlying asset, the lease component and nonlease component(s) associated with each lease as a single

component if two criteria are met:

(i)The timing and pattern of transfer of the lease component and the nonlease component(s) are the same; and

(ii)The lease component would be classified as an operating lease if it were accounted for separately.

Lease components consist primarily of fixed rental payments, which represent scheduled rental amounts due under our

leases, and contingent rental payments. Nonlease components consist primarily of tenant recoveries representing reimbursements of

rental operating expenses under our triple net lease structure, including recoveries for property taxes, insurance, utilities, repairs and

maintenance, and common area expenses.

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2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

If the lease component is the predominant component, we account for all revenues under such lease as a single component in

accordance with the lease accounting standard. Conversely, if the nonlease component is the predominant component, all revenues

under such lease are accounted for in accordance with the revenue recognition accounting standard. Our operating leases qualify for

the single component accounting, and the lease component in each of our leases is predominant. Therefore, we account for all

revenues from our operating leases under the lease accounting standard and classify these revenues as income from rentals in our

consolidated statements of operations.

We commence recognition of income from rentals related to the operating leases at the date the property is ready for its

intended use by the tenant and the tenant takes possession or controls the physical use of the leased asset. When a lease includes

construction of improvements, we determine whether the improvements are landlord or tenant assets. In determining if the

improvements are landlord or tenant improvements, we consider various factors, including, but not limited to, the following:

•Which party retains legal title to the improvements upon lease expiration;

•Whether the improvements are expected to have significant residual value at the end of the lease term;

•Whether the improvements are unique to the tenant;

•What happens to the improvements upon lease expiration (i.e., whether they are removed or preserved for the landlord);

•Which party bears all costs of the improvements (including the risk of cost overruns); and

•Which party supervises the construction of the improvements.

If the improvements are landlord assets, we capitalize such improvements. If the improvements are tenant assets, we do not

capitalize these assets. Improvements that qualify as tenant assets, if funded by us, are accounted for as lease incentives and

amortized as a reduction of revenue over the term of the lease. If the tenant funds improvements without reimbursement from us, and

we determine these improvements to be landlord assets, we consider the amount associated with the improvements to be non-cash

lease payments, which are recognized as incremental revenue over the term of the lease.

Income from rentals related to fixed rental payments under operating leases is recognized on a straight-line basis over the

respective operating lease terms. We classify amounts expected to be received in later periods as deferred rent in our consolidated

balance sheets. Amounts received currently but recognized as revenue in future periods are classified in accounts payable, accrued

expenses, and other liabilities in our consolidated balance sheets.

Income from rentals related to variable payments includes tenant recoveries and contingent rental payments. Tenant

recoveries, including reimbursements of utilities, repairs and maintenance, common area expenses, real estate taxes and insurance,

and other operating expenses, are recognized as revenue in the period during which the applicable expenses are incurred and the

tenant’s obligation to reimburse us arises. Income from rentals related to other variable payments is recognized when associated

contingencies are removed.

We assess collectibility from our tenants of future lease payments for each of our operating leases. If we determine that

collectibility is probable, we recognize income from rentals based on the methodology described above. If we determine that

collectibility is not probable, we recognize an adjustment to lower our income from rentals. Furthermore, we may recognize a general

allowance at a portfolio level (not the individual level) if we do not expect to collect future lease payments in full.

For each lease for which we determine that collectibility of future lease payments is not probable, we cease the recognition of

income from rentals on a straight-line basis and limit the recognition of income to the lesser of payments collected from the lessee or

lease income that would have been recognized on a straight-line basis. We do not resume straight-line recognition of income from

rentals for these leases until we determine that the collectibility of future payments related to these leases is probable. We also record a

general allowance related to the deferred rent balances that at the portfolio level (not the individual level) are not expected to be

collected in full through the lease term. As of March 31, 2026 and December 31, 2025, our general allowance balance aggregated

$14.3 million and $14.3 million, respectively.

Direct financing and sales-type leases

Income from rentals related to direct financing and sales-type leases is recognized over the lease term using the effective

interest rate method. At lease commencement, we derecognize the underlying asset classified within investments in real estate and

record net investment in a lease within other assets in our consolidated balance sheets. This initial net investment is determined by

aggregating the present values of the total future lease payments and the estimated residual value of the property, less any unearned

income related to a direct financing lease. Over the lease term, the investment in the lease accretes in value, producing a constant

periodic rate of return on the net investment in the lease. Income from these leases is classified in income from rentals in our

consolidated statements of operations. Our net investment is reduced over time as lease payments are received.

We evaluate our net investment in direct financing and sales-type leases for impairment under the current expected credit

losses accounting standard. For additional information, refer to “Provision for expected credit losses” in Note 2 – “Summary of

significant accounting policies” to our unaudited consolidated financial statements.

17

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

As a lessor, we classify a lease with variable lease payments that do not depend on an index or a rate as an operating lease

on the commencement date of the lease if both of the following criteria are met:

(i)The lease would have been classified as a sales-type lease or direct financing lease under the current lease accounting

standard; and

(ii)The sales-type lease or direct financing lease classification would have resulted in a selling loss at lease commencement.

We do not derecognize the underlying asset and do not recognize a loss upon lease commencement but continue to

depreciate the underlying asset over its useful life.

Lessee accounting

We have operating lease agreements in which we are the lessee consisting of ground and office leases. At the lease

commencement date (or at the acquisition date if the lease is acquired as part of a real estate acquisition), we are required to recognize

a liability to account for our future obligations under these operating leases, and a corresponding right-of-use asset.

The lease liability is measured based on the present value of the future lease payments, including payments during the term

under our extension options that we are reasonably certain to exercise. The present value of the future lease payments is calculated for

each operating lease using each respective remaining lease term and a corresponding estimated incremental borrowing rate, which is

the interest rate that we estimate we would have to pay to borrow on a collateralized basis over a similar term for an amount equal to

the lease payments. Subsequently, the lease liability is accreted by applying a discount rate established at the lease commencement

date to the lease liability balance as of the beginning of the period and is reduced by the payments made during the period. We classify

the operating lease liability in accounts payable, accrued expenses, and other liabilities in our consolidated balance sheets.

The right-of-use asset is measured based on the corresponding lease liability, adjusted for initial direct leasing costs and any

other consideration exchanged with the landlord prior to the commencement of the lease, as well as adjustments to reflect favorable or

unfavorable terms of an acquired lease when compared with market terms at the time of acquisition. Subsequently, the right-of-use

asset is amortized on a straight-line basis during the lease term. We classify the right-of-use asset in other assets in our consolidated

balance sheets.

Recognition of revenue arising from contracts with customers

We recognize revenues associated with transactions arising from contracts with customers, excluding revenues subject to the

lease accounting standard discussed in “Lease accounting” above, in accordance with the revenue recognition accounting standard. A

customer is distinguished from a noncustomer by the nature of the goods or services that are transferred. Customers are provided with

goods or services that are generated by a company’s ordinary output activities, whereas noncustomers are provided with nonfinancial

assets that are outside of a company’s ordinary output activities.

We generally recognize revenue representing the transfer of goods and services to customers in an amount that reflects the

consideration to which we expect to be entitled in the exchange. In order to determine the recognition of revenue from customer

contracts, we use a five-step model to (i) identify the contract with the customer, (ii) identify the performance obligations in the contract,

(iii) determine the transaction price, including variable consideration to the extent that it is probable that a significant future reversal will

not occur, (iv) allocate the transaction price to the performance obligations in the contract, and (v) recognize revenue when (or as) we

satisfy the performance obligation.

We identify contractual performance obligations and determine whether revenue should be recognized at a point in time or

over time, based on when control of goods and services transfers to a customer. We consider whether we control the goods or services

prior to the transfer to the customer in order to determine whether we should account for the arrangement as a principal or agent. If we

determine that we control the goods or services provided to the customer, then we are the principal to the transaction, and we recognize

the gross amount of consideration expected in the exchange. If we simply arrange but do not control the goods or services being

transferred to the customer, then we are considered to be an agent to the transaction, and we recognize the net amount of

consideration we are entitled to retain in the exchange.

Total revenues subject to the revenue recognition accounting standard and classified within income from rentals in our

consolidated statements of operations for the three months ended March 31, 2026 included $11.4 million primarily related to short-term

parking revenues associated with long-term lease agreements. Short-term parking revenues do not qualify for the single component

accounting policy, as discussed in “Lessor accounting” in Note 2 – “Summary of significant accounting policies,” due to the difference in

the timing and pattern of transfer of our parking service obligations and associated lease components within the same lease agreement.

We recognize short-term parking revenues in accordance with the revenue recognition accounting standard when the service is

provided and the performance obligation is satisfied, which normally occurs at a point in time.

18

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

Monitoring of tenant credit quality

During the term of each lease, we monitor the credit quality and any related material changes of our tenants by (i) monitoring

the credit rating of tenants that are rated by a nationally recognized credit rating agency, (ii) reviewing financial statements of the

tenants that are publicly available or that are required to be delivered to us pursuant to the applicable lease, (iii) monitoring news

reports regarding our tenants and their respective businesses, and (iv) monitoring the timeliness of lease payments.

Notes receivable

We carry notes receivable at amortized cost, adjusted for an estimated provision for expected credit losses. Interest income on

notes receivable is recognized using the effective interest rate method and is classified within other income in our consolidated

statements of operations. Direct costs incurred in originating notes, along with any premium or discount, are deferred and amortized as

an adjustment to interest income over the note’s term using the effective interest rate method. Notes receivable are classified within

other assets in our consolidated balance sheets. Refer to Note 8 – “Other assets” to our unaudited consolidated financial statements for

additional details.

Provision for expected credit losses

We are required to estimate and recognize lifetime expected losses, rather than incurred losses, for most financial assets

measured at amortized cost and certain other instruments, including trade, notes, and other receivables (excluding receivables arising

from operating leases), loans, held-to-maturity debt securities, net investments in leases arising from sales-type and direct financing

leases, and off-balance-sheet credit exposures (e.g., loan commitments). The recognition of such expected losses, even if the expected

risk of credit loss is remote, typically results in earlier recognition of credit losses. At each reporting date, we reassess our provision for

expected credit losses, and, if necessary, we recognize an adjustment for our current estimate of expected credit losses. Refer to

Note 5 – “Leases” and Note 8 – “Other assets” to our unaudited consolidated financial statements for additional details.

An assessment of the collectibility of operating lease payments and the recognition of an adjustment to lease income based on

this assessment is governed by the lease accounting standard discussed in “Lease accounting” earlier in Note 2 — “Summary of

significant accounting policies” to our unaudited consolidated financial statements.

Income taxes

We are organized and operate as a REIT pursuant to the Internal Revenue Code (the “Code”). Under the Code, a REIT that

distributes at least 90% of its REIT taxable income to its stockholders annually (excluding net capital gains) and meets certain other

conditions is not subject to federal income tax on its distributed taxable income, but could be subject to certain federal, foreign, state,

and local taxes. We distribute 100% of our taxable income annually; therefore, a provision for federal income taxes is not required. In

addition to our REIT returns, we file federal, foreign, state, and local tax returns for our subsidiaries. We file with jurisdictions located in

the U.S., Canada, and other international locations. Our tax returns are subject to routine examination in various jurisdictions for the

2020 through 2025 calendar years.

Employee and non-employee share-based payments

We have implemented an entity-wide accounting policy to account for forfeitures related to unmet service conditions of share-

based awards granted to employees and non-employees when they occur. Under this policy, when forfeitures occur, any previously

recognized expense related to those forfeited awards is reversed in the period of forfeiture.

Our employee and non-employee share-based awards are measured at fair value on the grant date and recognized over the

recipient’s required service period. For share-based awards with performance conditions, we continue to assess the probability of

achieving the performance conditions and recognize expense only when it becomes probable that the performance targets will be met.

Conversely, for share-based awards with market conditions, expense is recognized regardless of whether the market condition is met.

Dividends paid on share-based awards with nonforfeitable dividends are initially classified in retained earnings and reclassified

to compensation cost only if the underlying awards are forfeited. Conversely, for share-based awards with forfeitable dividends,

declared dividends are initially classified in retained earnings and in dividends payable within our consolidated balance sheets. If the

underlying awards are forfeited, the corresponding accrued dividend is reversed in the period of forfeiture. Upon vesting of the

underlying share-based awards with forfeitable dividends, the accumulated dividend payment is made and the dividend payable liability

is settled.

19

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

Forward equity sales agreements

From time to time, we enter into forward equity sales agreements and account for them in accordance with the accounting

guidance governing financial instruments and derivatives. Under the accounting guidance, our forward equity sales agreements are not

deemed to be liabilities as they do not embody obligations to repurchase our shares, nor do they embody obligations to issue a variable

number of shares for which the monetary value is predominantly fixed, varied with something other than the fair value of our shares, or

varied inversely in relation to our shares. We also evaluate whether the agreements meet the derivatives and hedging guidance scope

exception to be accounted for as equity instruments. Our forward equity sales agreements are classified as equity contracts based on

the following assessment: (i) none of the agreements’ exercise contingencies are based on observable markets or indices besides

those related to the market for our own stock price and operations; and (ii) none of the settlement provisions preclude the agreements

from being indexed to our own stock.

Hedge accounting

From time to time, we utilize derivative instruments to manage our exposure to certain risks. We are exposed to foreign

currency exchange rate risk related to our net investment in Canada. To mitigate the impact of fluctuations in the USD-CAD exchange

rate associated with our net investment in Canada, we use cross-currency swap agreements designated and qualifying as net

investment hedges under applicable derivatives and hedging standards.

We designate the USD-CAD cross-currency swap agreements as net investment hedges using the spot method to assess

hedge effectiveness. The spot component represents changes in fair value attributable to movements in the USD-CAD spot exchange

rate, which reflects the market exchange rate between the two currencies as of each reporting date. Changes in the fair value of the

designated spot component are recorded in other comprehensive income (loss) as part of the foreign currency translation adjustment,

to the extent the relationship is highly effective, until the net investment is sold or substantially liquidated. The related amounts due from

or due to counterparties are included in other assets or in accounts payable, accrued expenses, and other liabilities, respectively, within

our consolidated balance sheet.

We elected to account for the forward points (the portion of the derivative’s fair value attributable to the difference between the

forward exchange rate and spot exchange rate) as an excluded component in accordance with applicable derivatives and hedging

accounting standards. The excluded component is recognized over the life of the cross-currency swap agreements using a systematic

and rational basis (as interest settlements occur) and is classified within other income in our consolidated statement of operations.

Issuer and guarantor subsidiaries of guaranteed securities

Generally, a parent entity of an issuer that holds guaranteed securities must provide separate subsidiary issuer or guarantor

financial statements, unless it qualifies for disclosure exceptions. A parent entity may be eligible for disclosure exceptions if it meets the

following criteria:

(i)The subsidiary issuer or guarantor is a consolidated subsidiary of the parent company, and

(ii)The subsidiary issues a registered security that is:

•issued jointly and severally with the parent company, or

•fully and unconditionally guaranteed by the parent company.

A parent entity that meets the above criteria may instead present summarized financial information (“alternative disclosures”)

either within the consolidated financial statements or in “Item 2. Management’s discussion and analysis of financial condition and results

of operations” (“Item 2”). We evaluated the criteria and determined that we are eligible for the disclosure exceptions, which allow us to

provide alternative disclosures; as such, we present alternative disclosures in Item 2.

Loan fees

Fees incurred in obtaining long-term financing are capitalized and classified with the corresponding debt instrument appearing

on our consolidated balance sheets. Loan fees related to our unsecured senior line of credit are capitalized and classified within other

assets. Capitalized amounts are amortized over the term of the related loan, and the amortization is classified in interest expense in our

consolidated statements of operations.

Distributions from equity method investments

We use the “nature of the distribution” approach to determine the classification within our consolidated statements of cash

flows of cash distributions received from equity method investments, including our unconsolidated real estate joint ventures and equity

method non-real estate investments. Under this approach, distributions are classified based on the nature of the underlying activity that

generated the cash distributions. If we lack the information necessary to apply this approach in the future, we will be required to apply

the “cumulative earnings” approach as an accounting change on a retrospective basis. Under the cumulative earnings approach,

distributions up to the amount of cumulative equity in earnings recognized are classified as cash inflows from operating activities, and

those in excess of that amount are classified as cash inflows from investing activities.

20

2.SUMMARY OF SIGNIFICANT ACCOUNTING POLICIES (continued)

Restricted cash

We present cash and cash equivalents separately from restricted cash within our consolidated balance sheets. However, we

include restricted cash with cash and cash equivalents when reconciling the beginning-of-period and end-of-period total amounts shown

in the consolidated statements of cash flows. We provide a reconciliation between the consolidated balance sheets and the

consolidated statements of cash flows, which is required when the balance includes greater than one line item for cash, cash

equivalents, and restricted cash. We also provide a disclosure of the nature of the restrictions related to material restricted cash

balances.

Recent accounting pronouncements

On November 4, 2024, the FASB issued ASU 2024-03, Disaggregation of Income Statement Expenses, which will require

entities to provide enhanced disclosures related to certain expense categories included in line items on the statement of operations.

The ASU aims to increase transparency and provide investors with additional detailed information about the nature of expenses

reported on the face of the income statement. The new standard does not change the requirements for the presentation of expenses on

the face of the statement of operations.

Under this ASU, entities are required to disaggregate, in a tabular format, expense line items presented on the face of the

statement of operations — excluding earnings or losses from equity method investments — if they include any of the following expense

categories: purchases of inventory, employee compensation, depreciation, intangible asset amortization, and depreciation or depletion.

For any remaining items within each relevant expense line item, entities must provide a qualitative description of the nature of those

expenses. The new ASU is effective for annual reporting periods beginning after December 15, 2026 and interim reporting periods

beginning after December 15, 2027. Early adoption is permitted. We expect to adopt this ASU on January 1, 2027. Although the

adoption is not expected to have an impact on our financial statements, it is expected to result in incremental disclosures within the

notes to our consolidated financial statements.

21

3.INVESTMENTS IN REAL ESTATE

Our consolidated investments in real estate consisted of the following as of March 31, 2026 and December 31, 2025 (in

thousands):

March 31, 2026 December 31, 2025
Rental properties:
Land (related to rental properties) $3,639,219 $3,204,479
Buildings and building improvements 19,607,116 19,738,825
Other improvements 4,465,630 4,371,720
Rental properties 27,711,965 27,315,024
Current and future development and redevelopment projects 6,857,569 6,788,464
Gross investments in real estate 34,569,534 34,103,488
Less: accumulated depreciation (6,232,579) (5,970,171)
Investments in real estate assets held for sale, less accumulated depreciation(1) 493,161 556,679
Investments in real estate $28,830,116 $28,689,996

(1)Refer to “Assets held for sale” below.

Assets held for sale

As of March 31, 2026, we had 19 operating properties aggregating 1.4 million RSF and land parcels aggregating 1.6 million SF

that were classified as held for sale.

The disposal of properties classified as held for sale does not represent a strategic shift that has, or will have, a major effect on

our operations or financial results, as the dispositions relate to individual assets across multiple markets and do not represent the exit

from any significant market. Accordingly, these assets do not meet the criteria for classification as a discontinued operation. We cease

depreciation of our properties upon their classification as held for sale.

The following table presents the components of net assets related to real estate investments that met the criteria for

classification as held for sale as of March 31, 2026 and December 31, 2025 (in thousands):

March 31, 2026 December 31, 2025
Investments in real estate, less accumulated depreciation $493,161 $556,679
Other assets 41,409 37,859
Total assets 534,570 594,538
Total liabilities (11,841) (12,235)
Total accumulated other comprehensive loss (181) (566)
Net assets classified as held for sale $522,548 $581,737

For additional information, refer to “Real estate sales” in Note 2 – “Summary of significant accounting policies” to our unaudited

consolidated financial statements.

Sales of real estate assets and impairment of real estate

Impairment of real estate

During the three months ended March 31, 2026, we recognized impairment charges aggregating $5.5 million, classified within

impairment of real estate in our consolidated statement of operations. The impairment primarily reflects an incremental charge

recognized in connection with the amendment of the sales agreement during the three months ended March 31, 2026 related to our

Canada portfolio, which is classified as held for sale as of March 31, 2026.

22

3.INVESTMENTS IN REAL ESTATE (continued)

Other

ARE‑East River Science Park, LLC (“ARE”), a subsidiary of Alexandria Real Estate Equities, Inc., holds an option granted in

2006 to incorporate a land parcel adjacent to and north of the Alexandria Center® for Life Science – New York City campus (the “Option

Parcel”) into the existing ground lease, which would allow for the future development of an additional life science building within the

campus. ARE’s investment in pre‑construction costs related to the Option Parcel aggregated $180.6 million as of March 31, 2026.

On August 6, 2024, ARE filed a lawsuit in the U.S. District Court for the Southern District of New York against New York City

Health + Hospitals Corporation (“H+H”) and the New York City Economic Development Corporation (“EDC”) relating to disputes under

the ground lease and option arrangements governing the Option Parcel. ARE filed an amended complaint on January 24, 2025,

asserting claims for fraudulent inducement, breach of contract, breach of the implied covenant of good faith and fair dealing, and

declaratory relief concerning the continued validity of the option.

On March 27, 2026, the court granted defendants’ partial motion to dismiss the fraud in the inducement and implied covenant

of good faith and fair dealing claims. ARE intends to appeal this order at the appropriate time and to vigorously pursue its claims. The

court did not dismiss ARE’s claim for declaratory relief, which remains pending.

On April 10, 2026, H+H and EDC answered the amended complaint and asserted counterclaims seeking, among other things,

declaratory relief relating to the alleged expiration of the option and entitlement to a $5.0 million security deposit, and damages of at

least $3.8 million. As a result of the foregoing matters, the timing of any development of the Option Parcel is currently indeterminate.

Excluding the potential impact of the counterclaims filed by H+H and EDC, this matter exposes us to potential losses ranging from zero

to the full amount of the investment in the project aggregating $180.6 million as of March 31, 2026, depending on the resolution of the

remaining declaratory relief proceedings, the outcome of any appeal, and/or the ability to develop the project. We performed a

probability-weighted recoverability analysis based on initial estimates of various possible outcomes and determined no impairment was

present as of March 31, 2026.

Separately from, and not part of, the pending litigation related to the Option Parcel, on April 21, 2026, EDC, on behalf of H+H,

the landlord, and itself, lease administrator, delivered a notice alleging that ARE is in default of certain information delivery obligations

under the ground lease relating to the existing operating towers at the Alexandria Center® for Life Science – New York City campus,

including sublease, tax, and financial information. The notice asserts that the failure to cure the alleged defaults within the applicable

30-day cure period would result in daily charges of $1,000 increasing thereafter to up to $2,000. ARE disputes the allegations set forth

in the notice of default, and intends to vigorously defend against them.

23

4.CONSOLIDATED AND UNCONSOLIDATED REAL ESTATE JOINT VENTURES

From time to time, we enter into joint venture agreements through which we own a partial interest in real estate entities that

own, develop, and operate real estate properties. As of March 31, 2026, our real estate joint ventures held the following properties:

Property(1) Market Submarket Our Ownership<br><br>Interest
Consolidated real estate joint ventures:
50 and 60 Binney Street Greater Boston Cambridge/Inner Suburbs 34.0%
75/125 Binney Street Greater Boston Cambridge/Inner Suburbs 40.0%
100 and 225 Binney Street and 300 Third Street Greater Boston Cambridge/Inner Suburbs 30.0%
15 Necco Street Greater Boston Seaport Innovation District 56.7%
Alexandria Center® for Science and Technology –<br><br>Mission Bay(2) San Francisco Bay Area Mission Bay 25.0%
211 and 213 East Grand Avenue San Francisco Bay Area South San Francisco 30.0%
500 Forbes Boulevard San Francisco Bay Area South San Francisco 10.0%
Alexandria Center® for Life Science – Millbrae San Francisco Bay Area South San Francisco 48.6%
3215 Merryfield Row San Diego Torrey Pines 30.0%
Campus Point by Alexandria(3) San Diego University Town Center 57.2% (4)
5200 Illumina Way San Diego University Town Center 51.0%
9625 Towne Centre Drive San Diego University Town Center 30.0%
SD Tech by Alexandria(5) San Diego Sorrento Mesa 50.0%
Summers Ridge Science Park(6) San Diego Sorrento Mesa 30.0%
1201 and 1208 Eastlake Avenue East Seattle Lake Union 30.0%
400 Dexter Avenue North Seattle Lake Union 30.0%
800 Mercer Street Seattle Lake Union 60.0%
Unconsolidated real estate joint ventures:
1655 and 1725 Third Street San Francisco Bay Area Mission Bay 10.0%
101 West Dickman Street Maryland Beltsville 58.4% (7)

(1)Refer to the table on the next page that shows the categorization of our real estate joint ventures under the consolidation framework.

(2)Includes 1450, 1500, and 1700 Owens Street, and 455 Mission Bay Boulevard South.

(3)Includes 10200, 10290, and 10300 Campus Point Drive and 4135, 4155, 4165, 4224, and 4242 Campus Point Court.

(4)The noncontrolling interest share of our joint venture partner is anticipated to decrease to 25%, as we expect to fund the majority of future construction costs at the

campus until our ownership interest increases to 75%, after which future capital would be contributed pro rata with our partner.

(5)Includes 9605, 9645, 9675, 9725, 9735, 9805, 9808, 9855, and 9868 Scranton Road and 10055, 10065, and 10075 Barnes Canyon Road.

(6)Includes 9965, 9975, 9985, and 9995 Summers Ridge Road.

(7)Represents a joint venture with a local real estate operator in which our joint venture partner manages the day-to-day activities that significantly affect the economic

performance of the joint venture.

Our consolidation policy is described under “Consolidation” in Note 2 – “Summary of significant accounting policies” to our

unaudited consolidated financial statements. Consolidation accounting is highly technical, but its framework is primarily based on the

controlling financial interests and benefits of the joint ventures. We generally consolidate a joint venture that is a legal entity that we

control (i.e., we have the power to direct the activities of the joint venture that most significantly affect its economic performance)

through contractual rights, regardless of our ownership interest, and where we determine that we have benefits through the allocation of

earnings or losses and fees paid to us that could be significant to the joint venture (the “VIE model”).

We also generally consolidate joint ventures when we have a controlling financial interest through voting rights and where our

voting interest is greater than 50% (the “voting model”). Voting interest differs from ownership interest for some joint ventures. We

account for joint ventures that do not meet the consolidation criteria under the equity method of accounting by recognizing our share of

income and losses.

24

4.CONSOLIDATED AND UNCONSOLIDATED REAL ESTATE JOINT VENTURES (continued)

The table below shows the categorization of our real estate joint ventures under the consolidation framework:

Property Consolidation<br><br>Model Voting Interest Consolidation Analysis Conclusion
50 and 60 Binney Street VIE model Not applicable<br><br>under VIE<br><br>model Consolidated
75/125 Binney Street We have:
100 and 225 Binney Street and 300<br><br>Third Street
15 Necco Street (i) The power to direct the<br><br>activities of the joint venture<br><br>that most significantly affect its<br><br>economic performance; and
Alexandria Center® for Science and<br><br>Technology – Mission Bay
211 and 213 East Grand Avenue
500 Forbes Boulevard
Alexandria Center® for Life Science –<br><br>Millbrae (ii) Benefits that can be significant<br><br>to the joint venture.
3215 Merryfield Row
Campus Point by Alexandria
5200 Illumina Way Therefore, we are the primary<br><br>beneficiary of each VIE
9625 Towne Centre Drive
SD Tech by Alexandria
Summers Ridge Science Park
1201 and 1208 Eastlake Avenue East
400 Dexter Avenue North
800 Mercer Street
101 West Dickman Street We do not control the joint venture<br><br>and are therefore not the primary<br><br>beneficiary. Equity method<br><br>of accounting
1655 and 1725 Third Street Voting model Does not<br><br>exceed 50% Our voting interest is 50% or less.

Consolidated real estate joint ventures

99 Coolidge Avenue

In July 2025, we amended the agreement for our consolidated real estate joint venture at 99 Coolidge Avenue in our

Cambridge/Inner Suburbs submarket. Pursuant to the amendment, the carrying amount of our partner’s noncontrolling interest was

adjusted from $42.0 million to $48.7 million, and converted into a redeemable noncontrolling interest that accrued a fixed 4.05% annual

preferred return (“distributions”). In January 2026, the partner exercised its option to require us to purchase its preferred interest, and

the redemption was completed in January 2026 for $49.7 million, inclusive of unpaid distributions.

Consolidated VIEs’ balance sheet information

We, together with joint venture partners, hold interests in real estate joint ventures that we consolidate in our financial

statements. These existing joint ventures provide significant equity capital to fund a portion of our future construction spending, and our

joint venture partners may also contribute equity into these entities for financing-related activities.

25

4.CONSOLIDATED AND UNCONSOLIDATED REAL ESTATE JOINT VENTURES (continued)

The table below aggregates the balance sheet information of our consolidated VIEs (in thousands):

March 31, 2026 December 31, 2025
Investments in real estate $5,797,860 $6,129,668
Cash and cash equivalents 185,263 258,755
Other assets 724,923 712,154
Total assets $6,708,046 $7,100,577
Secured note payable $— $—
Other liabilities 287,285 324,513
Total liabilities 287,285 324,513
Redeemable noncontrolling interests 49,554
Alexandria Real Estate Equities, Inc.’s share of equity 2,800,347 3,098,932
Noncontrolling interests’ share of equity 3,620,414 3,627,578
Total liabilities and equity $6,708,046 $7,100,577

In determining whether to aggregate the balance sheet information of consolidated VIEs, we considered the similarity of each

VIE, including the primary purpose of these entities to own, manage, operate, and lease real estate properties owned by the VIEs, and

the similar nature of our involvement in each VIE as a managing member. Due to the similarity of the characteristics, we present the

balance sheet information of these entities on an aggregated basis. None of our consolidated VIEs’ assets have restrictions that limit

their use to settle specific obligations of the VIE. There are no creditors or other partners of our consolidated VIEs that have recourse to

our general credit, and our maximum exposure to our consolidated VIEs is limited to our variable interests in each VIE.

Noncontrolling interests in consolidated real estate joint ventures

Noncontrolling interests represent the third-party interests in consolidated real estate joint ventures in which we have a

controlling interest. Noncontrolling interests are adjusted for additional contributions and distributions, the proportionate share of the net

earnings or losses, and other comprehensive income or loss. Distributions, profits, and losses related to these entities are allocated in

accordance with the respective operating agreements. During the three months ended March 31, 2026 and 2025, we distributed

$60.1 million and $66.0 million, respectively, to our consolidated real estate joint venture partners.

Unconsolidated real estate joint ventures

Our investments in unconsolidated real estate joint ventures, accounted for under the equity method and classified in

investments in unconsolidated real estate joint ventures in our consolidated balance sheets, consisted of the following as of March 31,

2026 and December 31, 2025 (in thousands):

Property March 31, 2026 December 31, 2025
1655 and 1725 Third Street $19,333 $19,484
101 West Dickman Street 9,666 9,669
Other 1,521 1,524
$30,520 $30,677

Our maximum exposure to our 1655 and 1725 Third Street unconsolidated VIE is limited to our investment in this VIE. With

respect to our unconsolidated real estate joint venture at 101 West Dickman Street, we guarantee up to $6.7 million of the outstanding

balance related to the VIE’s secured construction loan.

Below are key terms of unconsolidated real estate joint ventures’ secured loans as of March 31, 2026 (dollars in thousands):

Interest<br><br>Rate(1) At 100%
Unconsolidated Joint Venture Maturity Date Stated Rate Aggregate<br><br>Commitment Debt Balance(2)
101 West Dickman Street 10/29/26 SOFR+1.95% (3) 5.68% $26,750 19,048
1655 and 1725 Third Street 2/10/35 6.37% 6.44% 500,000 496,967
$526,750 516,015

All values are in US Dollars.

(1)Includes interest expense and amortization of loan fees.

(2)Represents outstanding principal, net of unamortized deferred financing costs, as of March 31, 2026.

(3)This loan is subject to a fixed SOFR floor of 0.75%.

26

5.LEASES

Refer to “Lease accounting” in Note 2 – “Summary of significant accounting policies” to our unaudited consolidated financial

statements for information about lease accounting standards that set principles for the recognition, measurement, presentation, and

disclosure of leases for both parties to a lease agreement (i.e., lessees and lessors).

Leases in which we are the lessor

As of March 31, 2026, we had 339 properties aggregating 35.8 million operating RSF in key cluster locations, including

Greater Boston, the San Francisco Bay Area, San Diego, Seattle, Maryland, Research Triangle, and New York City. We primarily focus

on developing Class A/A+ properties in AAA life science innovation clusters that offer the scale and strategic design integral to our

Megacampus strategy. Strategically located near top academic and medical research institutions, our Megacampus ecosystems feature

curated amenities and services and convenient access to transit, creating environments that help our tenants attract and retain top

talent.

As of March 31, 2026, all leases in which we are the lessor were classified as operating leases, with the exception of one

direct financing and one sales-type lease. Our leases are described below.

Operating leases

As of March 31, 2026, our 339 properties were subject to operating lease agreements. Five of these properties are subject to

operating lease agreements that each contain a purchase option as described below:

(i)Two of these properties, representing two land parcels in the San Francisco Bay Area market, are subject to lease

agreements that each contain an option for the lessee to purchase the underlying asset from us at fair market value during

each of the 30-day periods commencing on the dates that are 15 years, 30 years, and 74.5 years after the rent

commencement date of October 1, 2017. The remaining lease term related to each of the two land parcels is 66.7 years.

(ii)Two operating properties in the Seattle market, held by a consolidated real estate joint venture, are subject to purchase

options held by our partner in this joint venture, which is also a tenant at these properties. One purchase option allows our

partner to purchase our 30% interest in one property for $40.0 million in 2031. Contingent upon the exercise of this option,

the second purchase option allows our partner to purchase our 30% interest in one property for $69.1 million in 2034. Our

partner’s remaining lease terms for these operating leases are 6.9 years and 18.5 years, respectively.

(iii)One property subject to an operating lease agreement contains a purchase option exercisable at fair market value in

March 2034.

Certain operating leases contain options for the tenant to extend their lease at prevailing market rates at the time of expiration.

In addition, certain operating leases contain an early termination option that requires advance notification and payment of an early

termination fee by the tenant.

At the commencement of each lease, we establish the lease term comprising the noncancelable period for each lease together

with periods covered by options to extend or terminate the lease that we determine the lessee is reasonably certain to exercise. Our

assessment of whether a lessee is reasonably certain to exercise or not exercise an option considers all economic factors relevant to

the assessment, including property-based, market-based, and tenant-based factors. We do not reassess the lease term or a lessee

option to purchase the underlying asset unless there is a lease modification that is not accounted for as a separate contract.

Future lease payments to be received under the terms of our operating lease agreements, excluding expense

reimbursements, in effect as of March 31, 2026 are outlined in the table below (in thousands):

Year Amount
2026 $1,220,483
2027 1,494,928
2028 1,359,084
2029 1,261,862
2030 1,193,688
Thereafter 7,152,763
Total $13,682,808

Refer to Note 3 – “Investments in real estate” to our unaudited consolidated financial statements for additional information

about our owned real estate assets, which are the underlying assets under our operating leases.

27

5.LEASES (continued)

Direct financing and sales-type leases

As of March 31, 2026, we have one direct financing lease agreement, with a net investment balance of $42.8 million, for a

parking structure with a remaining lease term of 66.7 years. The lessee has an option to purchase the underlying asset at fair market

value during each of the 30-day periods commencing on the dates that are 15 years, 30 years, and 74.5 years after the rent

commencement date of October 1, 2017.

As of March 31, 2026, we also have one sales-type lease for a property in the Seattle market. As of March 31, 2026, the net

investment in this lease is $16.6 million. At the end of the lease term in 2026, the property under this lease will transfer to the tenant for

a sales price of approximately $18.0 million.

As of March 31, 2026, our estimated provision for expected credit loss related to our direct financing lease and sales-type

lease aggregated $1.8 million, which was predominantly related to our direct financing lease. We estimate the provision for expected

credit loss related to our direct financing lease using a probability of default methodology, which incorporates the borrower’s investment-

grade credit rating from S&P Global Ratings, to evaluate the probability of default. Additionally, we incorporate the projected value of the

real estate securing the investments to estimate potential recoveries in the event of default, among other inputs. The estimate of the

expected credit loss related to our sales-type lease was determined using historical industry losses and transaction-specific information,

including the estimated fair value of the underlying real estate asset securing this transaction, the short-term nature of this lease, and

other available information. For further details, refer to “Provision for expected credit losses” in Note 2 – “Summary of significant

accounting policies” to our unaudited consolidated financial statements.

The components of our aggregate net investment in our direct financing and sales-type leases as of March 31, 2026 and

December 31, 2025 are summarized in the table below (in thousands):

March 31, 2026 December 31, 2025
Gross investment in direct financing and sales-type leases $265,618 $265,839
Less: unearned income on direct financing lease (204,355) (205,037)
Less: provision for expected credit losses (1,817) (1,817)
Net investment in leases $59,446 $58,985

Future lease payments to be received under the terms of our direct financing and sales-type leases as of March 31, 2026 are

outlined in the table below (in thousands):

Year Total
2026 $18,225
2027 2,097
2028 2,160
2029 2,224
2030 2,291
Thereafter 238,621
Total $265,618

Income from rentals

Our income from rentals includes revenue related to agreements for the rental of our real estate, which primarily includes

revenues subject to the lease accounting standard and the revenue recognition accounting standard as shown below (in thousands):

Three Months Ended March 31,
2026 2025
Income from rentals:
Revenues subject to the lease accounting standard:
Operating leases $640,659 $731,421
Direct financing and sales-type leases 964 810
Revenues subject to the lease accounting standard 641,623 732,231
Revenues subject to the revenue recognition accounting standard 11,390 10,944
Income from rentals $653,013 $743,175

28

5.LEASES (continued)

Revenues subject to the revenue recognition accounting standard and classified in income from rentals consist primarily of

short-term parking revenues that are not considered lease revenues under the lease accounting standard. Refer to “Revenues” and

“Recognition of revenue arising from contracts with customers” in Note 2 – “Summary of significant accounting policies” to our

unaudited consolidated financial statements for additional information.

Residual value risk management strategy

Our leases do not have guarantees of residual value on the underlying assets. We manage risk associated with the residual

value of our leased assets by (i) evaluating each potential acquisition of real estate to determine whether it meets our business

objective to invest primarily in high-demand markets, (ii) directly managing our leased properties, conducting frequent property

inspections, proactively addressing potential maintenance issues, and/or timely resolving any occurring issues, and (iii) carefully

selecting our tenants and monitoring their credit quality throughout their respective lease terms.

Leases in which we are the lessee

Operating lease agreements

We have operating lease agreements in which we are the lessee consisting of ground and office leases. Certain of these

leases have options to extend or terminate the contract terms upon meeting certain criteria. There are no notable restrictions or

covenants imposed by the leases, nor guarantees of residual value.

We recognize a right-of-use asset, which is classified within other assets in our consolidated balance sheets, and a related

liability, which is classified within accounts payable, accrued expenses, and other liabilities in our consolidated balance sheets, to

account for our future obligations under ground and office lease arrangements in which we are the lessee. Refer to “Lessee accounting”

in Note 2 – “Summary of significant accounting policies” to our unaudited consolidated financial statements.

As of March 31, 2026, the present value of the remaining contractual payments aggregating $767.2 million under our operating

lease agreements, including our extension options that we are reasonably certain to exercise, was $358.6 million. Our corresponding

operating lease right-of-use assets, adjusted for initial direct leasing costs and other consideration exchanged with the landlord prior to

the commencement of the lease, aggregated $693.8 million. As of March 31, 2026, the weighted-average remaining lease term of

operating leases in which we are the lessee was approximately 60 years, including extension options that we are reasonably certain to

exercise, and the weighted-average discount rate was 4.7%. The weighted-average discount rate is based on the incremental

borrowing rate estimated for each lease, which is the interest rate that we estimate we would have to pay to borrow on a collateralized

basis over a similar term for an amount equal to the lease payments.

Ground lease obligations as of March 31, 2026 included leases for 31 of our properties, which accounted for approximately 9%

of our total number of properties. Excluding one ground lease that expires in 2036 related to one operating property with a net book

value of $5.0 million as of March 31, 2026, our ground lease obligations have remaining lease terms ranging from approximately 28 to

97 years, including extension options that we are reasonably certain to exercise.

29

5.LEASES (continued)

The reconciliation of future lease payments under noncancelable operating leases in which we are the lessee to the operating

lease liability reflected in our unaudited consolidated balance sheet as of March 31, 2026 is in the table below (in thousands):

Year Total
2026 $17,206
2027 21,650
2028 21,318
2029 20,825
2030 20,743
Thereafter 665,496
Total future payments under our operating leases in which we are the lessee 767,238
Effect of discounting (408,628)
Operating lease liability $358,610

Lessee operating costs

Operating lease costs relate to our ground and office leases in which we are the lessee. Ground leases generally require fixed

annual rent payments and may also include escalation clauses and renewal options. For the three months ended March 31, 2026 and

2025, amounts paid and classified as operating activities in our unaudited consolidated statements of cash flows for leases in which we

are the lessee aggregated $6.2 million and $144.6 million, respectively. The decrease is primarily due to the ground lease prepayment

of $135.0 million made in January 2025 for a 24-year lease term extension to our existing ground lease agreement at the Alexandria

Technology Square® Megacampus in our Cambridge submarket.

Our operating lease obligations related to our office leases have remaining terms of up to 10 years, exclusive of extension

options. For the three months ended March 31, 2026 and 2025, our costs for operating leases in which we are the lessee were as

follows (in thousands):

Three Months Ended March 31,
2026 2025
Gross operating lease costs $8,417 $12,359
Capitalized lease costs (759) (693)
Expenses for operating leases in which we are the lessee $7,658 $11,666

6. CASH, CASH EQUIVALENTS, AND RESTRICTED CASH

Cash, cash equivalents, and restricted cash consisted of the following as of March 31, 2026 and December 31, 2025 (in

thousands):

March 31, 2026 December 31, 2025
Cash and cash equivalents $418,720 $549,062
Restricted cash:
Development escrows 2,129 2,142
Security deposits 1,804 1,729
Other 732 822
4,665 4,693
Total $423,385 $553,755

30

7.INVESTMENTS

We hold investments in publicly traded companies and privately held entities primarily involved in the life science industry. As a

REIT, we generally limit our ownership of each individual entity’s voting stock to less than 10%. We evaluate each investment to

determine whether we have the ability to exercise significant influence, but not control, over an investee. We evaluate investments in

which our ownership is equal to or greater than 20%, but less than or equal to 50%, of an investee’s voting stock with a presumption

that we have this ability. For our investments in limited partnerships that maintain specific ownership accounts, we presume that such

ability exists when our ownership interest exceeds 3% to 5%. In addition to our ownership interest, we consider whether we have a

board seat or whether we participate in the investee’s policy-making process, among other criteria, to determine if we have the ability to

exert significant influence, but not control, over an investee. If we determine that we have such ability, we account for the investment

under the equity method, as described below.

From time to time, we may hold equity investments in publicly traded companies that are subject to temporary contractual sale

restrictions. We do not recognize a discount related to such contractual sale restrictions.

Investments accounted for under the equity method

Under the equity method of accounting, we initially recognize our investment at cost and subsequently adjust the carrying

amount of the investment for our share of earnings or losses reported by the investee, distributions received, and other-than-temporary

impairments.

As of March 31, 2026, we had nine investments in limited partnerships maintaining specific ownership accounts for each

investor, which were accounted for under the equity method. These investments aggregated $367.9 million. Our ownership interest in

each of these nine investments was greater than 5%.

Investments that do not qualify for the equity method of accounting

For investees over which we determine that we do not have the ability to exercise significant influence or control, we account

for each investment depending on whether it is an investment in a (i) publicly traded company, (ii) privately held entity that reports NAV

per share, or (iii) privately held entity that does not report NAV per share, as described below.

Investments in publicly traded companies

Our investments in publicly traded companies are classified as investments with readily determinable fair values and are

presented at fair value in our consolidated balance sheets, with changes in fair value classified in investment income (loss) in our

consolidated statements of operations. The fair values for our investments in publicly traded companies are determined based on sales

prices or quotes available on securities exchanges.

Investments in privately held companies

Our investments in privately held entities without readily determinable fair values consist of (i) investments in privately held

entities that report NAV per share and (ii) investments in privately held entities that do not report NAV per share. These investments are

accounted for as follows:

Investments in privately held entities that report NAV per share

Investments in privately held entities that report NAV per share, such as our privately held investments in limited partnerships,

are presented at fair value using NAV as a practical expedient, with changes in fair value classified in investment income (loss) in our

consolidated statements of operations. We use NAV per share reported by limited partnerships generally without adjustment, unless we

are aware of information indicating that the NAV reported by a limited partnership does not accurately reflect the fair value of the

investment at our reporting date.

Investments in privately held entities that do not report NAV per share

Investments in privately held entities that do not report NAV per share are accounted for using a measurement alternative

under which these investments are measured at cost, adjusted for observable price changes and impairments, with changes classified

in investment income (loss) in our consolidated statements of operations.

An observable price arises from an orderly transaction for an identical or similar investment of the same issuer, which is

observed by an investor without expending undue cost and effort. Observable price changes result from, among other things, equity

transactions of the same issuer executed during the reporting period, including subsequent equity offerings or other reported equity

transactions related to the same issuer. To determine whether these transactions are indicative of an observable price change, we

evaluate, among other factors, whether these transactions have similar rights and obligations, including voting rights, distribution

preferences, and conversion rights to the investments we hold.

31

7.INVESTMENTS (continued)

Impairment evaluation of equity method investments and investments in privately held entities that do not report NAV per share

We monitor equity method investments and investments in privately held entities that do not report NAV per share for new

developments, including operating results, prospects and results of clinical trials, new product initiatives, new collaborative agreements,

capital-raising events, and merger and acquisition activities. These investments are evaluated on the basis of a qualitative assessment

for indicators of impairment by monitoring the presence of the following triggering events or impairment indicators:

(i)a significant deterioration in the earnings performance, credit rating, asset quality, or business prospects of the investee;

(ii)a significant adverse change in the regulatory, economic, or technological environment of the investee;

(iii)a significant adverse change in the general market condition, including the research and development of technology and

products that the investee is bringing or attempting to bring to the market;

(iv)significant concerns about the investee’s ability to continue as a going concern; and/or

(v)a decision by investors to cease providing support or reduce their financial commitment to the investee.

If such indicators are present, we are required to estimate the investment’s fair value and immediately recognize an

impairment charge in an amount equal to the investment’s carrying value in excess of its estimated fair value.

Investment income/loss recognition and classification

We recognize both realized and unrealized gains and losses in our consolidated statements of operations, classified in

investment income (loss) in our consolidated statements of operations. Unrealized gains and losses represent:

(i)changes in fair value for investments in publicly traded companies;

(ii)changes in NAV for investments in privately held entities that report NAV per share;

(iii)observable price changes for investments in privately held entities that do not report NAV per share; and

(iv)our share of unrealized gains or losses reported by our equity method investees.

Realized gains and losses on our investments represent the difference between proceeds received upon disposition of

investments and their historical or adjusted cost basis. For our equity method investments, realized gains and losses represent our

share of realized gains or losses reported by the investee. Impairments are realized losses, which result in an adjusted cost basis, and

represent charges to reduce the carrying values of investments in privately held entities that do not report NAV per share and equity

method investments, if impairments are deemed other than temporary, to their estimated fair value.

Funding commitments to investments in privately held entities that report NAV

We are committed to funding approximately $327.8 million for our investments in privately held entities that report NAV. Our

funding commitments expire at various dates over the next 12 years, with a weighted-average expiration of 8.0 years as of March 31,

  1. These investments are not redeemable by us, but we may receive distributions from these investments throughout their terms.

Our investments in privately held entities that report NAV generally have expected initial terms in excess of 10 years. The weighted-

average remaining term during which these investments are expected to be liquidated was 5.5 years as of March 31, 2026.

The following tables summarize our investments as of March 31, 2026 and December 31, 2025 (in thousands):

March 31, 2026
Cost Unrealized<br><br>Gains Unrealized<br><br>Losses Carrying<br><br>Amount
Publicly traded companies $83,916 $34,674 $(16,514) $102,076
Entities that report NAV 471,058 102,050 (38,132) 534,976
Entities that do not report NAV:
Entities with observable price changes 82,128 54,780 (10,991) 125,917
Entities without observable price changes 405,567 405,567
Investments accounted for under the equity method N/A N/A N/A 367,883
Total investments $1,042,669 $191,504 $(65,637) $1,536,419

32

7.INVESTMENTS (continued)

December 31, 2025
Cost Unrealized<br><br>Gains Unrealized<br><br>Losses Carrying<br><br>Amount
Publicly traded companies $54,752 $44,319 $(4,143) $94,928
Entities that report NAV 460,160 89,514 (37,298) 512,376
Entities that do not report NAV:
Entities with observable price changes 82,252 50,601 (9,615) 123,238
Entities without observable price changes 413,324 413,324
Investments accounted for under the equity method N/A N/A N/A 357,383
Total investments $1,010,488 $184,434 $(51,056) $1,501,249

Cumulative gains and losses (realized and unrealized) on investments in privately held entities that do not report NAV still held

as of March 31, 2026 aggregated to a loss of $125.7 million, which consisted of upward adjustments aggregating $54.8 million,

downward adjustments aggregating $11.0 million, and impairments aggregating $169.5 million.

Our investment income (loss) for the three months ended March 31, 2026 and 2025 consisted of the following (in thousands):

Three Months Ended March 31,
2026 2025
Realized gains $5,750 (1) $18,153
Unrealized losses (10,332) (68,145)
Investment loss $(4,582) (2) $(49,992)

(1)Consists of realized gains of $18.2 million, partially offset by impairment charges of $12.4 million during the three months ended March 31, 2026.

(2)Investment loss of $4.6 million for the three months ended March 31, 2026 also included $1.7 million of equity in losses of our equity method investments.

Additional details on our non-real estate investments still held as of the end of each period are presented below (in thousands):

Three Months Ended March 31,
2026 2025
Investments in privately held entities that do not report NAV still held as of the end of<br><br>each period:
Upward adjustments $6,044 $8,076
Downward adjustments and impairments (13,154) (17,052)
$(7,110) $(8,976)
Unrealized gains (losses) on non-real estate investments still held as of the end of<br><br>each period (excluding equity method investments) $3,877 $(37,398)

Refer to “Investments” in Note 2 – “Summary of significant accounting policies” to our unaudited consolidated financial

statements for additional information.

33

8. OTHER ASSETS

The following table summarizes the components of other assets as of March 31, 2026 and December 31, 2025 (in thousands):

March 31, 2026 December 31, 2025
Acquired in-place leases $189,108 $204,008
Deferred compensation plan 53,568 53,529
Deferred financing costs – unsecured senior line of credit 36,993 39,406
Deposits 28,472 28,618
Furniture, fixtures, equipment, and software 79,902 70,311
Net investment in leases 59,446 58,985
Notes receivable 260,167 258,033
Operating lease right-of-use assets 693,808 697,865
Other assets 87,894 87,036
Prepaid expenses 64,281 33,718
Property, plant, and equipment 129,504 130,263
Total $1,683,143 $1,661,772

Notes receivable

Our notes receivable as of March 31, 2026 and December 31, 2025 consisted of the following (dollars in thousands):

March 31, 2026
Weighted-Average
Notes Receivable Effective<br><br>Interest Rate Maturity<br><br>Date Balance December 31, 2025
Secured by real estate assets in San Diego 9.9% 1/10/29 $245,204 $240,476
Secured by real estate assets in Greater Boston 6.2% 12/16/29 15,495 18,089
Less: provision for expected credit losses (532) (532)
Notes receivable $260,167 $258,033

Our notes receivable represent held-to-maturity debt securities carried at amortized cost and are generally secured by real

estate. Under the current expected credit losses accounting standard, we are required to estimate and, if necessary, recognize a

provision for expected credit losses related to these notes. We do not have a history of losses on such securities; therefore, we utilize

available information on historical losses for the commercial real estate industry. We determine expected credit losses for our notes

receivable using historical industry losses and considering loan-specific information, including credit ratings of the borrowers, estimated

fair values of underlying real estate assets, loan-to-value ratios, the presence of guarantors, and/or other available information. During

the three months ended March 31, 2026, no adjustment to the provision for expected credit losses related to our notes receivable was

required. The provision is evaluated on an ongoing basis, with any necessary adjustments recognized in the corresponding period.

34

9.FAIR VALUE MEASUREMENTS

We provide fair value information about all financial instruments for which it is practicable to estimate fair value. We measure

and disclose the estimated fair value of financial assets and liabilities by utilizing a fair value hierarchy that distinguishes between data

obtained from sources independent of the reporting entity and the reporting entity’s own assumptions about market participant

assumptions. This hierarchy consists of three broad levels, as follows: (i) quoted prices in active markets for identical assets or liabilities

(Level 1), (ii) significant other observable inputs (Level 2), and (iii) significant unobservable inputs (Level 3). Significant other observable

inputs can include quoted prices for similar assets or liabilities in active markets, as well as inputs that are observable for the asset or

liability, such as interest rates, foreign exchange rates, and yield curves. Significant unobservable inputs are typically based on an

entity’s own assumptions, since there is little, if any, related market activity. In instances in which the determination of the fair value

measurement is based on inputs from different levels of the fair value hierarchy, the level in the fair value hierarchy within which the

entire fair value measurement falls is based on the lowest level of input that is significant to the fair value measurement in its entirety.

Our assessment of the significance of a particular input to the fair value measurement in its entirety requires judgment and considers

factors specific to the asset or liability.

Assets and liabilities measured at fair value on a recurring basis

The following table sets forth the assets and liabilities that we measure at fair value on a recurring basis by level in the fair

value hierarchy as of March 31, 2026 and December 31, 2025 (in thousands). There were no transfers of assets measured at fair value

on a recurring basis to or from Level 3 in the fair value hierarchy during the three months ended March 31, 2026.

Fair Value Measurement Using
Description Total Quoted Prices in<br><br>Active Markets<br><br>for Identical Assets<br><br>(Level 1) Significant<br><br>Other<br><br>Observable Inputs<br><br>(Level 2) Significant<br><br>Unobservable<br><br>Inputs<br><br>(Level 3)
Assets:
Investments in publicly traded companies:
As of March 31, 2026 $102,076 $102,076 $— $—
As of December 31, 2025 $94,928 $94,928 $— $—
Cross-currency swap agreements:
As of March 31, 2026 $2,409 $— $2,409 $—
Liabilities:
Cross-currency swap agreements:
As of December 31, 2025 $928 $— $928 $—

Our investments in publicly traded companies represent investments with readily determinable fair values, and are carried at

fair value, with changes in fair value classified in investment income (loss) in our consolidated financial statements. We also hold

investments in privately held entities, which consist of (i) investments that report NAV and (ii) investments that do not report NAV, as

further described below.

Our investments in privately held entities that report NAV, such as our privately held investments in limited partnerships, are

carried at fair value using NAV as a practical expedient, with changes in fair value classified in net income. As of March 31, 2026 and

December 31, 2025, the carrying values of investments in privately held entities that report NAV aggregated $535.0 million and

$512.4 million, respectively. These investments are excluded from the fair value hierarchy above as required by the fair value

accounting standard. We estimate the fair value of each of our investments in limited partnerships based on the most recent NAV

prepared by the general partner and reported by each limited partnership. As a result, the determination of fair values of our

investments in privately held entities that report NAV generally does not involve significant estimates, assumptions, or judgments on our

part.

Our cross-currency swap agreements are recognized at fair value. Refer to Note 2 – “Summary of significant accounting

policies” and Note 11 – “Hedge agreements” to our unaudited consolidated financial statements for additional information.

35

9.FAIR VALUE MEASUREMENTS (continued)

Assets and liabilities measured at fair value on a nonrecurring basis

The following table sets forth the assets measured at fair value on a nonrecurring basis by level within the fair value hierarchy

as of March 31, 2026 and December 31, 2025 (in thousands).

Fair Value Measurement Using
Description Carrying Amount Quoted Prices in<br><br>Active Markets<br><br>for Identical Assets<br><br>(Level 1) SignificantOtherObservable Inputs(Level 2) Significant<br><br>Unobservable<br><br>Inputs<br><br>(Level 3)
Real estate assets held for sale with carrying values<br><br>adjusted to fair value less costs to sell:
As of March 31, 2026 140,777 $— $140,777 (2)
As of December 31, 2025 581,737 $— $581,737 (2)
Investments in privately held entities that do not<br><br>report NAV:
As of March 31, 2026 137,303 $— 125,917 $11,386 (4)
As of December 31, 2025 139,251 $— 123,238 $16,013 (4)

All values are in US Dollars.

(1)These amounts are included in the total balances of our net assets classified as held for sale aggregating $522.5 million and $581.7 million as of March 31, 2026 and

December 31, 2025, respectively, disclosed in Note 3 – “Investments in real estate,” and represent assets held for sale as of March 31, 2026 and December 31, 2025,

for which impairments were recognized.

(2)These amounts represent the aggregate carrying amounts of assets held for sale after adjustments to their respective fair values less costs to sell based on executed

purchase and sale agreements, letters of intent, valuations provided by third-party real estate brokers, or market comparables from recent transactions.

(3)These amounts represent the total carrying amounts of our equity investments in privately held entities with observable price changes, which are included in the

investments balances of $1.54 billion and $1.50 billion in our unaudited consolidated balance sheets as of March 31, 2026 and December 31, 2025, respectively,

disclosed in Note 7 – “Investments” to our unaudited consolidated financial statements.

(4)These amounts are included in the investments in privately held entities without observable price changes balances aggregating $405.6 million and $413.3 million as of

March 31, 2026 and December 31, 2025, respectively, disclosed in Note 7 – “Investments” to our unaudited consolidated financial statements, and represent the carrying

amounts of investments in privately held entities that do not report NAV for which impairments have been recognized in accordance with the measurement alternative

guidance described in “Investments” in Note 2 – “Summary of significant accounting policies” to our unaudited consolidated financial statements.

Real estate assets classified as held for sale measured at fair value less costs to sell

Our real estate assets classified as held for sale and measured at fair value less costs to sell are presented in the table above.

These properties represent a subset of our total real estate assets classified as held for sale as of March 31, 2026 and December 31,

  1. The fair values for these real estate assets were estimated based on executed purchase and sale agreements, letters of intent,

valuations provided by third-party real estate brokers, or market comparables from recent transactions. Refer to “Investments in real

estate” in Note 2 – “Summary of significant accounting policies” and “Assets held for sale” in Note 3 – “Investments in real estate” to our

unaudited consolidated financial statements for additional information.

Investments in privately held entities that do not report NAV

Our investments in privately held entities that do not report NAV are measured at cost, adjusted for observable price changes

and impairments, with changes recognized in net income (loss). These investments are adjusted based on the observable price

changes in orderly transactions for the identical or similar investment of the same issuer. Further adjustments are not made until

another observable transaction occurs. Therefore, the determination of fair values of our investments in privately held entities that do

not report NAV does not involve significant estimates and assumptions or subjective and complex judgments.

We also subject our investments in privately held entities that do not report NAV to a qualitative assessment for indicators of

impairment. If indicators of impairment are present, we are required to estimate the investment’s fair value and immediately recognize

an impairment charge in an amount equal to the investment’s carrying value in excess of its estimated fair value.

The estimates of fair value typically incorporate valuation techniques that include an income approach reflecting a discounted

cash flow analysis, and a market approach that includes a comparative analysis of acquisition multiples and pricing multiples generated

by market participants. In certain instances, we may use multiple valuation techniques for a particular investment and estimate its fair

value based on an average of multiple valuation results.

Refer to Note 7 – “Investments” to our unaudited consolidated financial statements for additional information.

36

9.FAIR VALUE MEASUREMENTS (continued)

Assets and liabilities not measured at fair value in the statement of financial position but for which the fair value is disclosed

The fair value of our unsecured senior notes payable and the amounts outstanding on our unsecured senior line of credit and

commercial paper program were estimated using widely accepted valuation techniques, including discounted cash flow analyses using

significant other observable inputs such as available market information on discount and borrowing rates with similar terms, maturities,

and credit ratings. Because the valuations of our financial instruments are based on these types of estimates, the actual fair value of our

financial instruments may differ materially if our estimates do not prove to be accurate. Additionally, the use of different market

assumptions or estimation methods may have a material effect on the estimated fair value amounts.

As of March 31, 2026 and December 31, 2025, the book and estimated fair values of our unsecured senior notes payable and

the amounts outstanding under our unsecured senior line of credit and commercial paper program, including the level within the fair

value hierarchy for which the estimates were derived, were as follows (in thousands):

March 31, 2026
Book Value Fair Value Hierarchy Estimated<br><br>Fair Value
Quoted Prices in<br><br>Active Markets<br><br>for Identical Assets<br><br>(Level 1) Significant<br><br>Other<br><br>Observable Inputs<br><br>(Level 2) Significant<br><br>Unobservable<br><br>Inputs<br><br>(Level 3)
Liabilities:
Unsecured senior notes payable $11,166,009 $— $10,096,063 $— $10,096,063
Unsecured senior line of credit $— $— $— $— $—
Commercial paper program $1,353,986 $— $1,354,081 $— $1,354,081 December 31, 2025
--- --- --- --- --- ---
Book Value Fair Value Hierarchy Estimated<br><br>Fair Value
Quoted Prices in<br><br>Active Markets<br><br>for Identical Assets<br><br>(Level 1) Significant<br><br>Other<br><br>Observable Inputs<br><br>(Level 2) Significant<br><br>Unobservable<br><br>Inputs<br><br>(Level 3)
Liabilities:
Unsecured senior notes payable $12,047,394 $— $10,675,433 $— $10,675,433
Unsecured senior line of credit $— $— $— $— $—
Commercial paper program $353,161 $— $353,189 $— $353,189

The carrying values of cash and cash equivalents, restricted cash, tenant receivables, deposits, notes receivable, accounts

payable, accrued expenses, and other short-term liabilities approximate their fair value.

37

10.SECURED AND UNSECURED SENIOR DEBT

The following table summarizes our outstanding indebtedness and respective principal payments remaining as of March 31, 2026 (dollars in thousands):

Stated<br><br>Rate Interest<br><br>Rate(1) Maturity<br><br>Date(2) Principal Payments Remaining for the Periods Ending December 31, Unamortized<br><br>(Deferred<br><br>Financing<br><br>Cost),<br><br>(Discount)/<br><br>Premium
Debt 2026 2027 2028 2029 2030 Thereafter Principal Total
Unsecured senior line of credit and<br><br>commercial paper program(3) (3) 4.27% (3) 1/22/30 (3) $— $— $— $— $1,355,271 $— $1,355,271 $(1,285) $1,353,986
Unsecured senior notes payable 3.80% 3.96 4/15/26 (4) 350,000 350,000 (38) 349,962
Unsecured senior notes payable 3.95% 4.13 1/15/27 350,000 350,000 (426) 349,574
Unsecured senior notes payable 3.95% 4.07 1/15/28 425,000 425,000 (782) 424,218
Unsecured senior notes payable 4.50% 4.60 7/30/29 300,000 300,000 (749) 299,251
Unsecured senior notes payable 2.75% 2.87 12/15/29 400,000 400,000 (1,552) 398,448
Unsecured senior notes payable 4.70% 4.81 7/1/30 450,000 450,000 (1,594) 448,406
Unsecured senior notes payable 4.90% 5.05 12/15/30 700,000 700,000 (3,751) 696,249
Unsecured senior notes payable 3.375% 3.48 8/15/31 750,000 750,000 (3,543) 746,457
Unsecured senior notes payable 2.00% 2.12 5/18/32 900,000 900,000 (5,811) 894,189
Unsecured senior notes payable 1.875% 1.97 2/1/33 1,000,000 1,000,000 (6,023) 993,977
Unsecured senior notes payable 2.95% 3.07 3/15/34 800,000 800,000 (6,287) 793,713
Unsecured senior notes payable 4.75% 4.88 4/15/35 500,000 500,000 (4,385) 495,615
Unsecured senior notes payable 5.50% 5.66 10/1/35 550,000 550,000 (6,162) 543,838
Unsecured senior notes payable 5.25% 5.41 3/15/36 750,000 750,000 (11,130) 738,870
Unsecured senior notes payable 5.25% 5.38 5/15/36 400,000 400,000 (3,681) 396,319
Unsecured senior notes payable 4.85% 4.93 4/15/49 300,000 300,000 (2,727) 297,273
Unsecured senior notes payable 4.00% 3.91 2/1/50 390,801 390,801 5,463 396,264
Unsecured senior notes payable 3.00% 3.09 5/18/51 352,398 352,398 (4,454) 347,944
Unsecured senior notes payable 3.55% 3.64 3/15/52 475,406 475,406 (6,233) 469,173
Unsecured senior notes payable 5.15% 5.26 4/15/53 500,000 500,000 (7,316) 492,684
Unsecured senior notes payable 5.625% 5.71 5/15/54 600,000 600,000 (6,415) 593,585
Unsecured debt weighted-average interest<br><br>rate/Total 4.06% $350,000 $350,000 $425,000 $700,000 $2,505,271 $8,268,605 $12,598,876 $(78,881) $12,519,995

(1)Represents the weighted-average interest rate as of the end of the applicable period, including amortization of loan fees, amortization of debt premiums (discounts), and other bank fees.

(2)Reflects any extension options that we control.

(3)Refer to footnote 3 on the following page.

(4)In April 2026, we repaid our 3.80% unsecured senior notes payable upon maturity. No gain or loss was incurred in connection with this repayment.

38

10.SECURED AND UNSECURED SENIOR DEBT (continued)

The following table summarizes our unsecured senior debt and amounts outstanding under our unsecured senior line of credit

and commercial paper program as of March 31, 2026 (dollars in thousands):

Fixed-Rate<br><br>Debt Variable-Rate<br><br>Debt Weighted-Average
Interest Remaining<br><br>Term<br><br>(in years)
Total Percentage Rate(1)
Unsecured senior notes payable $11,166,009 $— 11,166,009 89.2% 4.03% 10.7
Unsecured senior line of credit<br><br>and commercial paper program 1,353,986 1,353,986 10.8 4.27 (2) 3.8 (3)
Total/weighted average $11,166,009 $1,353,986 12,519,995 100.0% 4.06% 10.0 (3)
Percentage of total debt 89.2% 10.8% 100%

All values are in US Dollars.

(1)Represents the weighted-average interest rate as of the end of the applicable period, including expense/income related to the amortization of loan fees, amortization of

debt premiums (discounts), and other bank fees.

(2)As of March 31, 2026, we had no outstanding balance on our unsecured senior line of credit and $1.35 billion of commercial paper notes outstanding.

(3)We calculate the weighted-average remaining term of our commercial paper notes by using the maturity date of our unsecured senior line of credit. Using the maturity

date of our outstanding commercial paper notes, the consolidated weighted-average maturity of our debt is 9.6 years. The commercial paper notes sold during the three

months ended March 31, 2026 were issued at a weighted-average yield to maturity of 4.08% and had a weighted-average maturity term of 13 days.

Issuance and repayments of unsecured senior notes payable

In February 2026, we completed tender offers to repurchase an aggregate debt principal amount of approximately $1.33 billion

of a portion of our outstanding 4.00% Senior Notes due 2050, 3.00% Senior Notes due 2051, and 3.55% Senior Notes due 2052. Cash

consideration paid was $952.2 million. The repurchase was primarily funded through the issuance of $750.0 million of 5.25% unsecured

senior notes due 2036, and approximately $200 million of short-term borrowings under our commercial paper program. In connection

with the debt repurchase, we recognized a gain on early extinguishment of debt aggregating $366.4 million, including the write-off of

unamortized debt issuance costs and other transaction-related costs.

In January 2026, we repaid $300.0 million of 4.30% unsecured senior notes payable upon maturity. No gain or loss was

incurred in connection with this repayment.

In April 2026, we repaid $350.0 million of 3.80% unsecured senior notes payable upon maturity. No gain or loss was incurred

in connection with this repayment.

$5.0 billion unsecured senior line of credit

As of March 31, 2026, our unsecured senior line of credit, which matures in 2030, including extension options under our

control, had aggregate commitments of $5.0 billion, and bore an interest rate of SOFR plus 0.835%. In addition to the cost of borrowing,

the unsecured senior line of credit is subject to an annual facility fee of 0.14% based on the aggregate commitments outstanding.

Based on achievement of certain annual sustainability metrics, the interest rate and facility fee rate are also subject to upward or

downward adjustments of up to four basis points with respect to the interest rate and up to one basis point with respect to the facility fee

rate.

During the three months ended March 31, 2026, we achieved certain annual sustainability targets, as described in our

unsecured senior line of credit agreement, which reduced the borrowing rate by four basis points for a one-year period to SOFR plus

0.835%, from SOFR plus 0.875%, and reduced the facility fee by one basis point to 0.14% from 0.15%. As of March 31, 2026, we had

no outstanding balance on our unsecured senior line of credit.

$2.50 billion commercial paper program

Our commercial paper program allows us to issue up to $2.50 billion of commercial paper notes that bear interest at short-term

fixed rates with a maturity of generally 30 days or less and a maximum maturity of 397 days from the date of issuance. This program is

back-stopped by our unsecured senior line of credit, and at all times we expect to retain a minimum undrawn amount of borrowing

capacity under our unsecured senior line of credit equal to the amount of commercial paper notes outstanding. We use the net

proceeds from the issuances of the notes for general working capital and other general corporate purposes, which may include, but are

not limited to, the repayment of other debt and selective development, redevelopment, or acquisition of properties. During the three

months ended March 31, 2026, the notes were issued at a weighted-average yield to maturity of 4.08% and had a weighted-average

maturity term of 13 days. As of March 31, 2026, we had $1.35 billion outstanding under our commercial paper program.

39

10.SECURED AND UNSECURED SENIOR DEBT (continued)

Interest expense

The following table summarizes interest expense for the three months ended March 31, 2026 and 2025 (in thousands):

Three Months Ended March 31,
2026 2025
Interest incurred $134,557 $130,941
Capitalized interest (69,973) (80,065)
Interest expense $64,584 $50,876

11. HEDGE AGREEMENTS

We have fixed-to-fixed cross-currency swap agreements designated as net investment hedges to mitigate the impact of

fluctuations in the USD–CAD exchange rate. The hedges were deemed effective on the commencement date. As of March 31, 2026,

the aggregate notional amount of the swaps was CAD $340.0 million, and the corresponding total USD notional was approximately

$246.8 million. Under the terms of the swap agreements, USD fixed interest amounts are payable to us and CAD fixed interest amounts

are payable to the counterparty. The swap agreements mature on April 30, 2026. As of March 31, 2026, the hedge relationships

remained highly effective. Refer to “Hedge accounting” in Note 2 – “Summary of significant accounting policies” to our unaudited

consolidated financial statements for additional information.

The tables below summarize the fair value of our cross-currency swap agreements designated as net investment hedges and

the impact on our consolidated financial statements. Amounts are presented in USD (in thousands).

Fair value of cross-currency swap agreements designated as net investment hedges

Balance Sheet Location March 31, 2026 December 31, 2025
Other assets $2,409 $—
Other liabilities $— $928

Effect on consolidated other comprehensive income

Location in Consolidated Statement of<br><br>Comprehensive Income Three Months Ended<br><br>March 31, 2026
Total unrealized gains recognized in other comprehensive<br><br>income Unrealized gains on foreign currency<br><br>translation, net $3,337

Effect on consolidated statements of operations

Location in Consolidated Statement of<br><br>Operations Three Months Ended<br><br>March 31, 2026
Total gain recognized in net income(1) Other income $2,779

(1)Represents net interest expense settlements and interest rate forward points excluded from assessment of hedge effectiveness. Refer to “Hedge accounting” in Note 2 –

“Summary of significant accounting policies” to our unaudited consolidated financial statements for additional information.

As of March 31, 2026, a majority of our assets in Canada were designated as held for sale. Unrealized gains or losses related

to our cross-currency swap agreements will be reclassified from accumulated other comprehensive income into net income upon sale

or substantially complete liquidation of our real estate investments in Canada.

40

12. ACCOUNTS PAYABLE, ACCRUED EXPENSES, AND OTHER LIABILITIES

The following table summarizes the components of accounts payable, accrued expenses, and other liabilities as of March 31,

2026 and December 31, 2025 (in thousands):

March 31, 2026 December 31, 2025
Accounts payable and accrued expenses $368,098 $510,580
Accrued construction 243,411 314,836
Acquired below-market leases 126,697 133,033
Conditional asset retirement obligations 34,267 34,342
Deferred rent liabilities 16,090 14,659
Operating lease liability 358,610 360,543
Unearned rent and tenant security deposits 860,519 876,252
Other liabilities 147,090 152,828
Total $2,154,782 $2,397,073

As of March 31, 2026 and December 31, 2025, our conditional asset retirement obligations primarily consisted of the soil and

groundwater remediation liabilities associated with certain properties. Some of our properties may contain asbestos or may be

subjected to other hazardous or toxic substances, which, under certain conditions, requires remediation. We engage independent

environmental consultants to conduct Phase I or similar environmental assessments at our properties. This type of assessment

generally includes a site inspection, interviews, and a public records review; asbestos, lead-based paint, and mold surveys; subsurface

sampling; and other testing. We recognize a liability for the fair value of a conditional asset retirement obligation when the fair value of

the liability can be reasonably estimated. In addition, environmental laws and regulations subject our tenants, and potentially us, to

liability that may result from our tenants’ routine handling of hazardous substances and wastes as part of their operations at our

properties. As of March 31, 2026, we are not aware of any additional environmental liability that we believe would require additional

disclosures or recognition in our consolidated financial statements.

41

13.EARNINGS PER SHARE

We grant two types of restricted stock awards: (i) restricted stock awards with nonforfeitable dividends and (ii) restricted stock

awards with forfeitable dividends.

Unvested restricted stock awards (“RSAs”) with nonforfeitable dividends are considered participating securities and included in

the computation of EPS using the two-class method. Under this method, we allocate net income (after amounts attributable to

noncontrolling interests) to common stockholders and these RSAs by using the weighted-average shares of each class outstanding for

quarter-to-date and year-to-date periods independently, based on their respective participation rights to dividends declared (or

accumulated) and undistributed earnings.

Unvested RSAs with forfeitable dividends do not qualify as participating securities under the two-class method because the

dividends are forfeited if the awards do not vest. As a result, undistributed earnings are not allocated to these awards prior to vesting,

and these awards have no effect on the computation of basic EPS while unvested. Once these awards vest, they are included in the

denominator of basic EPS, weighted for the portion of the reporting period they were vested. Prior to vesting, these awards are included

in the denominator of diluted EPS if they are dilutive, which is determined using the treasury stock method. Under this method,

incremental shares are calculated as the difference between the total unvested shares and the number of shares that could

hypothetically be repurchased using the assumed proceeds (including unrecognized compensation cost related to these awards).

These incremental shares are weighted for the portion of the reporting period they were unvested and are included in the diluted EPS

denominator only if their inclusion reduces EPS (i.e., if they are not antidilutive).

In addition, from time to time, we enter into forward equity sales agreements. We consider the potential dilution resulting from

the forward equity sales agreements on the EPS calculations. At inception, the agreements do not have an effect on the computation of

basic EPS as no shares are delivered until settlement. The common shares issued upon the settlement of the forward equity sales

agreements, weighted for the period these common shares were outstanding, are included in the denominator of basic EPS. To

determine the dilution resulting from the forward equity sales agreements during the period of time prior to settlement, we calculate the

number of weighted-average shares outstanding – diluted using the treasury stock method. As of March 31, 2026, no forward equity

sales agreements were outstanding.

The table below reconciles the numerators and denominators of the basic and diluted EPS computations for the three months

ended March 31, 2026 and 2025 (in thousands, except per share amounts):

Three Months Ended March 31,
2026 2025
Net income $398,377 $38,662
Net income attributable to noncontrolling interests (36,724) (47,601)
Net income attributable to unvested RSAs with nonforfeitable dividends (2,779) (2,660)
Numerator for basic and diluted EPS – net income (loss) attributable to Alexandria Real Estate<br><br>Equities, Inc.’s common stockholders $358,874 $(11,599)
Denominator for basic EPS – weighted-average shares of common stock outstanding 170,598 170,522
Dilutive effect of unvested RSAs with forfeitable dividends 269
Denominator for diluted EPS – weighted-average shares of common stock outstanding 170,867 170,522
Net income (loss) per share attributable to Alexandria Real Estate Equities, Inc.’s common<br><br>stockholders:
Basic $2.10 $(0.07)
Diluted $2.10 $(0.07)

42

14.STOCKHOLDERS’ EQUITY

Common equity transactions

Common stock repurchase program

On December 8, 2025, we announced that our Board of Directors authorized a new common stock repurchase program that

allows for the repurchase of up to $500.0 million of our common stock through December 31, 2026. This new program replaced our

prior stock repurchase program. As of the date of this report, no repurchases have been made under the new program and $500.0

million remains available for future share repurchases.

ATM common stock offering program

In February 2024, we entered into an ATM common stock offering program that allows us to sell up to an aggregate of

$1.50 billion of our common stock.

During the three months ended March 31, 2026, we had no activity under our ATM program. As of March 31, 2026, the

remaining aggregate amount available under our ATM program for future sales of common stock was $1.47 billion.

Dividends

During the three months ended March 31, 2026, we declared cash dividends on our common stock aggregating $125.5 million,

or $0.72 per share.

Accumulated other comprehensive loss

The change in accumulated other comprehensive loss attributable to Alexandria Real Estate Equities, Inc.’s stockholders

during the three months ended March 31, 2026 was entirely due to net unrealized losses of $1.5 million on foreign currency translation

related to our operations primarily in Canada.

The change in accumulated other comprehensive loss attributable to Alexandria Real Estate Equities, Inc.’s stockholders for

the three months ended March 31, 2026 was primarily due to unrealized foreign currency translation losses of $4.9 million related to our

operations in Canada and partially offset by $3.3 million of unrealized gains resulting from the changes in the fair value of our cross-

currency swap agreements due to the weakening of the Canadian dollar. Refer to Note 11 – “Hedge agreements” to our unaudited

consolidated financial statements for additional information.

Common stock, preferred stock, and excess stock authorizations

Our charter authorizes the issuance of 400.0 million shares of common stock, of which 170.7 million shares were issued and

outstanding as of March 31, 2026. Our charter also authorizes the issuance of up to 100.0 million shares of preferred stock, none of

which were issued and outstanding as of March 31, 2026. In addition, 200.0 million shares of “excess stock” (as defined in our charter)

are authorized, none of which were issued and outstanding as of March 31, 2026.

43

15.SEGMENT INFORMATION

We are a life science REIT focused on developing, redeveloping, and operating properties that provide space for lease to

tenants primarily in the life science industry. Our properties are leased predominantly through triple-net lease agreements and share

key characteristics, including generic and reusable improvements, consistent lease structures, and business and financial strategy. All

properties are located within North America, predominantly in the U.S., and operate within a comparable regulatory environment.

Operating segments

Our Chief Operating Decision Maker (“CODM”), represented by our Executive Chairman and our Chief Executive Officer,

evaluates operating results at the geographic market level to assess performance and allocate resources. Our operating segments align

with our markets, including Greater Boston, the San Francisco Bay Area, San Diego, and Seattle, among others. Regular market

performance updates are provided directly to the CODM. These updates include each market’s net operating income (“NOI”), which

serves as the profit or loss measure used by the CODM for performance assessment and resource allocation. NOI provides useful

information regarding performance of each market as it reflects income and expenses incurred in connection with real estate operations

in each market. This metric enables the CODM to evaluate the profitability and performance of each market on a consistent and

comparable basis, supporting decisions on capital resource allocation, including in connection with development, redevelopment,

acquisition, and disposition activities in each market.

Evaluation of economic similarity and aggregation of operating segments

In accordance with the segment reporting accounting standard, we evaluate the economic similarity of our operating

segments. Seven of our nine operating segments exhibit consistent long-term economic characteristics, including similar historical long-

term NOI margins, which are also expected to remain similar in the future. Additionally, these markets share similar operational

characteristics, including nature of services provided (i.e., leasing, operating, developing, and redeveloping life science properties),

tenant base (i.e., a variety of tenants involved in the life science industry), methods of operation (i.e., consistent lease structures,

property management practices, and business strategies), nature of the regulatory environment (consistent across North America,

where all our operating segments are located). Based on shared economic characteristics, we have aggregated our seven operating

segments into one reportable segment for segment reporting purposes. The remaining operating segments, which do not meet the

aggregation criteria and individually do not meet the quantitative thresholds to qualify as reportable segments, were included in the “all

other” category in the tables below.

The following table presents the reportable segment profit or loss measure, NOI, for the three months ended March 31, 2026

and 2025 (in thousands):

Three Months Ended March 31,
2026 2025
Reportable segment revenues:
Revenues from external customers $621,259 $699,199
Other income 4,535 6,527
Reportable segment total revenues 625,794 705,726
Reportable segment total rental operating expenses (212,658) (212,436)
Reportable segment net operating income (reportable segment profit or loss) $413,136 $493,290

Significant expenses included in the reportable segment profit or loss measure (i.e., NOI) are represented by the reportable

segment total rental operating expenses and are disclosed in the table above. These expenses primarily include property taxes, utilities,

repairs and maintenance, engineering, janitorial, and other costs.

44

15.SEGMENT INFORMATION (continued)

Presented below are reconciliations of the reportable segment total revenues to the consolidated revenues, the reportable

segment total rental operating expenses to consolidated rental operations, the reportable segment NOI to the consolidated net income,

and the reportable segment investments in real estate assets to the consolidated investments in real estate assets (in thousands):

Three Months Ended March 31,
2026 2025
Reconciliation of reportable segment revenues to consolidated total revenues:
Reportable segment total revenues $625,794 $705,726
All other revenues 45,228 52,432
Consolidated total revenues $671,022 $758,158
Reconciliation of reportable segment total rental operating expenses to consolidated rental<br><br>operations:
Reportable segment total rental operating expenses $(212,658) $(212,436)
All other rental operating expenses (11,484) (13,959)
Consolidated rental operations $(224,142) $(226,395)
Reconciliation of reportable segment net operating income to consolidated net losses:
Reportable segment net operating income (reportable segment profit or loss) $413,136 $493,290
All other revenues 45,228 52,432
All other rental operating expenses (11,484) (13,959)
Other items not allocated to segments:
General and administrative (34,685) (30,675)
Interest expense (64,584) (50,876)
Depreciation and amortization (305,441) (342,062)
Impairment of real estate (5,499) (32,154)
Equity in losses of unconsolidated real estate joint ventures (147) (507)
Investment loss (4,582) (49,992)
Gain on early extinguishment of debt 366,435
Gain on sale of real estate 13,165
Consolidated net income $398,377 $38,662 March 31, 2026 December 31, 2025
--- --- ---
Reconciliation of reportable segment assets to consolidated investments in real<br><br>estate assets:
Reportable segment investments in real estate $27,660,991 $27,510,082
All other investments in real estate 1,169,125 1,179,914
Consolidated investments in real estate $28,830,116 $28,689,996

16.SUBSEQUENT EVENTS

In April 2026, we repaid $350.0 million of 3.80% unsecured senior notes payable upon maturity. No gain or loss was incurred

in connection with this repayment.

45

ITEM 2. MANAGEMENT’S DISCUSSION AND ANALYSIS OF FINANCIAL CONDITION AND RESULTS OF OPERATIONS

Forward-looking statements

Certain information and statements included in this quarterly report on Form 10-Q, including, without limitation, statements

containing the words “forecast,” “guidance,” “goals,” “projects,” “estimates,” “anticipates,” “believes,” “expects,” “intends,” “may,” “plans,”

“seeks,” “should,” “targets,” or “will,” or the negative of those words or similar words, constitute “forward-looking statements” within the

meaning of Section 27A of the Securities Act of 1933, as amended, and Section 21E of the Securities Exchange Act of 1934, as

amended. Forward-looking statements involve inherent risks and uncertainties regarding events, conditions, and financial trends that

may affect our future plans of operations, business and financial strategy, results of operations, and financial position. A number of

important factors could cause actual results to differ materially from those included within or contemplated by the forward-looking

statements, including, but not limited to, the following:

•Operating factors, such as a failure to operate our business successfully in comparison to market expectations or in

comparison to our competitors, our inability to obtain capital when desired or refinance debt maturities when desired, and/

or a failure to maintain our status as a REIT for federal tax purposes;

•Market and industry factors, such as adverse developments concerning the life science industry and/or our tenants;

•Government factors, such as any unfavorable effects resulting from federal, state, local, and/or foreign government

policies, laws, and/or funding levels;

•Global factors, such as negative economic, social, political, financial, credit market, banking conditions, and/or regional

armed hostilities; and

•Other factors, such as climate change, cyber intrusions, and/or changes in laws, regulations, and financial accounting

standards.

This list of risks and uncertainties is not exhaustive. Additional information regarding risk factors that may affect us is included

under Part I; “Item 1A. Risk factors”; and “Item 7. Management’s discussion and analysis of financial condition and results of

operations” in our annual report on Form 10-K for the year ended December 31, 2025 and under respective sections in this quarterly

report on Form 10-Q. Readers of this quarterly report on Form 10-Q should also read our other documents filed publicly with the SEC

for further discussion regarding such factors.

46

Overview

We are a Maryland corporation formed in October 1994 that has elected to be taxed as a REIT for federal income tax

purposes. Alexandria Real Estate Equities, Inc. (NYSE: ARE), an S&P 500® company, is a best-in-class, mission-driven life science

REIT making a positive and lasting impact on the world. With our founding in 1994, Alexandria pioneered the life science real estate

niche. Alexandria is the preeminent and longest-tenured owner, operator, and developer of collaborative Megacampus ecosystems in

AAA life science innovation cluster locations, including Greater Boston, the San Francisco Bay Area, San Diego, Seattle, Maryland,

Research Triangle, and New York City. As of March 31, 2026, Alexandria has a total market capitalization of $20.44 billion and an asset

base that includes 35.8 million RSF of operating properties and 3.4 million RSF of Class A/A+ properties undergoing construction.

We develop dynamic Megacampus ecosystems that enable and inspire some of the world’s most brilliant minds and innovative

companies to create life-changing scientific and technological innovations. We believe in the utmost professionalism, humility, and

teamwork. Our tenants include multinational pharmaceutical companies; life science product, service, and device companies; public

and private biotechnology companies; advanced technologies companies; biomedical institutions; U.S. government institutions; and

others. Alexandria has a long-standing and proven track record of developing Class A/A+ properties clustered in highly dynamic and

collaborative Megacampus environments that enhance our tenants’ ability to successfully recruit and retain world-class talent and

inspire productivity, efficiency, creativity, and success. Alexandria also provides strategic capital to transformative life science

companies through our venture capital platform.

As of March 31, 2026:

•Investment-grade or publicly traded large cap tenants represented 55% of our annual rental revenue;

•Approximately 97% of our leases (on an annual rental revenue basis) contained effective annual rent escalations

approximating 3% that were either fixed or indexed based on a consumer price index or other index;

•Approximately 91% of our leases (on an annual rental revenue basis) were triple net leases, which require tenants to pay

substantially all real estate taxes, insurance, utilities, repairs and maintenance, common area expenses, and other

operating expenses (including increases thereto) in addition to base rent;

•Approximately 92% of our leases (on an annual rental revenue basis) provided for the recapture of capital expenditures

(such as HVAC maintenance and/or replacement, roof replacement, and parking lot resurfacing) that we believe would

typically be borne by the landlord in traditional office leases; and

•78% of our leasing activity during the last twelve months was generated from our existing tenant base.

A key element of our business and financial strategy is our unique focus on Class A/A+ properties primarily located in

collaborative Megacampus ecosystems in AAA life science innovation clusters. Our Megacampus ecosystems are designed for

optionality and scalability, offering our tenants a clear path to address their growth requirements, including through our future

developments and redevelopments. Strategically located near top academic and medical research institutions and equipped with

curated amenities and services and convenient access to transit, our Megacampus ecosystems are designed to support our tenants in

attracting and retaining top talent and in meeting our tenants’ growth needs, which we believe is a key driver of tenant demand for our

properties. Our strategy also includes drawing upon our deep, broad, and long-standing real estate and life science industry

relationships in order to retain tenants, identify and attract new and leading tenants, and source additional real estate.

47

Executive summary

Operating results

Three Months Ended March 31,
2026 2025
Net income (loss) attributable to Alexandria’s common stockholders – diluted:
In millions $358.9 $(11.6)
Per share $2.10 $(0.07)
Funds from operations attributable to Alexandria’s common stockholders – diluted, as<br><br>adjusted:
In millions $295.9 $392.0
Per share $1.73 $2.30

For additional information, refer to “Funds from operations and funds from operations, as adjusted, attributable to Alexandria

Real Estate Equities, Inc.’s common stockholders” under “Definitions and reconciliations.”

A best-in-class REIT with a high-quality, diverse tenant base, strong margins, and long lease terms

(As of March 31, 2026, unless stated otherwise)
Occupancy of operating properties 87.7%
Percentage of total annual rental revenue in effect from Megacampus platform 78%
Percentage of total annual rental revenue in effect from investment-grade or publicly traded large cap tenants 55%
Adjusted EBITDA margin for the three months ended March 31, 2026 66%
Percentage of leases containing annual rent escalations 97%
Weighted-average remaining lease term:
Top 20 tenants 9.9 years
All tenants 7.5 years
Strong tenant collections(1):
Rents and receivables for the three months ended March 31, 2026, collected as of the date of this report 99.9%

(1)Refer to “Tenant Collections” under “Definitions and reconciliations” for additional details.

Strong and flexible balance sheet with significant liquidity; top 15% credit rating ranking among all publicly traded U.S. REITs

•Net debt and preferred stock to Adjusted EBITDA of 6.8x and fixed-charge coverage ratio of 3.4x for the three months ended

March 31, 2026 annualized, with targets for the three months ending December 31, 2026 annualized of 5.6x-6.2x and

3.6x-4.1x, respectively.

•We expect improvement in our quarter annualized net debt and preferred stock to Adjusted EBITDA ratio in the second

half of 2026 as we complete dispositions and sales of partial interests.

•As of March 31, 2026:

•Our credit ratings from S&P Global Ratings and Moody’s Ratings were BBB+ and Baa1, respectively, which rank in the top

15% among all publicly traded U.S. REITs.

•Significant liquidity of $4.17 billion, or 3.7x of our debt maturities through 2028.

•Only 9% of our total debt matures through 2028.

•10.0-year weighted-average remaining debt term, the longest among S&P 500 REITs.

•Total debt and preferred stock to gross assets of 31%.

48

Solid leasing of development and redevelopment space

•Leasing volume of 647,356 RSF during the three months ended March 31, 2026.

•Leasing of development and redevelopment space aggregating 117,935 RSF during the three months ended March 31,

2026, up 135% from the prior five quarter average, excluding a build-to-suit lease executed in July 2025 with a long-

standing multinational pharmaceutical tenant.

•From April 1, 2026 through the date of this report, we have executed leases and/or letters of intent aggregating

276,188 RSF related to our development and redevelopment pipeline.

•72% of our leasing activity during the three months ended March 31, 2026 was generated from our existing tenant base.

Three Months Ended<br><br>March 31, 2026
Leasing activity in RSF:
Leasing of development and redevelopment space 117,935
Leasing of previously vacant space 148,734
Lease renewals and re-leasing of space 380,687
647,356
Lease renewals and re-leasing of space:
Rental rate increase (15.0)%
Rental rate increase (cash basis) (15.8)%

•Excluding the impact of one lease aggregating 47,719 RSF at 480 Arsenal Street in our Cambridge/Inner Suburbs submarket,

rental rates for renewed and re-leased space for the three months ended March 31, 2026 would have decreased by 10.1%

and 9.1% (cash basis). The space at 480 Arsenal Street was re-leased to an entertainment studio user to accommodate their

expansion needs and secure a long-term extension. In addition, the reorientation of this building layout provides flexibility to

market the remaining available space to a broader range of user demand.

Key operating metrics

•Total revenues of $671.0 million, down 11.5%, for the three months ended March 31, 2026, compared to $758.2 million for the

three months ended March 31, 2025. Excluding dispositions completed after January 1, 2025, total revenues would have

decreased by 5.1% for the three months ended March 31, 2026 compared to the three months ended March 31, 2025.

•Net operating income (cash basis) of $1.7 billion for the three months ended March 31, 2026, annualized, decreased by

$300.6 million, or 15.2%, compared to the three months ended March 31, 2025, annualized.

•Change in net operating income (cash basis) reflects the impact of operating properties disposed of after January 1, 2025.

Excluding these dispositions, net operating income (cash basis), annualized, for the three months ended March 31, 2026,

would have decreased by 8.9%.

•Same property net operating income decreased by 11.9% and 11.7% (cash basis) for the three months ended March 31, 2026,

compared to the three months ended March 31, 2025.

•The quarter-over-quarter decline was due to a decrease in same property occupancy, primarily driven by the previously

disclosed 2026 key lease expirations aggregating 657,492 RSF that became vacant during the three months ended March

31, 2026, with a weighted-average lease expiration date of January 2026, and by vacancy during the three months ended

December 31, 2025 at one property aggregating 170,618 RSF at Alexandria Center® for Advanced Technologies – South

San Francisco in our South San Francisco submarket. We expect our same property performance to improve in the

second half of 2026, primarily due to changes in same property occupancy, including the anticipated delivery of 1.1 million

RSF of vacant space that was leased but not yet delivered as of March 31, 2026, which has a weighted-average expected

delivery date of approximately September 2026, and is expected to generate annual rental revenue of approximately $68

million.

•Same properties average occupancy for the three months ended March 31, 2026 was 88.9%, compared to 94.0% average

same properties occupancy for the three months ended March 31, 2025.

Continued successful reduction and management of general and administrative expenses

•General and administrative expenses for the three months ended March 31, 2026 aggregated $34.7 million, which represents

a decrease of $7.4 million, or 18%, compared to the quarterly average for 2024. For the trailing twelve months ended

March 31, 2026, general and administrative expenses as a percentage of net operating income were 6.0%, approximately half

the average of other S&P 500 REITs for 2023–2025.

•During the year ended December 31, 2025, we achieved general and administrative expense reduction of $51.3 million, or

30%, compared to the year ended December 31, 2024, primarily as a result of cost-control and efficiency initiatives. Some of

these cost savings were temporary, and we anticipate that approximately half of the cost reduction achieved in 2025 will

continue in 2026.

49

Dividend strategy to share net cash flows from operating activities with stockholders while retaining a significant portion for reinvestment

•Common stock dividend declared of $0.72 per share for the three months ended March 31, 2026, consistent with the

preceding quarter. The declared dividend per common share reflects our commitment to maintaining the strength of our

balance sheet, enhancing financial flexibility, preserving liquidity, and sharing cash flows with our stockholders.

•Significant net cash provided by operating activities, as adjusted, retained for reinvestment aggregating $2.60 billion for the

years ended December 31, 2022 through 2025 and the midpoint of our 2026 guidance range.

•Dividend yield of 6.2% as of March 31, 2026 and dividend payout ratio of 42% for the three months ended March 31, 2026.

Ongoing execution of Alexandria’s capital recycling strategy

We plan to continue funding a significant portion of our capital requirements for the year ending December 31, 2026 through

dispositions of land, non-core assets, and core assets (primarily sales of partial interests).

(dollars in millions) Sales Price
Completed and pending transactions subject to non-refundable deposits, signed letters of intent, and/or<br><br>sale agreement negotiations as of the date of this report 151
Identified and in process 2,181
Additional projected 568
2026 guidance midpoint for dispositions and sales of partial interests 2,900

All values are in US Dollars.

Occupancy and leasing progress on temporary vacancy

Operating occupancy as of December 31, 2025 90.9%
Reduction in occupancy related to previously disclosed key lease expirations during the three months ended<br><br>March 31, 2026 (1.9) (1)
Other changes in occupancy (1.3) (2)
Operating occupancy as of March 31, 2026 87.7
Vacant space leased but not yet delivered 3.2 (3)
Operating occupancy as of March 31, 2026, including vacant space leased but not yet delivered 90.9%

(1)Represents previously disclosed key lease expirations aggregating 657,492 RSF, with a weighted-average lease expiration date of January 2026 and prior annual rental

revenue of approximately $41.6 million. These vacant spaces are currently 48% leased or under negotiation and the remaining 52% is being actively marketed for re-

lease.

(2)Includes i) 139,408 RSF, or 0.4%, resulting from spaces vacated by tenants winding down operations, which are being actively marketed for re-lease and ii) delivery of

50,531 vacant RSF, or 0.2%, at our 10075 Barnes Canyon Road development project located at our SD Tech by Alexandria Megacampus.

(3)Represents temporary vacancies aggregating 1.1 million RSF, primarily in the Greater Boston, San Francisco Bay Area, and Seattle markets, that are leased and

expected to be occupied upon completion of building and/or tenant improvements. The weighted-average expected delivery date is approximately September 2026, with

expected annual rental revenue of approximately $68 million.

Key capital metrics as of or for the three months ended March 31, 2026

•$20.44 billion in total market capitalization.

•$7.92 billion in total equity capitalization.

•Non-real estate investments aggregating $1.54 billion:

•Unrealized gains presented in our consolidated balance sheet were $125.9 million, comprising gross unrealized gains and

losses aggregating $191.5 million and $65.6 million, respectively.

•Investment loss of $4.6 million for the three months ended March 31, 2026 presented in our consolidated statement of

operations consisted of $18.2 million of realized gains, $10.3 million of unrealized losses, and $12.4 million of impairment

charges.

50

Key capital events

•In February 2026, we completed tender offers to repurchase an aggregate debt principal amount of $1.33 billion across a

portion of our outstanding 4.00% Senior Notes due 2050, 3.00% Senior Notes due 2051, and 3.55% Senior Notes due 2052.

Cash consideration paid was $952.2 million. In connection with the debt repurchase, we recognized a gain on early

extinguishment of debt of approximately $366.4 million, including the write-off of unamortized debt issuance costs and other

transaction-related costs.

•We funded the repurchases as follows:

•$750.0 million through the issuance of 5.25% unsecured senior notes due 2036; and

•Approximately $200 million through short-term borrowings under our commercial paper program, which will be repaid

through planned dispositions and sales of partial interests included in our 2026 guidance.

•The repurchase reduced debt and improved leverage by approximately 0.2x.

•This transaction did not have a significant impact to our FFO per share diluted, as adjusted, interest expense, or fixed-

charge coverage ratio.

•Following this transaction, our weighted-average remaining term of debt as of March 31, 2026 is 10.0 years, which

continues to be the longest among S&P 500 REITs.

•In January 2026 and April 2026, we repaid, upon maturity, $300.0 million of 4.30% unsecured senior notes payable and

$350.0 million of 3.80% unsecured senior notes payable, respectively. These repayments were funded temporarily with

borrowings under our commercial paper program, which will be repaid through planned dispositions and sales of partial

interests included in our 2026 guidance. No gain or loss was incurred in connection with these repayments.

•Under our common stock repurchase program authorized in December 2025, we may repurchase up to $500.0 million of our

common stock through December 31, 2026. As of the date of this report, no shares have been repurchased.

Reduction of capital spend and funding needs

•We are evaluating the business and financial strategy for certain projects aggregating 1.6 million RSF to reduce future

construction funding requirements within our active pipeline.

•Driven by demand for our Megacampuses and access to amenities at our 311 Arsenal Street and 3000 Minuteman Road

redevelopment projects, we executed letters of intent aggregating 242,408 RSF in April 2026. These letters of intent are

for lower-cost alternative uses for all or a portion of these projects, including advanced technology. If we are successful in

executing these potential leases, we expect to evaluate whether all or a portion of these projects will be placed back into

operation without the need to further redevelop.

•Non-income-producing assets are 17% as a percentage of gross assets, a reduction of 3% since December 31, 2024;

targeting a further reduction to 11% to 16% by December 31, 2026.

Alexandria’s development and redevelopment pipeline is anticipated to deliver $92 million of incremental annual net operating income

by 4Q26 primarily from projects that are 93% leased/negotiating.

•Annual net operating income (cash basis) from recently delivered projects is expected to increase by $25 million upon the

burn-off of initial free rent, which has a weighted-average remaining period of approximately four months.

•77% of the RSF in our total development and redevelopment pipeline is within our Megacampus ecosystems.

Development and Redevelopment Projects IncrementalAnnual Net Operating Income RSF Leased/<br><br>Negotiating<br><br>Percentage
(dollars in millions)
Expected to be placed into service:
Second quarter of 2026 through fourth quarter of 2026 92 601,589 (2) 93% (3)
Fiscal years 2027 through 2028 93 1,258,004 68%
185

All values are in US Dollars.

(1)Includes expected partial deliveries through 2026 from projects expected to stabilize in 2027-2028, including speculative future leasing that is not yet fully

committed. Refer to the initial and stabilized occupancy years under “New Class A/A+ development and redevelopment properties: under construction” in Item 2 for

additional information.

(2)Represents the RSF of projects expected to stabilize in 2026. Does not include RSF for partial deliveries through 2026 from projects expected to stabilize in

2027-2028.

(3)Represents the current leased/negotiating percentage of the 601,589 RSF of development and redevelopment projects that are expected to stabilize in 2026.

51

Trends that may affect our future results

Currently identified key market trends and uncertainties that had or may have a negative effect on our business are discussed

below. Although we seek to minimize the risks posed by these trends and uncertainties as discussed in the mitigating factors section

below, there can be no assurance that these measures will be successful in preventing material impacts on our future results of

operations, financial position, and cash flows. Refer to “Item 1A. Risk factors” within Part I in our annual report on Form 10-K for the

year ended December 31, 2025 for discussion of additional risks we face.

New supply and reduced demand for life science space may continue to negatively affect our rental rates, occupancy, and

operating results.

•Influx of supply. During and after the COVID-19 pandemic, the shift toward hybrid and remote work arrangements as well as

exceptionally strong demand for life science space, driven by public health urgency and supported by historically low interest

rates, prompted certain office and other real estate investors to repurpose underutilized office spaces into laboratory facilities,

initiating a wave of new development activity across the sector. Our success and the success of other laboratory operators

prompted new and existing developers to commence speculative redevelopment and/or development laboratory projects in

anticipation of demand for such facilities. These conversion and speculative development projects have contributed to a

significant influx of new laboratory properties in our top three markets—Greater Boston, San Diego, and the San Francisco

Bay Area. Life science real estate availability in these top markets—measured as the percentage of life science RSF available

relative to total life science RSF—rose to approximately 29% during 2025, from approximately 4% in 2021. This surge created

supply that materially exceeded current demand. As pandemic-driven urgency faded, the amount of available space became

the dominant factor influencing tenant activity, with absorption unable to match the influx of supply.

•Decrease in demand. Adding to these challenges, life science tenant demand—after reaching historically high levels in 2021—

has moderated significantly. The average tenant demand, measured by life science tenants’ RSF requirements, has declined

by more than 60% during 2025 compared to 2021 across our top three markets: Greater Boston, San Francisco Bay Area, and

San Diego. This reflected a shift from extraordinary tenant demand driven by pandemic-related urgency to levels more

consistent with historical pre-pandemic norms, particularly those observed during 2016-2018. Importantly, this shift occurred

amid substantially higher available supply, as discussed above, further negatively impacting occupancy and rental rates in top

markets.

Exacerbating the recent demand trend, the life science industry faced an unusual convergence of macroeconomic, regulatory,

policy, and political challenges in 2025 which persist today, all of which are critical facets of the life science industry. These

included consequential shifts in leadership at the Health and Human Services agency (“HHS”), tariff-related measures,

operational, leadership, and staff disruptions at the NIH and the FDA, threatened reductions in NIH funding of biomedical

research and proposals to limit NIH funding of indirect grant costs, heightened scrutiny of pharmaceutical pricing, and

increased global competition from China, discussed below. Collectively, these factors, including those described below,

increased uncertainty, leading tenants to defer leasing commitments and expansion decisions pending greater clarity. As a

result, absorption of available space has been notably slower.

◦Prolonged biotech bear market and capital constraints. The life science sector experienced the fifth consecutive year

of a broad-based biotech bear market in 2025. Life science venture capital fundraising declined to its lowest level since

2016, reducing overall levels of venture capital funds available to deploy in the future. Life science venture funds also

continued to be highly risk averse, focusing investments on clinical-stage and asset-based opportunities that may not

drive significant laboratory space needs. The initial public offering market for biotech companies has remained largely

closed, eliminating a key source of liquidity and growth capital. Elevated interest rates have further constrained access to

debt and equity financing. These factors have slowed company formation, reduced headcount growth, and delayed

laboratory expansion decisions, directly impacting leasing demand for specialized life science space.

◦Regulatory and policy factors affecting absorption. At the same time, the regulatory environment experienced

significant disruption. The FDA saw more than 50% turnover in senior leadership during the first half of 2025,

accompanied by employee layoffs and delays in regulatory review decisions. Changing expectations related to clinical trial

requirements and flexibility for rare diseases with large unmet needs created additional uncertainty around development

timelines for certain regulated products. These conditions have reduced some tenants’ near-term confidence in expansion

and capital investment decisions.

Biomedical research institutions have faced increased uncertainty around federal funding policies throughout 2025. The

proposed 15% cap on NIH institutional indirect grant spending, subsequently ruled unlawful by an appellate court, raised

concerns for biomedical research institutions about the ability to recover infrastructure and operating costs, which

materially constrained incremental real estate demand among certain federally supported entities.

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Further, government actions aimed at reducing U.S. prescription drug prices have heightened uncertainty regarding future

returns on pharmaceutical and biotechnology investments. This has weighed on risk appetite across the sector and

constrained investment into some areas of research and development. As a result, some tenants have delayed or scaled

back expansion plans, reducing leasing activity and occupancy levels.

At the same time, global competition for life science research has intensified, with certain foreign markets, especially

China, rapidly gaining ground as biotechnology leaders through centralized funding and faster regulatory timelines.

Coupled with immigration-related restrictions implemented in the U.S. during 2025 that limit access to international

research talent, these policy actions not only affect current activities but also pose a significant threat to the long-term

viability of the U.S. biomedical industry. The cumulative effect of these developments may significantly reduce tenant

demand for U.S. life science real estate. Refer to “Item 1A. Risk factors” in this annual report on Form 10-K for more

information.

•Impact on our business. The surge in supply and decrease in demand have led to industry-wide elevated vacancy rates,

slower leasing activity, pressure on rental rates, higher lease concessions, and increased competition for tenants. Our

operating occupancy declined from 90.9% as of December 31, 2025 to 87.7% as of March 31, 2026, and we project our

operating occupancy to be approximately 87.0% as of December 31, 2026, representing the midpoint of our guidance range

for occupancy percentage in North America as of December 31, 2026.

To remain competitive, we have realized lower rental rate changes on renewed and re-leased spaces and have offered more

tenant improvement allowances or additional tenant concessions, including free rent, to retain existing tenants, or attract new

tenants. We project our rental rate changes on renewed and re-leased spaces to be approximately (5.0)% for the year ending

December 31, 2026, representing the midpoint of our guidance range. Furthermore, to maintain long-term tenant relationships

and sustain occupancy levels within our core assets, our existing operating properties may require additional revenue- and

non-revenue-enhancing capital expenditures earlier than typically expected.

The table below reflects a trend of increasing revenue- and non-revenue-enhancing capital expenditures, including tenant

improvement expenditures. The table also presents the trend, on a per RSF basis, of increasing tenant improvements, leasing

commissions, and free rent concessions, and of less favorable changes in rental rates related to our renewed/re-leased

spaces, as well as decreases in our operating occupancy (dollars in thousands, except per RSF amounts):

Revenue- and<br><br>Non-Revenue-<br><br>Enhancing<br><br>Capital<br><br>Expenditures Tenant<br><br>Improvements/<br><br>Leasing<br><br>Commissions<br><br>per RSF Free Rent<br><br>Concessions per<br><br>Annum<br><br>(leases executed in<br><br>trailing 12 months) Rental Rate<br><br>Changes<br><br>(on renewed/<br><br>re-leased<br><br>spaces) Operating<br><br>Occupancy<br><br>(as of each<br><br>period end)
2024 $273,377 $46.89 0.7 months 16.9% 94.6%
2025 $324,293 $55.34 1.5 months 7.0% 90.9%
Three months ended March 31, 2026 $164,382 $59.92 2.0 months (15.0)% 87.7%
Midpoint of 2026 guidance range $510,000 N/A (5.0)% 87.0%

Additionally, we have key lease expirations with expected downtime in 2026, primarily in the Greater Boston, San Francisco

Bay Area, and San Diego markets, aggregating 747,383 RSF as of March 31, 2026 with a weighted-average lease expiration

date of June 2026. These spaces are expected to become vacant at lease expiration and re-leased to new tenants. We expect

downtime on the 747,383 RSF to be approximately 6 to 24 months on a weighted-average basis. In addition, we have

identified 1.5 million RSF of key lease expirations in 2027 that are expected to have downtime of approximately 6 to 24 months

on a weighted-average basis. Considering elevated new laboratory supply in these markets, there can be no assurance that

we will be able to re-lease some or all of this space on acceptable terms, without significant capital expenditures, or within

anticipated time frames, even at reduced rates.

As of March 31, 2026, we anticipate that 1.9 million RSF of our projects undergoing construction will be placed into service

from April 1, 2026 through 2028 and will generate $185 million in future incremental annual net operating income. These

projects are 77% leased or under lease negotiations as of March 31, 2026. Furthermore, we have additional 1.6 million RSF of

projects under evaluation which are 23% leased or under lease negotiations. For these projects, we are evaluating business

and financial strategy, including continuing construction, repositioning for office or other non-laboratory use, selling, or pausing

development or redevelopment. If we decide to sell or pause, such actions could be dilutive to our funds from operations and

operating metrics. Alternatively, if we decide to invest limited capital, we may place some or all of these projects into

operations, which could temporarily reduce our operating occupancy until the projects are leased and occupied.

Landlord-funded tenant improvement allowances have increased significantly for first-generation space, including development

and redevelopment projects, with most space in shell condition requiring landlords to fund the full build-out cost. This trend

places additional pressure on projected returns and overall economics, and further challenges our ability to attract and secure

tenants for the remaining unleased RSF related to these projects at the expected rates, or at all, which could result in a

shortfall or delay in the commencement of the projected incremental annual net operating income.

53

Unfavorable macroeconomic and capital market conditions may continue to adversely affect the value of our real estate and

non-real estate portfolios, which could result in additional significant impairments and may impact our ability to raise capital

efficiently to further our business objectives.

The effective execution of our development and redevelopment activities is contingent on access to the capital required to fund

these projects. We expect funding for construction spending in 2026 to aggregate $1.75 billion at the midpoint of our 2026

guidance range for construction spending. This includes significant remaining construction costs to complete our active

pipeline and anticipated increases in both revenue- and non-revenue-enhancing capital expenditures in our operating portfolio.

As a result, our capital plan and leverage management strategy have increased our reliance on real estate dispositions and

sales of partial interests to generate capital. However, current real estate market conditions, including lower property

valuations and increased capitalization rates, will likely adversely affect the timing and pricing of such transactions.

•Lower property valuations and increased capitalization rates. A portion of our projected construction spending and other uses

of capital is expected to be funded through dispositions and sales of partial interests in core, land, and non-core real estate

assets. Real estate investments are generally less liquid than many other investment types, which can present challenges in

selling our properties in a timely manner or at desirable prices, especially in an environment of oversupply.

In addition to the factors discussed above specifically affecting demand for life science space, broader real estate demand has

also been impacted by macroeconomic conditions, particularly elevated interest rates. Following the onset of the COVID-19

pandemic, the U.S. Federal Reserve reduced the federal funds target range to 0%–0.25% in March 2020 and maintained that

near-zero range until March 2022. To address inflation concerns, the U.S. Federal Reserve then increased the target range

rapidly, reaching 5.25%–5.50% in July 2023, where it remained for an extended period. Although the U.S. Federal Reserve

reduced the federal funds target range to 4.25%–4.50% during 2024, and to 3.50%–3.75% during 2025, interest rates remain

elevated. This continues to limit access to debt and/or equity financing for prospective buyers of real estate assets. All other

aspects being equal, such challenges for buyers contribute to an excess of properties available for sale, which exerts

downward pressure on property valuations and elevates capitalization rates, adversely impacting the sales proceeds we can

generate from our real estate asset sales.

The oversupply of life science real estate assets, discussed above, combined with high interest rates and reduced market

liquidity, has contributed to a prolonged period of lower property valuations and higher capitalization rates, resulting in

significant real estate impairments and making it more challenging to execute asset sales within the expected timelines and at

favorable pricing. In 2026, we expect to complete dispositions and sales of partial interests of approximately $2.90 billion at the

midpoint of our 2026 guidance range. However, we may not be able to achieve this and/or other targets disclosed in our 2026

guidance as a result of the uncertainties discussed in this section as well as in “Item 1A. Risk factors” within “Part II – Other

information” of this quarterly report on Form 10-Q and “Item 1A. Risk factors” within Part I in our annual report on Form 10-K

for the year ended December 31, 2025.

The table below presents total dispositions and a trend of increasing impairments of real estate and capitalization rates

associated with dispositions and sales of partial interests in our real estate assets over the last several years (dollars in

thousands), which is partly attributable to the quality of core and non-core assets sold during each period.

Aggregate Sales Price<br><br>of Dispositions and<br><br>Sales of Partial<br><br>Interests Impairment of<br><br>Real Estate Capitalization<br><br>Rates(1) Capitalization<br><br>Rates<br><br>(Cash Basis)(1)
2024 $1,382,453 $223,068 7.7% 6.5%
2025 $1,813,778 $2,202,818 7.7% 7.5% (2)
Three months ended March 31, 2026 $— $5,499 N/A
Midpoint of 2026 guidance range $2,900,000 (3)

(1)Capitalization rates are calculated only for stabilized operating assets sold. Refer to “Capitalization rates” under “Definitions and reconciliations” for additional

information.

(2)Represents the weighted-average capitalization rate for stabilized operating assets sold in 2025, which accounted for only 20% of the aggregate sales price

of dispositions and sales of partial interests in 2025.

(3)We are not able to forecast impairments or capitalization rates for future periods without unreasonable effort due to the inherent difficulty of forecasting the

timing and amount of transactions that depend on market conditions outside of our control.

For additional information about our dispositions and real estate impairments recognized during the three months ended March

31, 2026, refer to “Sales of real estate assets and impairment of real estate” in Note 3 – “Investments in real estate” to our

unaudited consolidated financial statements in Item 1.

54

For 2026, we have established a disposition program with expected sales of approximately $2.90 billion at the midpoint of our

2026 guidance range for real estate dispositions and sales of partial interests.

In 2026 we are committed to dispose of certain assets classified as held for sale with an aggregate book value of

$522.5 million as of March 31, 2026. To achieve the midpoint of our 2026 guidance range of $2.90 billion for dispositions and

sales of partial interests, we continue to evaluate a significant number of disposition targets, including non-core operating

properties, both stabilized and unstabilized, and land parcels.

Under GAAP, real estate assets are evaluated for impairment upon indication of potential impairment:

•For real estate assets held and used, impairments are recognized if the sum of expected future undiscounted cash

flows, including estimated proceeds from eventual disposition, is less than the carrying amount. In such cases, the

carrying amount is reduced to estimated fair value.

•For real estate assets held for sale, impairments are recognized if fair value less costs to sell is less than the carrying

amount.

•In evaluating potential disposition targets that do not meet the criteria for held for sale classification, we apply a

probability-weighted approach, and in each case, no impairment charge is currently required.

If circumstances change, including changes in expected cash flows, capitalization rates, or market conditions, we may incur

additional material real estate impairments in 2026. For additional information on accounting for real estate impairments, refer

to “Impairment of long-lived assets” in Note 2 – “Summary of significant accounting policies” to our unaudited consolidated

financial statements in Item 1.

We expect to substantially complete our large-scale non-core asset sales program in 2026. As of March 31, 2026, 78% of our

annual rental revenue is from our Megacampus platform, and we expect this percentage to continue to grow over time, in part

through our disposition program.

•Increased cost and limited availability of capital. Our 2026 guidance assumes a reduction of our outstanding debt by

approximately $1.68 billion, at the midpoint of our 2026 guidance range.

•In February 2026, we completed tender offers to repurchase an aggregate debt principal amount of $1.33 billion

across a portion of our outstanding 4.00% Senior Notes due 2050, 3.00% Senior Notes due 2051, and 3.55% Senior

Notes due 2052. The tender offers were completed at an average discount of approximately 28%, for a total cash

payment of $952.2 million, resulting in the extinguishment of approximately $380 million of debt. We funded the

$952.2 million payment through the issuance of $750.0 million of 5.25% unsecured senior notes due 2036 and

approximately $200 million of short-term borrowings under our commercial paper program.

•In January 2026 and April 2026, we repaid, upon maturity, $300.0 million of 4.30% unsecured senior notes and

$350.0 million of 3.80% unsecured senior notes, respectively. These repayments, aggregating $650 million, were

temporarily funded through borrowings under our commercial paper program.

Although we repaid a portion of our outstanding debt during 2026, these repayments have been fully financed through the

issuance of new debt. As a result, we have not yet made progress toward our targeted $1.68 billion net debt reduction.

Accordingly, achievement of this target debt reduction remains dependent on our ability to generate proceeds during 2026

from planned real estate dispositions and sales of partial interests.

These expectations assume our ability to execute our planned real estate dispositions and partial interest sales on acceptable

terms. If we are unable to sell real estate assets at our targeted prices or within our expected timeframes, we may need to

reduce the projected amount of debt repayment, delay the timing of such repayment, and/or increase our reliance on additional

debt financing to fund the approximately $1.75 billion of construction spending, based on the midpoint of our 2026 guidance

range. Elevated interest rates may result in debt financing options that are costlier, less accessible, or even unavailable,

potentially limiting our ability to complete our development and redevelopment projects on schedule and thereby delaying our

expected incremental annual net operating income generation.

The table below reflects interest rates related to unsecured senior notes payable that we have issued over the last several

years and in February 2026 (dollars in thousands). There is no assurance that high debt costs will not continue into the future.

Unsecured Senior<br><br>Notes Payable Issued Interest Rate(1)
2024 $1,000,000 5.57%
2025 $550,000 5.66%
February 2026 issuance $750,000 5.41%

(1)Includes amortization of loan fees, amortization of debt premiums (discounts), and other bank fees.

55

•Capitalized Interest.

The table below presents gross interest expense, capitalized interest, and interest expense (in thousands):

Gross Interest Expense Capitalized Interest Interest Expense
2024 $516,799 $(330,961) $185,838
2025 $557,122 $(330,424) $226,698
Three months ended March 31, 2026 $134,557 $(69,973) $64,584
Midpoint of 2026 guidance range $505,000 $(245,000) $260,000

For 2026, we expect capitalized interest of approximately $245 million at the midpoint of our guidance range. The decrease

compared to 2025 reflects our actions taken in response to the market conditions, including re-evaluating certain projects,

ceasing or pausing certain pre-construction activities on land and uncommitted projects to conserve capital, and disposing of

certain assets. As a result, we expect our interest expense to increase to approximately $260 million (at the midpoint of our

2026 guidance range) in 2026 from $226.7 million in 2025. Continued macroeconomic and capital market pressures may

necessitate further reevaluation of our plans, including temporary suspension of our construction projects, delay of future

projects, or the sale of non-income-producing properties, which could further reduce our capitalized interest and increase

interest expense.

•Volatility in the valuation of non-real estate investments. We hold strategic investments in publicly traded companies and

privately held entities primarily involved in the life science industry. These investments are subject to market-and sector-

specific risks that can substantially affect their valuation. Like many other industries, the life science industry is susceptible to

macroeconomic challenges, such as ongoing economic and geopolitical uncertainty and a tighter capital environment. These

factors may lead to increased volatility in the valuation of our non-real estate investments.

In such an environment, distributions from our investments—which we may receive as dividends, as liquidation distributions

from our investments in limited partnerships, or as a result of mergers and acquisitions involving our privately held investees—

may be limited and could result in lower realized gains. Gross unrealized gains related to our non-real estate investments held

as of March 31, 2026, December 31, 2025, and December 31, 2024 aggregated to $191.5 million, $184.4 million, and $228.1

million, respectively. These unrealized amounts are subject to market fluctuations and may not ultimately be realized. We may

not receive distributions from our investments or may face difficulties in monetizing our non-real estate investments at optimal

prices. There can be no assurance that we will be able to realize gains in the future. In periods with limited or no realized

gains, our FFO per share, as adjusted, may be adversely affected.

For the three months ended March 31, 2026, we recognized $18.2 million in realized gains on non-real estate investments and

are projecting realized gains of $75 million in 2026 at the midpoint of our guidance range. During the three months ended

March 31, 2026, we also recognized impairment charges and unrealized losses that reflect continued valuation pressures. The

table below presents components of investment income (loss) on our non-real estate investments (in thousands):

Non-Real Estate Investments
Realized<br><br>Gains Significant<br><br>Realized<br><br>Losses Impairments Unrealized<br><br>(Losses)<br><br>Gains Investment<br><br>Loss
2024 $117,214 $— $(58,090) $(112,246) $(53,122)
2025 $115,722 $(103,329) $(95,716) $26,980 $(56,343)
Three months ended March 31, 2026 $18,198 $— $(12,448) $(10,332) $(4,582)
Midpoint of 2026 guidance range $75,000 N/A(1)

(1)We are not able to forecast investment income (loss) of future periods without unreasonable effort and therefore do not provide the information on a forward-

looking basis. This is due to the inherent difficulty of forecasting the timing and/or amount of items that depend on market conditions outside of our control.

Unfavorable market conditions could also lead to additional impairments of our investments in privately held entities that do not

report NAV per share, as well as other‑than‑temporary impairments of our non‑real‑estate investments accounted for under the

equity method.

56

The realization of any of the foregoing risks could continue to have material adverse impacts on our revenues and operating

performance, including but not limited to our income from rentals, net operating income, results of operations, funds from operations,

operating margins, initial stabilized yields (unlevered) on new or existing construction projects, occupancy, rental rates, EPS, FFO per

share, FFO per share, as adjusted, and net cash provided by operating activities, as adjusted. These impacts have adversely affected,

and could continue to adversely affect, our Adjusted EBITDA, which in turn may continue to negatively impact our key metrics such as

Adjusted EBITDA margin, net debt and preferred stock to Adjusted EBITDA, and fixed-charge coverage ratios. This may also impact

our credit ratings and credit rating outlooks. To preserve liquidity and mitigate an increase to our net debt and preferred stock to

Adjusted EBITDA ratio resulting from declines in Adjusted EBITDA, we may seek additional capital by pursuing additional sales of real

and non-real estate assets, or through equity offerings, which could be dilutive to existing stockholders. A reduction in earnings and/or

net cash provided by operating activities, as adjusted, could potentially necessitate or make advisable a reduction in our dividends per

share, as determined by our board of directors. Any of the foregoing could further negatively affect our business and the market value

of our common stock.

•Mitigating factors:

•Reinforcing the Megacampus platform as our core growth engine. We believe our Megacampus strategy represents

our most powerful competitive advantage in an oversupplied life science real estate market. Our Megacampus

ecosystems are large-scale environments designed to meet the evolving needs of the world’s leading scientific and

technological organizations, located in life science innovation hubs in close proximity to top academic and medical

research institutions. This proximity is a key driver of tenant demand. These campuses are used in two distinct ways: (i) to

house the research operations of our tenants, and (ii) to recruit and retain the best talent available from a limited pool,

which underscores why their scale, strategic design, and location are critical. With our Megacampus ecosystems, we aim

to provide a superior set of amenities, services, and access to transit. With inspiring design and people-centric amenities,

we believe these campuses enhance our tenants’ confidence in using these spaces as effective recruiting tools. In

contrast, we believe that a significant amount of the competitive supply in the market today consists of isolated facilities

that provide operational space but lack the scale and strategic design that our Megacampus ecosystems deliver.

Our Megacampus ecosystems, which offer both high visibility and a clear path for growth, are designed for scalability to

accommodate our tenants’ growth. Our future development and redevelopment projects aggregate 21.9 million RSF as of

March 31, 2026, of which 77% is concentrated within our Megacampus ecosystems. Their strategic locations and path for

growth serve as powerful incentives for tenants to lease space from us.

We believe our Megacampus strategy has enabled us to capture a greater share of available leasing demand relative to

competitors in our core life science markets, even as overall supply has increased. The strength of this strategy is

reflected in the 2026 performance metrics below, achieved despite challenging macroeconomic, regulatory, policy, and

geopolitical environments:

•Occupancy of 87.7% as of March 31, 2026:

•Outperforms market occupancy levels in our top three markets: Greater Boston, San Francisco Bay Area, and

San Diego.

•Additional 3.2% occupancy is expected from temporary vacancies aggregating 1.1 million RSF (3.2% of total

operating RSF) as of March 31, 2026. These vacant spaces are currently leased and are expected to become

occupied upon completion of building and/or tenant improvements, with a weighted‑average expected occupancy

date of September 2026. These spaces are primarily located in our Greater Boston, San Francisco Bay Area,

and Seattle markets and are expected to generate annual rental revenue of approximately $68 million upon lease

commencement.

•93% of development and redevelopment projects under construction that are expected to stabilize in 2026 are leased

or under lease negotiation.

•Expected incremental annual net operating income from projects anticipated to be placed into service from the

second quarter of 2026 to the end of 2028:

•$92 million from deliveries in remaining quarters of 2026.

•$93 million from 2027-2028 deliveries.

•Strength of our brand. As a recognized leader in the life science and real estate sectors, Alexandria has successfully

built a diverse and high-quality tenant base. Over the past three decades, we have fostered long-standing relationships

and strategic partnerships with our tenants, which have enabled us to maintain strong occupancy levels and leasing

volume, and growth in net operating income and cash flows and to effectively navigate through various economic cycles.

Key indicators of our brand strength include the following:

•As of March 31, 2026, 78% of our leasing activity during the last twelve months was from our existing tenant base.

•As of March 31, 2026, 87% of our top 20 tenant annual rental revenue was derived from investment-grade or publicly

traded large cap companies.

•Our tenant collections have remained consistently high, averaging 99.8% since the beginning of 2021 through

March 31, 2026.

57

•Prudent financial management. Our strong and flexible balance sheet and prudent balance sheet management are key

factors in our ability to navigate macroeconomic uncertainties and capitalize on new opportunities. The strength of our

financial position is highlighted by several key indicators:

•Our significant liquidity of $4.17 billion as of March 31, 2026 provides us the flexibility to address our operational

needs and to pursue strategic opportunities.

•We expect to self-fund a large portion of our capital requirements through the following sources in 2026:

•$525 million in net cash provided by operating activities, as adjusted, at the midpoint of our 2026 guidance range.

•$120.0 million in capital contributions to fund construction expected from our existing consolidated real estate

joint venture partners from April 1, 2026 through 2027 and beyond.

•$2.90 billion from dispositions and sales of partial interests in real estate assets at the midpoint of our 2026

guidance range.

•As of March 31, 2026, our credit ratings from S&P Global Ratings and Moody’s Ratings were BBB+ and Baa1,

respectively, which rank in the top 15% among all publicly traded U.S. REITs.

•Net debt and preferred stock to Adjusted EBITDA ratio target is 5.6x to 6.2x for the fourth quarter of 2026, annualized.

•As of March 31, 2026, our fixed-rate debt represents 89.2% of our total debt, which provides predictability in debt

servicing costs. Since 2022, our quarter-end fixed-rate debt has averaged 96.4%.

•Our debt maturity schedule is well laddered, which provides us with financial flexibility and reduces short-term

refinancing risks. As of March 31, 2026, only 9% of our debt matures through 2028.

•As of March 31, 2026, the weighted-average remaining term of our debt is 10.0 years, which is the longest among

S&P 500 REITs, and demonstrates our strategic approach to debt management and our focus on maintaining

manageable annual debt maturities.

•Operational excellence of our team. Alexandria focuses on operational excellence in the direct asset management and

operations of our Labspace® asset base. Our asset management and operations team is composed of highly experienced,

educated, and professionally credentialed facilities specialists. This expertise, essential in ensuring a secure and efficient

environment for groundbreaking scientific research, has been cultivated and maintained over many years.

The demanding nature of laboratory-based scientific research requires strict adherence to safety standards set by local,

state, and federal regulatory bodies. Key compliance aspects include good manufacturing practices (“GMP”) and Clinical

Laboratory Improvement Amendments (“CLIA”) certifications, adherence to national biosafety level guidelines, proper

permitting and handling of hazardous waste generation and chemical storage, maintenance of safety stations, effective

management of ultra-low temperature freezers, and careful licensing and management of radioactive materials.

•Other mitigating factors

•Improvement in office market. The increase in demand for premium office space since 2024, primarily driven by the

technology sector, particularly companies focused on AI, absorbed some of the market’s supply previously anticipated

for life science use and is now being repositioned back into office space. High ceilings, improved ventilation systems,

and abundant natural light, which are all features of life science real estate, have become highly desirable, appealing

to office and advanced technologies tenants. We expect this trend may lead to the exit from the life science sector of

inexperienced life science real estate developers and expedite the resolution of the oversupply impacting the sector.

•Proactive reduction in capital spending and funding needs. To address higher capital costs and slower market

absorption, we implemented a disciplined reduction in construction spending. Based on the midpoint of our 2026

guidance range, our average annual construction spending is expected to decrease to approximately $1.74 billion for

2024–2026, representing a reduction of approximately $1.02 billion, or 37%, compared to the 2021–2023 average.

Our 2026 construction spending is primarily focused on:

•Leasing vacant space at operating properties

•Completing active committed construction projects

•Limiting future pipeline pre-construction activity

This strategy supports a more self-funded capital plan while preserving flexibility for future growth opportunities.

•Decrease in general and administrative expenses. Over the past several years, we have implemented comprehensive

measures to reduce our expenditures across our organization, including our general and administrative expenses, by

implementing a variety of cost-control and efficiency initiatives, including, but not limited to:

•Personnel-related matters, including:

•Reduction in headcount over the last two years.

•Restructuring of various compensation plans.

•Streamlining of business processes:

•Implementation of systems upgrades, process improvements, and smarter technology.

•Renegotiation of contracts related to legal, technology, and operational support services, and

elimination of redundancies through better alignment and consolidation of roles.

58

As a result, we have achieved the following outcomes:

•During the three months ended March 31, 2026, general and administrative expenses aggregated

$34.7 million, which represents a decrease of $7.4 million, or 18%, compared to the quarterly average for

2024.

•We expect $76 million of cumulative savings in 2025 and 2026 (based upon the midpoint of our guidance

range for 2026 general and administrative expenses), compared to 2024.

•For the trailing twelve months ended March 31, 2026, our general and administrative expenses as a

percentage of net operating income were 6.0%, approximately half the average of other S&P 500 REITs for

2023–2025.

We believe the mitigating factors discussed above will help us manage prolonged market volatility while maintaining the

flexibility to act on strategic opportunities. Through disciplined execution of non-core asset recycling, targeted capital

allocation, continued focus on our Megacampus platform, moderated construction spending, and preservation of balance sheet

strength, we are building a resilient platform designed to deliver sustainable future growth and value creation across multiple

cycles. We believe these actions position us to emerge from the current cycle in a position of strength.

59

Operating summary

Same Property Performance:<br><br>Net Operating Income Changes Rental Rate Changes:<br><br>Renewed/Re-Leased Space
Margins(3) Favorable Lease Structure(4)
Operating Adjusted EBITDA Strategic Lease Structure by Owner and<br><br>Operator of Collaborative Megacampus Ecosystems
67% 66% Increasing cash flows
Percentage of leases containing annual<br><br>rent escalations 97%
Stable cash flows
Long-Duration Lease Terms(5) Percentage of triple net leases 91%
9.9 Years 7.5 Years Lower capex burden
Percentage of leases providing for the<br><br>recapture of capital expenditures 92%
Top 20 Tenants All Tenants
Net Debt and Preferred Stock<br><br>to Adjusted EBITDA(6) Fixed-Charge Coverage Ratio(6)

13

37

25

1

(2)

(1)

(1)

(2)

49

5.6x to 6.2x

61

3.6x to 4.1x

Refer to “Same properties” and “Definitions and reconciliations” in Item 2 for additional details. “Definitions and reconciliations” contains the definitions of “Fixed-charge

coverage ratio,” “Net debt and preferred stock to Adjusted EBITDA,”, “Net operating income”, and “Adjusted EBITDA” and their respective reconciliations from the most directly

comparable financial measures presented in accordance with GAAP.

(1)Refer to footnote 1 under “Same properties” in Item 2 for additional details.

(2)Refer to footnote 2 under “Leasing activity” in Item 2 for additional details.

(3)For the three months ended March 31, 2026.

(4)Percentages calculated based on our annual rental revenue in effect as of March 31, 2026.

(5)Represents the weighted-average remaining term based on annual rental revenue in effect as of March 31, 2026.

(6)Quarter annualized.

60

Stable Cash Flows From Our High-Quality, Diverse Tenant Mix
Investment-Grade or Publicly Traded Large Cap Tenants
87% 55%
of ARE’s Top 20 Tenant<br><br>Annual Rental Revenue of ARE’s Total Annual<br><br>Rental Revenue

8796093022384

1

(1)

(2)

(3)

Percentage of ARE’s

Annual Rental Revenue

As of March 31, 2026. Annual rental revenue represents amounts in effect as of March 31, 2026. Refer to “Definitions and reconciliations” in Item 2 for additional information.

(1)Represents the percentage of our annual rental revenue generated by professional services, finance, construction/real estate companies, and retail-related tenants.

(2)76% of our annual rental revenue from advanced technologies tenants is from investment-grade or publicly traded large cap tenants.

(3)81% of our annual rental revenue from biomedical institutions is from investment-grade or publicly traded large cap tenants.

61

Leasing activity

The following table summarizes our leasing activity at our properties:

Three Months Ended Year Ended
March 31, 2026 December 31, 2025
(Dollars per RSF) Including Straight-Line Rent Cash Basis Including Straight-Line Rent Cash Basis
Leasing activity:
Renewed/re-leased space(1)
Rental rate changes (15.0)% (15.8)% 7.0% 3.5%
New rates 49.88 53.43 52.71 $53.66
Expiring rates 58.67 63.42 49.27 $51.87
RSF 380,687 2,543,473
Tenant improvements/leasing commissions 59.92 55.34
Weighted-average lease term 8.4 years 9.0 years
Previously vacant/developed/redeveloped space leased
New rates 53.34 52.10 72.30 $67.56
Previously vacant RSF 148,734 944,362
Developed/redeveloped RSF(3) 117,935 704,821
Weighted-average lease term 14.0 years 13.8 years
Leasing activity summary (totals):
New rates 51.30 52.88 60.42 $59.13
RSF 647,356 4,192,656
Weighted-average lease term 12.0 years 11.9 years
Lease expirations(1)
Expiring rates 56.43 62.28 54.22 $55.56
RSF 1,340,809 4,460,081

All values are in US Dollars.

Leasing activity includes 100% of results for properties in North America in which we have an investment.

(1)Excludes month-to-month leases aggregating 103,632 RSF and 58,516 RSF as of March 31, 2026 and December 31, 2025, respectively. During the trailing twelve

months ended March 31, 2026, we granted free rent concessions averaging 2.0 months per annum.

(2)Includes the impact of one lease aggregating 47,719 RSF at 480 Arsenal Street in our Cambridge/Inner Suburbs submarket, which was signed to an entertainment

studio user to accommodate their expansion needs and secure a long-term extension. The reorientation of the building layout needed to accommodate the tenant also

provides flexibility to market the remaining available space to a broader range of user demand. Excluding this lease, rental rates for renewed and re-leased space for the

three months ended March 31, 2026 would have decreased by 10.1% and 9.1% (cash basis) and tenant improvements and leasing commissions would have been

$46.18 per RSF.

(3)Refer to “New Class A/A+ development and redevelopment properties: summary of pipeline” in Item 2 for additional information, including total project costs.

(4)Includes the largest life science lease in company history, executed in July 2025 with a long-standing multinational pharmaceutical tenant. The 16-year expansion

build-to-suit lease aggregates 466,598 RSF and is located at the Campus Point by Alexandria Megacampus in our University Town Center submarket. Excluding

this lease, developed/redeveloped space for the year ended December 31, 2025 was 238,223 RSF.

(5)Includes previously disclosed 2026 key lease expirations aggregating 657,492 RSF that became vacant during the three months ended March 31, 2026, with a

weighted-average lease expiration date of January 2026.

62

Summary of contractual lease expirations

The following table summarizes the contractual lease expirations at our properties as of March 31, 2026:

Year RSF Percentage of<br><br>Occupied RSF Annual Rental Revenue<br><br>(per RSF)(1)
2026 (2) 1,485,686 4.9% 55.84 4.6%
2027 3,367,997 11.2% 56.51 10.5%
2028 3,687,608 12.3% 49.66 10.1%
2029 1,831,388 6.1% 44.65 4.5%
2030 2,446,190 8.1% 42.70 5.8%
2031 3,562,366 11.8% 54.91 10.8%
2032 886,704 2.9% 57.83 2.8%
2033 2,177,834 7.2% 50.72 6.1%
2034 2,727,148 9.1% 66.91 10.1%
2035 1,042,126 3.5% 58.19 3.4%
Thereafter 6,868,641 22.9% 82.42 31.3%

All values are in US Dollars.

Contractual lease expirations for properties classified as held for sale as of March 31, 2026 are excluded from the information on this page.

(1)Amounts in effect as of March 31, 2026.

(2)Excludes month-to-month leases aggregating 103,632 RSF as of March 31, 2026.  Refer to “Leasing activity” in Item 2 for additional details.

63

The following tables present our lease expirations by market for the remainder of 2026 and for 2027, as of March 31, 2026:

2026 Contractual Lease Expirations (in RSF)
Market Leased Negotiating/<br><br>Anticipating Remaining<br><br>Expiring<br><br>Leases Total(2) Annual Rental<br><br>Revenue<br><br>(per RSF)(1)
Greater Boston 119,822 11,894 110,339 242,055 $42.93
San Francisco Bay Area 4,908 21,671 115,876 142,455 74.74
San Diego 104,945 104,945 57.70
Seattle 10,553 12,908 39,002 62,463 24.74
Maryland 8,155 48,302 56,457 26.82
Research Triangle 41,518 11,913 19,783 73,214 43.58
New York City 36,501 36,501 103.49
Texas
Non-cluster/other markets 20,213 20,213 57.70
Subtotal 184,956 58,386 494,961 738,303 51.86
Key lease expirations with expected downtime 17,271 81,642 648,470 747,383 (3) 59.77
Total 202,227 140,028 1,143,431 1,485,686 $55.84
Percentage of expiring leases 14% 9% 77% 100%
2027 Contractual Lease Expirations (in RSF)
Market Leased Negotiating/<br><br>Anticipating Remaining<br><br>Expiring<br><br>Leases Total Annual Rental<br><br>Revenue<br><br>(per RSF)(1)
Greater Boston 50,375 24,386 110,518 185,279 $67.27
San Francisco Bay Area 1,873 8,927 183,000 193,800 72.78
San Diego 62,415 335,515 397,930 45.00
Seattle 9,435 93,839 305,254 408,528 47.30
Maryland 191,188 191,188 30.63
Research Triangle 30,696 283,614 314,310 34.13
New York City 100,787 100,787 97.97
Texas 65,551 26,160 91,711 26.10
Non-cluster/other markets
Subtotal 92,379 255,118 1,536,036 1,883,533 49.28
Key lease expirations with expected downtime 1,484,464 1,484,464 (4) 65.64
Total 92,379 255,118 3,020,500 3,367,997 $56.51
Percentage of expiring leases 3% 8% 89% 100%

Contractual lease expirations for properties classified as held for sale as of March 31, 2026 are excluded from the information on this page.

(1)Amounts in effect as of March 31, 2026.

(2)Excludes month-to-month leases aggregating 103,632 RSF as of March 31, 2026.  Refer to “Leasing activity” in Item 2 for additional details.

(3)Represents 2026 key lease expirations with expected downtime aggregating 747,383 RSF with a weighted-average expiration date of June 2026 and annual rental

revenue aggregating $44.7 million as of March 31, 2026. This includes i) 137,316 RSF generating $13.3 million of annual rental revenues at Alexandria Stanford Life

Science District, where we are evaluating a redevelopment for advanced technology space, ii) 108,136 RSF generating $8.1 million of annual rental revenues at

Alexandria Center® at One Kendall Square, where expected downtime is primarily driven by tenant relocations to other ARE-developed properties, and iii) 99,271 RSF

generating $7.9 million of annual rental revenues at Alexandria Center® for Life Science – Eastlake, where the tenant is also relocating to another ARE-developed

property. Additionally, we have identified prospective tenants or are in early discussions for 111,983 RSF in our Greater Boston and San Francisco Bay Area markets. We

continue to evaluate business plans and re-leasing strategies for these projects to maximize occupancy and rental revenue. We expect weighted-average downtime to

be approximately 6 to 24 months.

(4)Represents space with an annual rental revenue of $97.4 million, with a weighted-average lease expiration date of February 2027, and a weighted-average downtime of

approximately 6 to 24 months. Of the expiring 1.5 million RSF, we have identified prospective tenants or are in early discussions for 532,585 RSF. Compared to the

information previously reported as of December 31, 2025, 2027 key lease expirations increased primarily due to a 232,902-RSF lease expiration with a single tenant at

our Alexandria Center® for Life Science – Waltham Megacampus with $27.0 million of annual rental revenue, for which we no longer expect the tenant to renew. Other

2199023255553

2199023255575

key considerations of this expiring 1.5 million RSF include:

Progress on 2027 Key Lease Expirations
36% 61% 533K<br><br>RSF
of ARR Relocating to<br><br>ARE Development/<br><br>Redevelopment<br><br>Projects Average RSF<br><br>Expansion by<br><br>Relocating<br><br>Tenants Under<br><br>Negotiation<br><br>(36% of<br><br>Expiring RSF)

Located on Megacampuses

(based on RSF)

Laboratory Space

(based on RSF)

64

Top 20 tenants

87% of Top 20 Tenant Annual Rental Revenue Is From Investment-Grade

or Publicly Traded Large Cap Tenants(1)

Our properties are leased to a high-quality and diverse group of tenants, with no individual tenant accounting for greater than

5.8% of our annual rental revenue in effect as of March 31, 2026. The following table sets forth information regarding leases with our 20

largest tenants in North America based upon annual rental revenue in effect as of March 31, 2026 (dollars in thousands, except average

market cap amounts):

Remaining<br><br>Lease<br><br>Term(1)<br><br>(in Years) Aggregate<br><br>RSF Annual<br><br>Rental<br><br>Revenue(1) Percentage<br><br>of Annual<br><br>Rental<br><br>Revenue(1) Investment-Grade<br><br>Credit Ratings Average<br><br>Market<br><br>Cap<br><br>(in billions)
Tenant Moody’s S&P
1 Bristol-Myers Squibb Company 5.9 1,226,762 $ 107,021 5.8% A2 A $102.8
2 Eli Lilly and Company 9.2 1,053,592 92,049 5.0 Aa3 A+ $826.6
3 Moderna, Inc. 12.7 462,100 71,571 3.9 $12.4
4 AstraZeneca PLC 5.9 611,326 55,480 3.0 A1 A+ $255.3
5 Takeda Pharmaceutical Company Limited 10.6 386,111 41,673 2.3 Baa1 BBB+ $48.9
6 Eikon Therapeutics, Inc.(2) 13.2 299,638 38,907 2.1 $0.7
7 Illumina, Inc. 5.6 792,687 29,977 1.6 Baa3 BBB $16.6
8 United States Government 4.4 414,499 29,334 (3) 1.6 Aaa AA+ $—
9 Uber Technologies, Inc. 56.5 (4) 1,009,188 27,865 1.5 Baa1 BBB $178.6
10 Boston Children's Hospital 11.0 309,231 26,294 1.4 Aa2 AA $—
11 Novartis AG 2.2 (5) 321,743 25,111 1.4 Aa3 AA- $273.6
12 Sanofi 4.8 267,278 22,045 1.2 Aa3 AA $119.9
13 Alphabet Inc. 2.1 418,600 21,837 1.2 Aa2 AA+ $2,946.3
14 New York University 6.3 218,983 21,073 1.1 Aa2 AA- $—
15 Massachusetts Institute of Technology 3.8 242,428 20,529 1.1 Aaa AAA $—
16 Charles River Laboratories, Inc. 9.4 238,938 20,045 1.1 $8.1
17 Merck & Co., Inc. 7.8 308,356 19,610 1.1 Aa3 A+ $231.2
18 Vaxcyte, Inc. 8.8 230,755 18,672 1.0 $5.5
19 Altos Labs, Inc.(6) 15.0 158,990 18,407 1.0 $—
20 Amgen Inc. 9.7 317,157 18,181 1.0 Baa1 BBB+ $167.7
Total/weighted-average 9.9 (4) 9,288,362 $ 725,681 39.4%

Annual rental revenue and RSF include 100% of each property managed by us in North America. Refer to “Annual rental revenue” and “Investment-grade or publicly traded large

cap tenants” under “Definitions and reconciliations” in Item 2 for additional details, including our methodologies of calculating annual rental revenue from unconsolidated real

estate joint ventures and average market capitalization, respectively.

(1)Based on total annual rental revenue in effect as of March 31, 2026.

(2)Eikon Therapeutics, Inc. is a public biotechnology company led by Roger Perlmutter, a biopharmaceutical executive who previously served as an executive vice president

of Merck & Co., Inc. As of December 31, 2025, the company held $336 million in cash and marketable securities.

(3)Includes leases, which are not subject to annual appropriations, with governmental entities such as the NIH and the General Services Administration. Approximately 2% of

the annual rental revenue derived from our leases with the United States Government is cancellable prior to the lease expiration date.

(4)Includes (i) ground leases for land at 1455 and 1515 Third Street (two buildings aggregating 422,980 RSF) and (ii) leases at 1655 and 1725 Third Street (two buildings

aggregating 586,208 RSF) in our Mission Bay submarket owned by our unconsolidated real estate joint venture in which we have an ownership interest of 10%. Annual

rental revenue is presented using 100% of the annual rental revenue from our consolidated properties and our share of annual rental revenue from our unconsolidated real

estate joint ventures. Excluding these ground leases, the weighted-average remaining lease term for our top 20 tenants was 8.1 years as of March 31, 2026.

(5)Includes one lease at 100 Technology Square at Alexandria Technology Square® Megacampus in our Cambridge submarket aggregating 255,441 RSF, which generates

annualized rental revenue of $21.0 million and expires in March 2028. We are actively marketing the space for re-lease.

(6)Altos Labs, Inc. is a private biotechnology company led by Hal Barron, M.D., former Chief Scientific Officer and President, R&D at GlaxoSmithKline. Altos Labs launched

with $3.0 billion in private funding in 2022, and is backed by a group of prominent investors.

65

Locations of properties

Our properties are strategically located in AAA life science innovation cluster markets. The following table sets forth the total

RSF, number of properties, and annual rental revenue in effect as of March 31, 2026 in each of our markets in North America (dollars in

thousands, except per RSF amounts):

RSF Number of<br><br>Properties Annual Rental Revenue
Market Operating Development Redevelopment Total % of Total Total % of Total Per RSF
Greater Boston 9,338,588 566,673 1,361,372 11,266,633 29% 63 $697,286 38% $89.09
San Francisco Bay Area 6,083,765 212,657 84,157 6,380,579 16 52 332,008 18 69.11
San Diego 6,225,980 893,525 7,119,505 18 61 295,434 15 53.67
Seattle 2,846,292 227,577 3,073,869 8 39 120,494 7 48.21
Maryland 3,676,566 3,676,566 9 47 153,371 8 45.94
Research Triangle 3,435,988 3,435,988 9 36 89,401 5 27.74
New York City 727,674 727,674 2 2 66,510 4 95.46
Texas 1,651,094 66,350 1,717,444 4 13 37,887 2 28.04
Non-cluster/other markets(1) 417,523 417,523 1 7 10,903 1 30.36
Properties held for sale 1,444,387 1,444,387 4 19 32,567 2 31.27
North America 35,847,857 1,900,432 1,511,879 39,260,168 100% 339 $1,835,861 100% $59.91
3,412,311

Summary of occupancy percentages in North America

The following table sets forth the occupancy percentages for our operating properties and our operating and redevelopment

properties in each of our North America markets, excluding properties held for sale, as of the following dates:

Operating Properties Operating and Redevelopment Properties
Market 3/31/26 12/31/25 3/31/25 3/31/26 12/31/25 3/31/25
Greater Boston 83.8% (2) 86.4% 91.8% 73.1% 75.1% 78.4%
San Francisco Bay Area 87.6 (2) 90.9 90.3 86.4 89.4 86.3
San Diego 88.4 (2) 97.2 94.3 88.4 97.2 94.3
Seattle 87.8 88.4 91.5 87.8 88.4 91.5
Maryland 92.3 (3) 93.6 94.1 92.3 93.6 94.1
Research Triangle 93.8 (4) 95.2 93.4 93.8 95.2 93.4
New York City 95.8 96.4 87.6 95.8 96.4 87.6
Texas 81.8 79.9 82.1 78.7 76.5 78.9
Subtotal 87.8 90.9 91.8 84.0 86.9 87.1
Canada N/A (1) N/A (1) 94.6 N/A N/A 82.4
Non-cluster/other markets 86.0 91.2 73.0 86.0 91.2 73.0
North America 87.7% (5) 90.9% 91.7% 84.1% 86.9% 86.9%

(1)10 properties in Canada were classified as held for sale during the three months ended December 31, 2025 and the one remaining property in Canada is now included

in our non-cluster market.

(2)Decline in occupancy since December 31, 2025 was primarily attributable to previously disclosed key lease expirations with expected downtime, including:

•Greater Boston: 45,636 RSF at Alexandria Center® at Kendall Square in our Cambridge submarket and 52,467 RSF in our Route 128 submarket.

•San Francisco Bay Area: 78,962 RSF at Alexandria Stanford Life Science District in our Greater Stanford submarket, where we are evaluating a redevelopment for

advanced technology use, and 42,299 RSF in our South San Francisco submarket which is currently under negotiation.

•San Diego:

•163,648 RSF at a property in our University Town Center submarket, for which we have executed a letter of intent to re-lease the entire premises;

•118,225 RSF at One Alexandria Square in our Torrey Pines submarket. Of this space, 58,282 RSF has already been re-leased, with occupancy expected to

phase in commencing during the fourth quarter of 2026; and

•83,354 RSF at 5810 and 5820 Nancy Ridge Drive in our Sorrento Mesa submarket which has been re-leased, with occupancy expected to commence during

the fourth quarter of 2026.

(3)Decline in occupancy was primarily driven by the expiration of one private biotechnology lease aggregating 30,161 RSF in our Rockville submarket.

(4)Decline in occupancy was primarily due to 30,667 RSF of vacancy from a tenant winding down operations in the Research Triangle submarket.

(5)Includes temporary vacancies as of March 31, 2026 aggregating 1.1 million RSF, or 3.2% of total operating RSF, primarily in the Greater Boston, San Francisco Bay

Area, and Seattle markets, which are leased and expected to be occupied upon completion of building and/or tenant improvements. The weighted-average expected

delivery date is approximately September 2026, with expected annual rental revenue of approximately $68 million.

66

Investments in real estate

A key component of our business model is our disciplined allocation of capital to the development and redevelopment of new

Class A/A+ properties, and property enhancements identified during the underwriting of certain acquired properties, primarily located in

collaborative Megacampus ecosystems in AAA life science innovation clusters. These projects are focused on providing high-quality,

generic, and reusable spaces that meet the real estate requirements of a wide range of tenants. Upon completion, each development or

redevelopment project is expected to generate increases in rental income, net operating income, and cash flows. Our development and

redevelopment projects are generally in locations that are highly desirable to high-quality entities, which we believe may result in higher

occupancy levels, longer lease terms, higher rental income, higher returns, and greater long-term asset value. Our pre-construction

activities are undertaken in order to prepare the property for its intended use and include entitlements, permitting, design, site work, and

other activities preceding commencement of construction of aboveground building improvements.

Our investments in real estate consisted of the following as of March 31, 2026 (dollars in thousands):

Development and Redevelopment
Under Construction
Operating 2Q26–4Q26<br><br>Stabilization 2027–2028<br><br>Stabilization Evaluating<br><br>Business<br><br>and<br><br>Financial<br><br>Strategy Future Subtotal Total
Square footage
Operating 34,403,470 34,403,470
Future Class A/A+ development and<br><br>redevelopment properties 601,589 1,258,004 1,552,718 20,014,546 23,426,857 23,426,857
Future development and redevelopment square<br><br>feet currently included in rental properties(1) (1,516,872) (1,516,872) (1,516,872)
Total square footage, excluding properties held for<br><br>sale 34,403,470 601,589 1,258,004 1,552,718 18,497,674 21,909,985 56,313,455
Properties held for sale 1,444,387 1,619,425 1,619,425 3,063,812
Total square footage 35,847,857 601,589 1,258,004 1,552,718 20,117,099 23,529,410 59,377,267
Investments in real estate
Gross book value as of March 31, 2026(2) $28,135,300 $629,966 $1,130,851 $1,356,515 $3,971,142 7,088,474 $35,223,774
Properties held for sale 423,335 230,905 230,905 654,240
Total gross investment in real estate, excluding<br><br>properties held for sale $27,711,965 $629,966 $1,130,851 $1,356,515 $3,740,237 6,857,569 $34,569,534

All values are in US Dollars.

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20%

Development/<br><br>Redevelopment<br><br>Under Construction
Land/Future<br><br>Development

17%

11%–16%

Non-Income-Producing Assets(4) as a Percentage of Gross Assets

(1)Refer to “Investments in real estate” under “Definitions and reconciliations” in Item 2 for additional details, including future development and redevelopment square feet

currently included in rental properties.

(2)Balances exclude accumulated depreciation and our share of the cost basis associated with our properties held by our unconsolidated real estate joint ventures, which is

classified as investments in unconsolidated real estate joint ventures in our consolidated balance sheet.

(3)Our share of investment in our development and redevelopment pipeline as of March 31, 2026 is $6.45 billion.

(4)Excludes properties classified as held for sale, of which land parcels represented approximately 1% of total non-income-producing assets as of December 31, 2024 and

2025.

67

Dispositions and sales of partial interests

Our completed dispositions and sales of partial interests of real estate assets during the three months ended March 31, 2026 and pending dispositions as of the date of this report

consisted of the following (dollars in thousands):

Price(Our Share)
Property
Completed in April 2026 2,250
Our share of pending dispositions subject to non-refundable deposits, signed letters of intent, and/or purchase and sale agreement negotiations 149,106
Completed and pending 2026 dispositions as of April 27, 2026 151,356
Dispositions and sales of partial interests identified and in process(1) 2,181,275
Additional projected 567,369
2,900,000
2026 guidance range for dispositions and sales of partial interests
Midpoint 2,900,000
Weighted-average projected disposition date August 2026

All values are in US Dollars.

(1) Consists of assets classified as held for sale as of March 31, 2026, and projected sales of noncontrolling ownership interests in real estate assets for which consolidation is expected to continue.

68

New Class A/A+ development and redevelopment properties

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INCREMENTAL ANNUAL NET OPERATING INCOME

GROWTH EXPECTED FROM ALEXANDRIA’S

DEVELOPMENT AND REDEVELOPMENT DELIVERIES

Near-Term<br><br>Deliveries Intermediate-Term<br><br>Deliveries
2Q26–4Q26 2027–2028
$92M $93M
93%<br><br>Leased/Negotiating 68%<br><br>Leased/Negotiating
601,589 RSF 1.3 million RSF

(1)

(2)

(3)

(4)

For the definition of “Net operating income” and a reconciliation from the most directly comparable GAAP measure, refer to the “Definitions and reconciliations” in Item 2.

(1)Includes expected partial deliveries through 2026 from projects expected to stabilize in 2027-2028, including speculative future leasing that is not yet fully committed. Our share of incremental annual net operating income from

projects expected to be placed into service primarily commencing through 2026 is projected to be $70 million. Refer to the initial and stabilized occupancy years under “New Class A/A+ development and redevelopment properties:

under construction” in Item 2 for additional details.

(2)Our share of incremental annual net operating income from projects expected to stabilize in 2027-2028 is projected to be $60 million.

(3)Represents the current leased/negotiating percentage of development and redevelopment projects that are expected to stabilize through the end of 2026.

(4)Represents the RSF related to projects expected to stabilize in 2026. Does not include RSF for partial deliveries through 2026 from projects expected to stabilize in 2027-2028.

69

New Class A/A+ development and redevelopment properties: recent deliveries

99 Coolidge Avenue 10075 Barnes Canyon Road 8800 Technology Forest Place
Greater Boston/<br><br>Cambridge/Inner Suburbs San Diego/Sorrento Mesa Texas/Greater Houston
146,147 RSF 253,079 RSF 57,042 RSF
100% Occupancy 80% Occupancy 100% Occupancy

The following table presents development and redevelopment of new Class A/A+ projects placed into service during the three months ended March 31, 2026 (dollars in thousands):

Property/Market/Submarket 1Q26<br><br>Delivery<br><br>Date(1) Our<br><br>Ownership<br><br>Interest RSF Placed in Service Occupancy<br><br>Percentage(2) Total Project
Prior to<br><br>1/1/26 1Q26 Total Initial<br><br>Stabilized
RSF Investment
Development projects
99 Coolidge Avenue/Greater Boston/Cambridge/Inner Suburbs 1/27/26 100% 129,413 16,734 146,147 100% 320,809 444,000 6.0% 6.8%
10075 Barnes Canyon Road/San Diego/Sorrento Mesa 3/31/26 50.0% 171,469 81,610 (3) 253,079 80% 253,079 314,000 5.5 5.7
Redevelopment projects
8800 Technology Forest Place/Texas/Greater Houston 2/5/26 100% 50,094 6,948 57,042 100% 123,392 112,000 6.3 6.0
Weighted average/total 1/28/26 350,976 105,292 456,268 697,280 870,000 5.9% 6.3%

All values are in US Dollars.

(1)Represents the average delivery date for deliveries that occurred during the three months ended March 31, 2026, weighted by annual rental revenue.

(2)Occupancy reflects total operating RSF placed in service as of each respective delivery date when the space was placed into service. Subsequent occupancy changes are not reflected.

(3)Includes 50,531 RSF that were vacant and/or unleased at delivery.

70

New Class A/A+ development and redevelopment properties: under construction

99 Coolidge Avenue 50 and 60 Sylvan Road(1) 1450 Owens Street
Greater Boston/<br><br>Cambridge/Inner Suburbs Greater Boston/Route 128 San Francisco Bay Area/<br><br>Mission Bay
174,662 RSF 267,015 RSF 212,657 RSF
84% Leased/Negotiating 74% Leased/Negotiating 51% Leased/Negotiating 269 East Grand Avenue 4135 Campus Point Court 10200 Campus Point Drive 701 Dexter Avenue North
--- --- --- ---
San Francisco Bay Area/<br><br>South San Francisco San Diego/<br><br>University Town Center San Diego/<br><br>University Town Center Seattle/Lake Union
84,157 RSF 426,927 RSF 466,598 RSF 227,577 RSF
40% Leased/Negotiating 100% Leased 100% Leased 23% Leased/Negotiating

(1)Image represents 60 Sylvan Road on the Alexandria Center® for Life Science – Waltham Megacampus. The project is expected to capture demand in our Route 128 submarket.

71

New Class A/A+ development and redevelopment properties: under construction (continued)

The following tables set forth a summary of our new Class A/A+ development and redevelopment properties under construction as of March 31, 2026 (dollars in thousands):

Property/Market/Submarket Located<br><br>on Mega-<br><br>campus Square Footage Percentage Occupancy(1)
Dev/<br><br>Redev In Service CIP Total Leased Leased/<br><br>Negotiating Initial Stabilized
Under construction
2Q26–4Q26 stabilization
99 Coolidge Avenue/Greater Boston/Cambridge/Inner Suburbs X Dev 146,147 174,662 320,809 84% 84% 4Q23 4Q26
4135 Campus Point Court/San Diego/University Town Center X Dev 426,927 426,927 100 100 3Q26 3Q26
146,147 601,589 747,736 93 93
2027–2028 stabilization
50 and 60 Sylvan Road/Greater Boston/Route 128 X Redev 267,015 267,015 74 74 4Q26 2027
1450 Owens Street/San Francisco Bay Area/Mission Bay X Dev 212,657 212,657 51 51 2027 2027
269 East Grand Avenue/San Francisco Bay Area/South San Francisco X Redev 84,157 84,157 40 40 2H26 2027
10200 Campus Point Drive/San Diego/University Town Center(2) X Dev 466,598 466,598 100 100 2028 2028
701 Dexter Avenue North/Seattle/Lake Union X Dev 227,577 227,577 23 23 3Q26 2027
1,258,004 1,258,004 68 68
Total 146,147 1,859,593 2,005,740 77% 77%
Evaluating business and financial strategy with earliest potential lab<br><br>delivery in 2028(3)
311 Arsenal Street/Greater Boston/Cambridge/Inner Suburbs X Redev 56,904 333,758 390,662 9% 28%
421 Park Drive/Greater Boston/Fenway X Dev 392,011 392,011 13 13
40 Sylvan Road/Greater Boston/Route 128 X Redev 329,049 329,049
3000 Minuteman Road/Greater Boston/Other X Redev 431,550 431,550 37
8800 Technology Forest Place/Texas/Greater Houston Redev 57,042 66,350 123,392 46 46
113,946 1,552,718 1,666,664 9% 23%

(1)Initial occupancy dates are subject to leasing and/or market conditions. Stabilized occupancy may vary depending on single tenancy versus multi-tenancy. Multi-tenant projects may increase in occupancy over time.

(2)Represents a single-tenant project that expands the existing Campus Point by Alexandria Megacampus, where we currently have a 57.2% ownership interest. The project is fully leased to a longtime multinational pharmaceutical tenant that

currently occupies one building within the Megacampus aggregating 52,853 RSF, which generated annual rental revenue of $4.1 million as of March 31, 2026. The tenant is expected to vacate this building during 2028. We expect to fund

the majority of future construction costs at the Megacampus until our ownership interest increases to 75%, after which future capital would be contributed pro rata with our joint venture partner.

(3)We are evaluating multiple options, including whether to continue construction of laboratory improvements, pause construction, pursue lower-investment construction alternatives (including a pivot to advanced technology use), or

disposition, based upon future leasing interest. Under a lower-investment, we would expect lower rent and tenant improvement requirements, and we would evaluate whether all or a portion of the property would be placed back into

operation. If the properties are completed as laboratory/office, we do not expect significant deliveries until 2028 at the earliest.

72

New Class A/A+ development and redevelopment properties: under construction (continued)

Our<br><br>Ownership<br><br>Interest At 100% Unlevered Yields
Property/Market/Submarket In Service Initial Stabilized<br><br>(Cash Basis)
Under construction
2Q26–4Q26 stabilization with 93% leased/negotiating
99 Coolidge Avenue/Greater Boston/Cambridge/Inner Suburbs 100% 203,361 185,307 55,332 444,000 6.0% 6.8%
4135 Campus Point Court/San Diego/University Town Center 57.2% 444,659 79,341 524,000 9.8% 6.2%
203,361 629,966
2027–2028 stabilization with 68% leased/negotiating(1)
50 and 60 Sylvan Road/Greater Boston/Route 128 100% 360,504 TBD
1450 Owens Street/San Francisco Bay Area/Mission Bay 25.0% 251,678
269 East Grand Avenue/San Francisco Bay Area/South San Francisco 100% 121,604
10200 Campus Point Drive/San Diego/University Town Center(2) 57.2% 76,310 583,690 660,000 7.3% 6.5%
701 Dexter Avenue North/Seattle/Lake Union 100% 320,755 TBD
1,130,851
Total 203,361 1,760,817 1,020,000 2,990,000
Our share of investment(3)(4) 200,000 1,350,000 680,000 2,230,000
Evaluating business and financial strategy with earliest potential lab<br><br>delivery in 2028(5)
311 Arsenal Street/Greater Boston/Cambridge/Inner Suburbs 100% 28,081 310,025 TBD
421 Park Drive/Greater Boston/Fenway 100% 606,090
40 Sylvan Road/Greater Boston/Route 128 100% 229,568
3000 Minuteman Road/Greater Boston/Other 100% 168,783
8800 Technology Forest Place/Texas/Greater Houston 100% 65,564 42,049
93,645 1,356,515

All values are in US Dollars.

Refer to “Initial stabilized yield (unlevered)” under “Definitions and reconciliations” in Item 2 for additional information.

(1)We expect to provide total estimated costs and related yields for each project over the next several quarters.

(2)Refer to footnote 2 on the prior page for additional details.

(3)Represents dollar amount rounded to the nearest $10 million and includes preliminary estimated amounts for projects listed as TBD.

(4)Represents our share of investment based on our current ownership percentage upon completion of development or redevelopment projects. Our share of investment will be adjusted as our ownership percentage increases at the Campus

Point project.

(5)Refer to footnote 3 on the prior page for additional details.

73

New Class A/A+ development and redevelopment properties: summary of pipeline

77% of Our Total Development and Redevelopment Pipeline RSF

Is Within Our Megacampus Ecosystems

The following table summarizes the key information for all our development and redevelopment projects in North America as of March 31, 2026 (dollars in thousands):

Market<br><br>Property/Submarket Our<br><br>Ownership<br><br>Interest Book Value Development and Redevelopment<br><br>Square Footage
Future Total(1)
Greater Boston
Megacampus: The Arsenal on the Charles/Cambridge/Inner Suburbs 100% 322,671 333,758 34,157 367,915
311 Arsenal Street
Megacampus: 480 Arsenal Way and 446, 458, and 500 Arsenal Street, and 99 Coolidge Avenue/Cambridge/<br><br>Inner Suburbs 100% 209,508 174,662 560,000 734,662
446, 458, and 500 Arsenal Street, and 99 Coolidge Avenue
Megacampus: Alexandria Center® for Life Science – Fenway/Fenway 100% 606,090 392,011 392,011
421 Park Drive
Megacampus: Alexandria Center® for Life Science – Waltham/Route 128 100% 655,867 596,064 515,000 1,111,064
40, 50, and 60 Sylvan Road, and 35 Gatehouse Drive
Megacampus: 30, 200, and 3000 Minuteman Road/Other 100% 194,127 431,550 350,000 781,550
3000 Minuteman Road
Megacampus: Alexandria Technology Square®/Cambridge 100% 8,858 100,000 100,000
10 Necco Street/Seaport Innovation District 100% 107,101 175,000 175,000
215 Presidential Way/Route 128 100% 6,816 112,000 112,000
Other development and redevelopment projects 100% 165,576 740,000 740,000
2,276,614 1,928,045 2,586,157 4,514,202
Refer to “Megacampus” under “Definitions and reconciliations” in Item 2 for additional information.<br><br>(1)Represents total square footage upon completion of development or redevelopment of one or more new Class A/A+ properties. Square footage presented includes the RSF of buildings currently in operation at properties that also have<br><br>future development or redevelopment opportunities. Upon expiration of existing in-place leases, we intend to demolish or redevelop the existing property subject to market conditions and leasing. Refer to “Investments in real estate” under<br><br>“Definitions and reconciliations” in Item 2 for additional information, including development and redevelopment square feet currently included in rental properties.

All values are in US Dollars.

74

New Class A/A+ development and redevelopment properties: summary of pipeline (continued)

Market<br><br>Property/Submarket Our<br><br>Ownership<br><br>Interest Book Value Development and Redevelopment<br><br>Square Footage
Future Total(1)
San Francisco Bay Area
Megacampus: Alexandria Center® for Science and Technology – Mission Bay/Mission Bay 25.0% 251,678 212,657 212,657
1450 Owens Street
Megacampus: Alexandria Center® for Advanced Technologies – South San Francisco/South San Francisco 100% 128,259 84,157 90,000 174,157
211(2) and 269 East Grand Avenue
Megacampus: Alexandria Center® for Advanced Technologies – Tanforan/South San Francisco 100% 443,324 1,930,000 1,930,000
1122, 1150, and 1178 El Camino Real
Alexandria Center® for Life Science – Millbrae/South San Francisco 48.6% 162,361 348,401 348,401
201 and 231 Adrian Road and 30 Rollins Road
Megacampus: Alexandria Center® for Life Science – San Carlos/Greater Stanford 100% 493,425 1,497,830 1,497,830
960 Industrial Road, 987 and 1075 Commercial Street, and 888 Bransten Road
2100, 2200, 2300, and 2400 Geng Road/Greater Stanford 100% 128,360 240,000 240,000
1,607,407 296,814 4,106,231 4,403,045
San Diego
Megacampus: Campus Point by Alexandria/University Town Center 57.2% (3) 709,266 893,525 866,816 1,760,341
10010(4), 10140(4), and 10200 Campus Point Drive and 4135, 4165, 4224, and 4275(4) Campus Point Court
11255 and 11355 North Torrey Pines Road/Torrey Pines 100% 163,973 215,000 215,000
Megacampus: One Alexandria Square/Torrey Pines 100% 69,826 125,280 125,280
10975 and 10995 Torreyana Road
Megacampus: 5200 Illumina Way/University Town Center 51.0% 17,939 451,832 451,832
9625 Towne Centre Drive/University Town Center 30.0% 837 100,000 100,000
Megacampus: Sequence District by Alexandria/Sorrento Mesa 100% 49,696 1,661,915 1,661,915
6290, 6310, 6340, 6350, and 6450 Sequence Drive
Megacampus: SD Tech by Alexandria/Sorrento Mesa 50.0% 127,187 493,845 493,845
9805 Scranton Road and 10065 Barnes Canyon Road
4075 Sorrento Valley Boulevard/Sorrento Valley 100% 29,598 144,000 144,000
Other development and redevelopment projects (2) 78,404 475,000 475,000
1,246,726 893,525 4,533,688 5,427,213
Refer to “Megacampus” under “Definitions and reconciliations” in Item 2 for additional information.<br><br>(1)Represents total square footage upon completion of development or redevelopment of one or more new Class A/A+ properties. Square footage presented includes the RSF of buildings currently in operation at properties that also have<br><br>future development or redevelopment opportunities. Upon expiration of existing in-place leases, we intend to demolish or redevelop the existing property subject to market conditions and leasing. Refer to “Investments in real estate” under<br><br>“Definitions and reconciliations” in Item 2 for additional information, including development and redevelopment square feet currently included in rental properties.<br><br>(2)Includes a property in which we own a partial interest through a real estate joint venture. Refer to Note 4 – “Consolidated and unconsolidated real estate joint ventures” to our unaudited consolidated financial statements in Item 1 for<br><br>additional details.<br><br>(3)The noncontrolling interest share of our real estate joint venture partner is anticipated to decrease to 25%, as we expect to fund the majority of future construction costs at the campus until our ownership interest increases to 75%, after<br><br>which future capital would be contributed pro rata with our partner.<br><br>(4)We have a 100% interest in this property.

All values are in US Dollars.

75

New Class A/A+ development and redevelopment properties: summary of pipeline (continued)

Market<br><br>Property/Submarket Our<br><br>Ownership<br><br>Interest Book Value Development and Redevelopment<br><br>Square Footage
Future Total(1)
Seattle
Megacampus: Alexandria Center® for Advanced Technologies – South Lake Union/Lake Union (2) 617,648 227,577 1,057,400 1,284,977
601 and 701 Dexter Avenue North and 800 Mercer Street
1010 4th Avenue South/SoDo 100% 63,523 544,825 544,825
410 West Harrison Street/Elliott Bay 100% 25,224 91,000 91,000
Megacampus: Alexandria Center® for Advanced Technologies – Canyon Park/Bothell 100% 20,692 230,000 230,000
21660 20th Avenue Southeast
Other development and redevelopment projects 100% 157,385 706,087 706,087
884,472 227,577 2,629,312 2,856,889
Maryland
Megacampus: Alexandria Center® for Life Science – Shady Grove/Rockville 100% 28,803 296,000 296,000
9830 Darnestown Road
28,803 296,000 296,000
Research Triangle
Megacampus: Alexandria Center® for Life Science – Durham/Research Triangle 100% 167,871 2,060,000 2,060,000
Megacampus: Alexandria Center® for Advanced Technologies and AgTech – Research Triangle/Research<br><br>Triangle 100% 115,384 1,170,000 1,170,000
4 and 12 Davis Drive
Megacampus: Alexandria Center® for Sustainable Technologies/Research Triangle 100% 56,960 750,000 750,000
120 TW Alexander Drive, 2752 East NC Highway 54, and 10 South Triangle Drive
Other development and redevelopment projects 100% 1,647 25,000 25,000
341,862 4,005,000 4,005,000
New York City
Megacampus: Alexandria Center® for Life Science – New York City/New York City 100% 180,617 550,000 (3) 550,000
180,617 550,000 550,000
Refer to “Megacampus” under “Definitions and reconciliations” in Item 2 for additional information.<br><br>(1)Represents total square footage upon completion of development or redevelopment of one or more new Class A/A+ properties. Square footage presented includes the RSF of buildings currently in operation at properties that also have<br><br>inherent future development or redevelopment opportunities. Upon expiration of existing in-place leases, we intend to demolish or redevelop the existing property. Refer to “Investments in real estate” under “Definitions and reconciliations”<br><br>for additional information, including development and redevelopment square feet currently included in rental properties.<br><br>(2)We have a 100% interest in 601 and 701 Dexter Avenue North aggregating 415,977 RSF and a 60% interest in the future development project at 800 Mercer Street aggregating 869,000 RSF.<br><br>(3)During the three months ended September 30, 2024, we filed a lawsuit against the New York City Health + Hospitals Corporation and the New York City Economic Development Corporation for fraud and breach of contract concerning our<br><br>option to ground lease a land parcel to develop a future world-class life science building within the Alexandria Center® for Life Science – New York City Megacampus. Refer to “Legal proceedings” in Item 1 under Part II – Other Information<br><br>for additional details.

All values are in US Dollars.

76

New Class A/A+ development and redevelopment properties: summary of pipeline (continued)

Market<br><br>Property/Submarket Our<br><br>Ownership<br><br>Interest Book Value Development and Redevelopment<br><br>Square Footage
Future Total(1)
Texas
Alexandria Center® for Advanced Technologies at The Woodlands/Greater Houston 100% 45,105 66,350 116,405 182,755
8800 Technology Forest Place
1001 Trinity Street and 1020 Red River Street/Austin 100% 137,652 250,010 250,010
Other development and redevelopment projects 100% 60,808 344,000 344,000
243,565 66,350 710,415 776,765
Other development and redevelopment projects 100% 47,503 597,743 597,743
Total pipeline as of March 31, 2026, excluding properties held for sale 6,857,569 3,412,311 20,014,546 23,426,857
Properties held for sale 230,905 1,619,425 1,619,425
Total pipeline as of March 31, 2026 7,088,474 3,412,311 21,633,971 25,046,282

All values are in US Dollars.

Refer to “Megacampus” under “Definitions and reconciliations” in Item 2 for additional information.

(1)Total square footage includes 1,516,872 RSF of buildings currently in operation that we expect to demolish or redevelop and commence future construction subject to market conditions and leasing. Refer to “Investments in real estate”

under “Definitions and reconciliations” in Item 2 for additional information, including development and redevelopment square feet currently included in rental properties.

(2)Includes $3.12 billion of projects that are currently under construction.

77

Results of operations

Same properties

We supplement an evaluation of our results of operations with an evaluation of operating performance of certain of our

properties, referred to as “Same Properties.” For additional information on the determination of our Same Properties portfolio, refer to

“Same property comparisons” under “Definitions and reconciliations” in Item 2. The following table presents information regarding our

Same Properties for the three months ended March 31, 2026:

Three Months Ended<br><br>March 31, 2026
Percentage change in net operating income over comparable period from prior year (11.9)% (1)
Percentage change in net operating income (cash basis) over comparable period from prior year (11.7)% (1)
Operating margin 65%
Number of Same Properties 294
RSF 31,965,127
Occupancy – current-period average 88.9%
Occupancy – same-period prior-year average 94.0%

(1)The quarter-over-quarter decline was due to a decrease in same property occupancy, primarily driven by the previously disclosed 2026 key lease expirations

aggregating 657,492 RSF that became vacant during the three months ended March 31, 2026, with a weighted-average lease expiration date of January 2026, and by

vacancy during the three months ended December 31, 2025 at one property aggregating 170,618 RSF at Alexandria Center® for Advanced Technologies – South San

Francisco in our South San Francisco submarket. We expect our same property performance to improve in the second half of 2026, primarily due to changes in same

property occupancy. This includes the anticipated delivery of 1.1 million RSF of vacant space that was leased but not yet delivered as of March 31, 2026, which has a

weighted-average expected delivery date of approximately September 2026, and is expected to generate annual rental revenue of approximately $68 million.

The following table reconciles the number of Same Properties to total properties for the three months ended March 31, 2026:

Development and redevelopment – under construction Properties
99 Coolidge Avenue 1
1450 Owens Street 1
421 Park Drive 1
4135 Campus Point Court 1
701 Dexter Avenue North 1
10200 Campus Point Drive 1
40, 50, and 60 Sylvan Road 3
269 East Grand Avenue 1
8800 Technology Forest Place 1
311 Arsenal Street 1
3000 Minuteman Road 2
14
Development – placed into service after January 1, 2025
230 Harriet Tubman Way 1
500 North Beacon Street and 4 Kingsbury Avenue 2
10935, 10945, and 10955 Alexandria Way 3
10075 Barnes Canyon Road 1
7
Acquisitions after January 1, 2025
Other 2
2
Unconsolidated real estate JVs 3
Properties held for sale 19
Total properties excluded from Same Properties 45
Same Properties 294
Total properties in North America as of March 31, 2026 339

78

Comparison of results for the three months ended March 31, 2026 to the three months ended March 31, 2025

The following table presents a comparison of the components of net operating income for our Same Properties and Non-Same

Properties for the three months ended March 31, 2026, compared to the three months ended March 31, 2025 (dollars in thousands).

Refer to “Definitions and reconciliations” in Item 2 for definitions of “Tenant recoveries” and “Net operating income” and their

reconciliations from the most directly comparable financial measures presented in accordance with GAAP, income from rentals and net

income, respectively.

Three Months Ended March 31,
2026 2025 $ Change % Change
Income from rentals:
Same Properties $431,442 $473,233 $(41,791) (8.8)%
Non-Same Properties 43,344 78,879 (35,535) (45.1)
Rental revenues 474,786 552,112 (77,326) (14.0)
Same Properties 166,149 165,419 730 0.4
Non-Same Properties 12,078 25,644 (13,566) (52.9)
Tenant recoveries 178,227 191,063 (12,836) (6.7)
Income from rentals 653,013 743,175 (90,162) (12.1)
Same Properties 250 303 (53) (17.5)
Non-Same Properties 17,759 14,680 3,079 21.0
Other income 18,009 14,983 3,026 20.2
Same Properties 597,841 638,955 (41,114) (6.4)
Non-Same Properties 73,181 119,203 (46,022) (38.6)
Total revenues 671,022 758,158 (87,136) (11.5)
Same Properties 208,056 196,587 11,469 5.8
Non-Same Properties 16,086 29,808 (13,722) (46.0)
Rental operations 224,142 226,395 (2,253) (1.0)
Same Properties 389,785 442,368 (52,583) (11.9)
Non-Same Properties 57,095 89,395 (32,300) (36.1)
Net operating income $446,880 $531,763 $(84,883) (16.0)%
Net operating income – Same Properties $389,785 $442,368 $(52,583) (11.9)%
Straight-line rent revenue (13,114) (15,469) 2,355 (15.2)
Amortization of acquired below-market leases and deferred<br><br>revenue related to tenant-funded and -built landlord<br><br>improvements (10,063) (11,558) 1,495 (12.9)
Net operating income – Same Properties (cash basis) $366,608 $415,341 $(48,733) (11.7)%

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Income from rentals

Total income from rentals for the three months ended March 31, 2026 decreased by $90.2 million, or 12.1%, to $653.0 million,

compared to $743.2 million for the three months ended March 31, 2025, due to a decrease in rental revenues, as discussed below.

Rental revenues

Total rental revenues for the three months ended March 31, 2026 decreased by $77.3 million, or 14.0%, to $474.8 million,

compared to $552.1 million for the three months ended March 31, 2025. The decrease was related to dispositions of real estate assets

within our Non-Same Properties since January 1, 2025 and a decrease in Same Properties’ average occupancy, as discussed below.

Same Properties’ rental revenues for the three months ended March 31, 2026 decreased by $41.8 million, or 8.8%, to

$431.4 million, compared to $473.2 million for the three months ended March 31, 2025. This decrease is primarily attributable to a

decrease in Same Properties’ average occupancy to 88.9% for the three months ended March 31, 2026 from 94.0% for the three

months ended March 31, 2025. This decline includes the impact of key lease expirations during the first quarter of 2026, including: i)

45,636 RSF at Alexandria Center® at Kendall Square Megacampus in our Cambridge submarket, ii) 163,648 RSF at 9625 Towne

Center Drive in our University Town Center submarket, and iii) 118,225 RSF at One Alexandria Square Megacampus in our Torrey

Pines submarket. The decrease was further impacted by vacancy at one property aggregating 170,618 RSF at Alexandria Center® for

Advanced Technologies – South San Francisco Megacampus in our South San Francisco submarket during the three months ended

December 31, 2025, which has been re-leased with occupancy expected to commence during the second half of 2026.

Tenant recoveries

Tenant recoveries for the three months ended March 31, 2026 decreased by $12.8 million, or 6.7%, to $178.2 million,

compared to $191.1 million for the three months ended March 31, 2025, primarily in connection with dispositions of real estate assets

within our Non-Same Properties since January 1, 2025.

The decrease in Non-Same Properties tenant recoveries was partially offset by the increase of $730 thousand, or 0.4%, to

$166.1 million in Same Properties’ tenant recoveries for the three months ended March 31, 2026, compared to $165.4 million for the

three months ended March 31, 2025.

Tenant recoveries for our Same Properties increased only modestly during the period despite higher operating costs primarily

due to the decrease in Same Property occupancy to 88.9% as of March 31, 2026, from 94.0% as of March 31, 2025, which reduced the

proportion of expenses recoverable from tenants. As of March 31, 2026, 91% of our leases (on an annual rental revenue basis) were

triple net leases, which require tenants to pay substantially all real estate taxes, insurance, utilities, repairs and maintenance, common

area expenses, and other operating expenses (including increases thereto) in addition to base rent.

Rental operations

Total rental operating expenses for the three months ended March 31, 2026 decreased by $2.3 million, or 1.0%, to

$224.1 million, compared to $226.4 million for the three months ended March 31, 2025. The decrease was primarily due to the

decrease in Non-Same Properties’ rental operating expenses of $13.7 million primarily resulting from the real estate dispositions since

January 1, 2025, partially offset by higher rental operating expenses related to our Same Properties, as discussed below.

Same Properties’ rental operating expenses increased by $11.5 million, or 5.8%, to $208.1 million during the three months

ended March 31, 2026, compared to $196.6 million for the three months ended March 31, 2025, primarily due to: (i) a $4.5 million

increase in utilities expenses, reflecting increases in electricity rates in our Maryland market, our assumption of utilities costs for certain

vacant spaces that were previously paid directly by tenants, primarily in our San Diego market, and increased electric and gas

consumption in our Greater Boston market due to a more severe winter in 2026, (ii) a $3.8 million increase in property taxes, primarily

resulting from a 22% commercial tax rate increase in Cambridge, effective in 2026, as well as changes in leasing arrangements with

certain new tenants in our San Francisco Bay Area market under which we pay property taxes and are reimbursed by tenants,

compared to previous leases where tenants paid taxes directly, and  (iii) a $3.6 million increase in repairs and maintenance expenses,

reflecting more severe winter conditions in our Greater Boston, Maryland, and Research Triangle markets.

Depreciation and amortization

Depreciation and amortization expense for the three months ended March 31, 2026 decreased by $36.6 million, or 10.7%, to

$305.4 million, compared to $342.1 million for the three months ended March 31, 2025. The decrease was primarily as a result of real

estate dispositions of real estate assets since January 1, 2025.

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Impairment of real estate

During the three months ended March 31, 2026, we recognized impairment charges aggregating $5.5 million, classified in

impairment of real estate in our consolidated statement of operations. The impairment primarily reflects an incremental charge

recognized in connection with the amendment of the sales agreement for our Canada portfolio, which was classified as held for sale as

of December 31, 2025. We expect to complete the sale in 2026.

During the three months ended March 31, 2025, we recognized real estate impairment charges aggregating $32.2 million,

related to a ground lease entered into in 2021 for a future development site in our San Francisco Bay Area market.

General and administrative expenses

General and administrative expenses for the three months ended March 31, 2026 increased by $4.0 million, or 13.1%, to

$34.7 million, compared to $30.7 million for the three months ended March 31, 2025. The increase primarily reflects the timing of the

restructuring of compensation plans and other cost-control and efficiency initiatives during the three months ended March 31, 2025.

Notwithstanding this anticipated increase compared to three months ended March 31, 2025, general and administrative

expenses for the three months ended March 31, 2026 decreased by $12.4 million, or 26.3%, compared to $47.1 million for the three

months ended March 31, 2024, reflecting the continued benefit from cost‑efficiency initiatives implemented in prior periods. Based on

the midpoint of our 2026 guidance range, we continue to expect approximately $76 million of cumulative general and administrative

expense savings in 2025 and 2026 compared to 2024.

As a percentage of net operating income, our general and administrative expenses for the trailing twelve months ended

March 31, 2026 and 2025 were 6.0% and 6.9%, respectively.

Interest expense

Interest expense for the three months ended March 31, 2026 and 2025 consisted of the following (dollars in thousands):

Three Months Ended March 31,
Component 2026 2025 Change
Gross interest $134,557 $130,941 $3,616
Capitalized interest (69,973) (80,065) 10,092
Interest expense $64,584 $50,876 $13,708
Average debt balance outstanding(1) $12,901,002 $12,815,953 $85,049
Weighted-average annual interest rate(2) 4.2% 4.1% 0.1%

(1)Represents the average debt balance outstanding during the respective periods.

(2)Represents annualized total interest incurred divided by the average debt balance outstanding during the respective periods.

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The net change in interest expense during the three months ended March 31, 2026, compared to the three months ended

March 31, 2025, resulted from the following (dollars in thousands):

Component Effective Date Change
Increases in interest incurred due to:
Issuances of debt:
550 million of unsecured senior notes payable due 2035 5.66% February 2025 $3,544
750 million of unsecured senior notes payable due 2036 5.41% February 2026 3,956
Higher average outstanding balances under commercial paper program and/or unsecured senior line of credit 13,173
Total increases 20,673
Decreases in interest incurred due to:
Repayments of debt:
Repaid 525 million of 1 billion of unsecured senior notes due 2052 3.64% February 2026 (2,537)
Repaid 498 million of 850 million of unsecured senior notes due 2051 3.09% February 2026 (2,036)
Repaid 309 million of 700 million of unsecured senior notes due 2050 3.91% February 2026 (1,678)
300 million of unsecured senior notes payable due 2026 4.50% January 2026 (2,748)
600 million of unsecured senior notes payable due 2025 3.62% April 2025 (5,219)
Secured notes payable 7.18% August 2025 (2,658)
Other decrease in interest (181)
Total decreases (17,057)
Change in gross interest 3,616
Decrease in capitalized interest 10,092
Total change in interest expense $13,708

All values are in US Dollars.

(1)Represents the weighted-average interest rate as of the end of the applicable period, including amortization of loan fees, amortization of debt premiums (discounts), and

other bank fees.

Investment loss

During the three months ended March 31, 2026, we recognized investment loss aggregating $4.6 million, which consisted of

$18.2 million of realized gains, $10.3 million of unrealized losses, and $12.4 million of impairment charges.

During the three months ended March 31, 2025, we recognized investment loss aggregating $50.0 million, which consisted of

$29.3 million of realized gains, $68.1 million of unrealized losses, and $11.2 million of impairment charges.

For more information about our investments, refer to Note 7 – “Investments” and “Investments” in Note 2 – “Summary of

significant accounting policies” to our unaudited consolidated financial statements in Item 1.

Gain on early extinguishment of debt

During the three months ended March 31, 2026, we recognized a gain on early extinguishment of debt aggregating

$366.4 million, net of the write-off of unamortized debt issuance costs and other transaction-related costs, related to the completion of

the February 2026 tender offers to repurchase $1.33 billion of debt principal across a portion of our outstanding 4.00% Senior Notes

due 2050, 3.00% Senior Notes due 2051, and 3.55% Senior Notes due 2052 for $952.2 million.

Other comprehensive (loss) income

Other comprehensive (loss) income primarily comprised unrealized foreign currency translation gains or losses related to our

operations in Canada. Total other comprehensive loss for the three months ended March 31, 2026 aggregating $1.5 million included

$4.9 million of cumulative translation loss related to our operations in Canada, resulting from the CAD’s weakening against the USD

during this period and partially offset by $3.3 million of unrealized gains related to the change in the fair value of our cross-currency

swap agreements, resulting from the Canadian dollar’s strengthening since the execution of these agreements on July 29, 2025 through

March 31, 2026. Refer to Note 11 – “Hedge Agreements” to our unaudited consolidated financial statements in Item 1 for additional

information.

Total other comprehensive income of $50 thousand for the three months ended March 31, 2025 is primarily due to unrealized

foreign currency translation gains related to our operations in Canada.

82

Summary of capital expenditures

Our construction spending for the three months ended March 31, 2026 and projected spending for the year ending December

31, 2026 consisted of the following (in thousands):

Three Months<br><br>Ended March 31,<br><br>2026 Projected Midpoint for Year Ending December 31, 2026
Construction of Class A/A+ properties:
Active construction projects
Development and redevelopment under construction(1) $ 184,054
Future pipeline pre-construction
Primarily Megacampus expansion pre-construction work (entitlement,<br><br>design, and site work) 56,610 (2)
Revenue- and non-revenue-enhancing capital expenditures(3) 164,382 (4)
Construction spending (before contributions from noncontrolling interests or<br><br>tenants) 405,046
Contributions from noncontrolling interests (consolidated real estate joint<br><br>ventures) (23,804) (5)
Tenant-funded and -built landlord improvements (2,694)
Total construction spending $ 378,548
2026 guidance range for construction spending 1,500,000 – 2,000,000

All values are in US Dollars.

(1)Includes smaller conversions to laboratory space through redevelopment.

(2)Approximately 75% represents capitalized costs.

(3)Represents revenue- and non-revenue-enhancing capital expenditures before contributions from noncontrolling interests and tenant-funded and tenant-built landlord

improvements.

(4)The top two revenue- and non-revenue-enhancing capital expenditure projects in 2026 represent approximately 58% of the total spending within this category. The first

project relates to a property located at the Alexandria Center® for Advanced Technologies – South San Francisco Megacampus in our South San Francisco submarket,

which is leased to a new tenant and is undergoing its first major renovation in 12 years. The second project relates to two properties at the Alexandria Technology

Square® Megacampus in our Cambridge submarket, which are undergoing their first major renovation in 16 years.

(5)Represents contractual capital commitments from existing consolidated real estate joint venture partners to fund construction.

Projected capital contributions from partners in consolidated real estate joint ventures to fund construction

The following table summarizes projected capital contributions from partners in our existing consolidated joint ventures to fund

construction through 2027 and beyond (in thousands):

Projected timing Amount(1)
April 1, 2026 through December 31, 2026 $76,000
2027 and beyond 44,000
Total $120,000

(1)Amounts represent reductions to our consolidated construction spending.

83

Average real estate basis used for capitalization of interest

Our construction spending includes capitalized interest. The table below provides key categories of interest capitalized during

the three months ended March 31, 2026 (in thousands):

Average Real Estate Basis Capitalized
Amount
Construction of Class A/A+ properties:
Development and redevelopment of projects under construction and repositioning<br><br>projects:
2026 stabilization 330,399
2027–2028 stabilization 768,594
Evaluating business and financial strategy(1) 1,277,710
Repositioning and smaller redevelopment projects(2) 1,538,022
3,914,725 57%
Land/future development projects with critical key pre-construction milestones through:
2026(3) 1,159,948
2027(3) 567,606
2028 and beyond(4) 1,217,819
2,945,373 43
Total average real estate basis capitalized(5) 6,860,098 100%

All values are in US Dollars.

(1)Includes projects aggregating 1.6 million RSF for which we are evaluating business and financial strategy and that are expected to reach anticipated construction or

delivery milestones, on a weighted-average real estate investment basis by March 2027. We are evaluating multiple options, including whether to continue construction

of laboratory improvements, pause construction, pursue lower-investment construction alternatives (including a pivot to advanced technology use), or disposition. Upon

achievement of these milestones, if we choose not to pursue future construction or other activities, capitalized interest and other project costs may no longer qualify for

capitalization. Refer to “New Class A/A+ development and redevelopment properties: under construction” in Item 2 for additional information.

(2)Includes the real estate basis related to the 1.1 million RSF of vacant space as of March 31, 2026 that is leased with future delivery. The weighted-average expected

delivery date is approximately September 2026.

(3)Includes future pipeline projects that are expected to reach anticipated pre-construction milestones, including various phases of entitlement, design, site work, and other

activities necessary to begin aboveground vertical construction, on a weighted-average real estate investment basis by August 2026 and April 2027, for the 2026 and

2027 milestones, respectively. At each milestone date, we will evaluate whether to proceed with additional pre-construction and/or construction activities based on

leasing demand and/or market conditions, pause future investments, or consider for potential disposition.

(4)Includes future Megacampus development projects at Alexandria Center® for Advanced Technologies – Tanforan in our South San Francisco submarket and Alexandria

Center® for Life Science – San Carlos in our Greater Stanford submarket, which represents approximately 68% of the total average capitalized real estate basis with

2028 and beyond milestones during the three months ended March 31, 2026. These projects are located at transit-friendly sites with future access to exceptional

amenities.

(5)In addition to capitalized interest, we incur additional capitalized project costs, including property taxes, insurance, payroll, and other costs directly related and essential

to the construction of Class A/A+ properties. If we cease activities necessary to prepare a project for its intended use, costs related to such project are expensed as

incurred. Annualized capitalized operating expenses and payroll represent approximately 2% and 1%, respectively, of the total average real estate basis subject to

capitalization for the three months ended March 31, 2026.

84

Projected results

Our 2026 guidance includes certain forward-looking non-GAAP financial measures, such as funds from operations as adjusted, net debt and preferred stock to Adjusted EBITDA –

fourth quarter of 2026 annualized, fixed-charge coverage ratio – fourth quarter of 2026 annualized, and net cash provided by operating activities, as adjusted, that differ from measures

calculated in accordance with GAAP. These non-GAAP measures are in addition to, and not a substitute for or superior to, financial measures prepared in accordance with GAAP and should

be considered in conjunction with our GAAP financial measures. We are unable to provide corresponding GAAP measures on a forward-looking basis, or a reconciliation from these GAAP

measures to the non-GAAP measures on a forward-looking basis, as we are not able to forecast the net income or loss of future periods without unreasonable effort. This is due to the

inherent difficulty of forecasting the timing and/or amount of items that depend on market conditions outside of our control, including the timing of dispositions, capital events, and financing

decisions, as well as quarterly and annual components such as gain on sales of real estate, impairments of real estate and non-real estate investments, and unrealized gains or losses on

non-real estate investments. Our attempt to predict these amounts may produce significant but inaccurate estimates, which would be potentially misleading for our investors. Refer to

“Definitions and reconciliations” included in Item 2 for additional details about these non-GAAP measures.

Projected 2026 Funds From Operations per Share Attributable to Alexandria’s Common Stockholders –<br><br>Diluted As of 4/27/26 As of 1/26/26
Funds from operations per share and funds from operations per share, as adjusted(1) 6.30 to 6.50 6.25 to 6.55
Midpoint 6.40

All values are in US Dollars.

Key Credit Metric Targets As of 4/27/26 As of 1/26/26 Key Changes
Net debt and preferred stock to Adjusted EBITDA – fourth quarter of 2026 annualized 5.6x to 6.2x 5.6x to 6.2x No Change
Fixed-charge coverage ratio – fourth quarter of 2026 annualized 3.6x to 4.1x 3.6x to 4.1x

Refer to “Definitions and reconciliations” in Item 2 for additional details on key credit metrics.

(1)Refer to “Funds from operations and funds from operations, as adjusted, attributable to Alexandria’s common stockholders” under “Definitions and reconciliations” in Item 2 for additional details.

85

Capital resources

We expect that our principal liquidity needs for the year ending December 31, 2026 will be satisfied by the following multiple sources of capital, as shown in the table below. There

can be no assurance that our sources and uses of capital will not be materially higher or lower than these expectations.

Key Sources and Uses of Capital<br><br>(In millions) As of 4/27/26 As of 1/26/26<br><br>Midpoint
Range Midpoint
Sources of capital:
Reduction in debt $(1,075) $(2,275) (1,675) $(1,675)
Net cash provided by operating activities, as adjusted 475 575 525 525
Dispositions and sales of partial interests(1) 2,100 3,700 2,900 2,900
Total sources of capital $1,500 $2,000 1,750 $1,750
Uses of capital:
Construction $1,500 $2,000 1,750 $1,750
Total uses of capital $1,500 $2,000 1,750 $1,750
Reduction in debt (included above):
Repayment of secured notes payable with 2026 maturities(2) $(650) $(650) (650) (650)
Tender offers for partial principal repayments of unsecured senior notes payable(3) (952) (952) (952)
Issuance of unsecured senior notes payable(3) 750 750 750
Unsecured senior line of credit, commercial paper program, and other (223) (1,423) (823) (1,025)
Reduction in debt $(1,075) $(2,275) (1,675) $(1,675)

All values are in US Dollars.

(1)We expect our dispositions and sales of partial interests for the year ending December 31, 2026 to consist of land, non-core assets, and core assets (primarily sales of partial interests). As of the date of this report, our share of pending

dispositions subject to non-refundable deposits, signed letters of intent, or purchase and sale agreement negotiations aggregated $149.1 million. We have an additional $2.18 billion of dispositions and sales of partial interests identified and

in process.

(2)In January 2026 and April 2026, we repaid, upon maturity, $300.0 million of 4.30% unsecured senior notes payable and $350.0 million of 3.80% unsecured senior notes payable, respectively. These repayments were funded temporarily with

borrowings under our commercial paper program, which will be repaid through planned dispositions and sales of partial interests included in our 2026 guidance. No gain or loss was incurred in connection with these repayments.

(3)In February 2026, we completed tender offers to repurchase debt principal aggregating $1.33 billion across a portion of our outstanding 4.00% Senior Notes due 2050, 3.00% Senior Notes due 2051, and 3.55% Senior Notes due 2052 for

$952.2 million and recognized a gain on early extinguishment of debt of $366.4 million, including the write-off of unamortized debt issuance costs and other transaction-related costs. The tender offers were primarily funded through the

issuance of $750.0 million unsecured senior notes payable, due 2036, with an interest rate of 5.25%. Refer to Note 10 – “Secured and unsecured senior debt” in Item 1 for additional information.

The key assumptions behind the sources and uses of capital in the table above include favorable real estate transaction and capital market environments, performance of our core

operating properties, lease-up and delivery of current and future development and redevelopment projects, and leasing activity. Our expected sources and uses of capital are subject to a

number of variables and uncertainties, including those discussed as “Forward-looking statements” under Part I; “Item 1A. Risk factors”; and “Item 7. Management’s discussion and analysis of

financial condition and results of operations” in our annual report on Form 10-K for the year ended December 31, 2025; as well as in “Item 2. Trends that may affect our future results” within

“Part II – Other information” of this quarterly report on Form 10-Q. We expect to update our forecast for key sources and uses of capital on a quarterly basis.

Refer to “Key assumptions” on the following page.

86

Key Assumptions(1)<br><br>(Dollars in millions) As of 4/27/26 As of 1/26/26 Key Changes<br><br>to Midpoint
Low High Low High
Occupancy of operating properties as of December 31, 2026 86.2% 87.8% 87.7% 89.3% 150 bps reduction(3)
Same property performance:
Net operating income (10.5)% (8.5)% (9.5)% (7.5)% 100 bps reduction(3)
Net operating income (cash basis) (10.5)% (8.5)% (9.5)% (7.5)% 100 bps reduction(3)
Lease renewals and re-leasing of space:
Rental rate changes (9.0)% (1.0)% (2.0)% 6.0% 700 bps reduction(4)
Rental rate changes (cash basis) (15.0)% (7.0)% (12.0)% (4.0)% 300 bps reduction(4)
Straight-line rent revenue 55 85 $65 $95 $10 million reduction(4)
General and administrative expenses 134 154 $134 $154 No Change
Capitalization of interest 225 265 $225 $275 $5 million reduction(5)
Interest expense 240 280 $230 $280 $5 million increase(5)
Realized gains on non-real estate investments(6) 60 90 $60 $90 No Change

All values are in US Dollars.

(1)Our assumptions presented in the table above are subject to a number of variables and uncertainties, including those discussed as “Forward-looking statements” under Part I; “Item 1A. Risk factors”; and “Item 7. Management’s discussion

and analysis of financial condition and results of operations” in our annual report on Form 10-K for the year ended December 31, 2025, as well as in “Item 2. Trends that may affect our future results” within “Part II – Other information” of this

quarterly report on Form 10-Q. To the extent our full-year earnings guidance is updated during the year, we will provide additional disclosure supporting reasons for any significant changes to such guidance.

(2)Our guidance for operating occupancy percentage as of December 31, 2026 and for same property performance net operating income changes assumes an approximate 1% and 2% benefit, respectively, related to a range of assets with

vacancy that could potentially be sold during 2026 and/or qualify for designation as held for sale by December 31, 2026, but that had not yet met such criteria as of March 31, 2026. Refer to the chart below for key drivers of the changes to

certain key projected operating metrics and assumptions included in our 2026 guidance.

(3)Decline primarily relates to changes to a range of properties that could potentially be sold during 2026 that were assumed in our prior guidance. Refer to the chart below.

(4)Decline primarily related to the re-lease of two spaces subject to tenant wind-downs. Refer to the chart below.

(5)The $5 million reduction and corresponding $5 million increase to the midpoints of our guidance ranges for 2026 capitalized interest and 2026 interest expense, respectively, are primarily driven by the anticipated earlier completion of

certain milestones on several projects. Refer to the discussion of fourth quarter of 2026 FFO per share – diluted, as adjusted and “Capitalized interest” item on the following page, and “Average real estate basis used for capitalization of

interest” in Item 2 for additional details.

(6)Represents realized gains and losses included in funds from operations per share – diluted, as adjusted. Excludes unrealized gains and losses and significant gains and impairments realized on non-real estate investments, if any. Refer to

Note 7 – “Investments” to our unaudited consolidated financial statements in Item 1 for additional details.

Occupancy of Operating Properties Same Property Performance Lease Renewals and<br><br>Re-leasing of Space Straight-Line Rent Revenue
Key Drivers of Changes to Certain 2026 Projected<br><br>Operating Metrics and Assumptions As of<br><br>December 31, 2026 Benefit From<br><br>Potential Held<br><br>For Sale<br><br>Assets(1) Net Operating<br><br>Income Changes<br><br>(Cash Basis) Net Operating<br><br>Income Changes Benefit From<br><br>Potential<br><br>Held For<br><br>Sale Assets(1) Rental Rate<br><br>Changes Rental Rate<br><br>Changes<br><br>(Cash Basis)
Guidance ranges as of 1/26/26 87.7% to 89.3% 2% (9.5)% to (7.5)% (9.5)% to (7.5)% 3% (2.0)% to 6.0% (12.0)% to (4.0)% 65M to 95M
Changes to range of properties that could potentially be sold<br><br>during 2026 that were assumed in prior 2026 guidance(1) (1.3) (1) (1.0) (1.0) (1)
Primarily related to the re-lease of two spaces subject to<br><br>tenant wind-downs(2) (0.2) (7.0) (3.0) (10)
Total changes to 2026 guidance midpoints (1.5) (1) (1.0) (1.0) (1) (7.0) (3.0) (10)
Guidance ranges as of 4/27/26 86.2% to 87.8% 1% (10.5)% to (8.5)% (10.5)% to (8.5)% 2% (9.0)% to (1.0)% (15.0)% to (7.0)% 55M to 85M

All values are in US Dollars.

(1)Our prior guidance for occupancy percentage as of December 31, 2026 and 2026 same property performance net operating income changes assumed a benefit of approximately 2% and 3%, respectively, related to a range of assets with

vacancy that could potentially be sold in 2026 and/or qualify for classification as held for sale by December 31, 2026 but had not yet met such criteria as of December 31, 2025. Our updated guidance for these metrics assumes a reduced

benefit of approximately 1% and 2%, respectively, related to a range of assets with vacancy that could potentially be sold during 2026 and/or qualify for classification as held for sale by December 31, 2026, primarily due to our revised

expectation that we may no longer sell as many assets with significant vacancy driven, in part, by positive leasing prospects for certain spaces that are currently vacant.

(2)Includes the impacts of i) one lease aggregating 81,220 RSF in our Torrey Pines submarket, for which we proactively addressed a tenant wind-down by terminating the existing lease and executing a new lease in April 2026 with a growth-

stage life science company advancing next-generation therapeutics to accommodate their expansion needs within our portfolio, with expected delivery in early 2027 following the completion of tenant improvements, and ii) one lease

aggregating 47,719 RSF at 480 Arsenal Street in our Cambridge/Inner Suburbs submarket, with the space re-leased during the three months ended March 31, 2026. Refer to "Leasing activity" in Item 2 for additional details.

(3)Primarily attributable to the write-off of a deferred rent receivable in April 2026 of approximately $5 million in connection with the lease termination and a payment of $10.5 million from a tenant in our Torrey Pines submarket discussed in

footnote 2 above.

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Fourth quarter of 2026 FFO per share – diluted, as adjusted

•On December 3, 2025, we provided guidance for our projected fourth quarter of 2026 FFO per share – diluted, as adjusted, to be within a range of $1.40 to $1.60 as part of

our path forward. The decline in our quarterly FFO from first quarter of 2026 to fourth quarter of 2026 is due, in part, to projected dispositions and partial interest sales of

land, non-core, and core properties aggregating $2.9 billion at our guidance midpoint, currently with a weighted-average projected completion date in August 2026.

•As of April 27, 2026, we continue to refine this guidance and currently expect to be within a range of $1.40 to $1.50. The updated range reflects an assumption for lower

capitalized interest, and a corresponding increase in interest expense, in the fourth quarter of 2026 due to earlier than anticipated completion of construction/pre-

construction milestones on several projects (refer to the “Capitalized interest” item below for additional details).

549755813889

$1.40 – $1.50

1)Potential tenant wind-downs

•Our 2026 guidance includes a $25 million to $30 million reduction in funds from operations for potential tenant wind-downs, of which approximately $6 million was

recognized during the three months ended March 31, 2026.

•Based upon current market conditions, we estimate at least a similar level of reduction in funds from operations may be necessary through 2026 and beyond.

2)Development-related other income

•During the three months ended March 31, 2026, we recognized development fees and other related revenues of approximately $2.5 million, most of which are expected to

cease by the end of 2026 as we complete the respective projects.

3)Development and redevelopment projects under business and financial strategy evaluation

•We have five development and redevelopment projects for which the business and financial strategies continue to be evaluated, including whether to continue construction

of laboratory improvements, pause construction, pursue lower-investment construction alternatives (including a pivot to advanced technology use), or disposition. Refer to

“New Class A/A+ development and redevelopment properties: under construction” in Item 2 for additional details.

◦If we elect to continue to pursue construction of laboratory improvements for these projects, the earliest deliveries of these projects are in 2028.

◦If we elect to pursue lower-investment construction alternatives (including a pivot to advanced technology use), these projects could deliver earlier than 2028. The

incremental capital required for alternative use construction, and corresponding rental rates earned, is generally lower compared to laboratory improvements.

4)Capitalized interest

•For the three months ended March 31, 2026, average real estate basis capitalized of $6.86 billion comprised the following:

•$2.64 billion of active development and redevelopment of projects under construction that are expected to stabilize through 2028 and are 77% leased and repositioning

projects;

•$1.28 billion of development and redevelopment projects for which we are evaluating the business and financial strategy with weighted-average critical key construction

milestones by March 2027;

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•$1.16 billion of land with weighted-average critical key pre-construction milestones by August 2026;

•$567.6 million of land with weighted-average critical key pre-construction milestones by April 2027; and

•$1.22 billion of land with critical key pre-construction milestones through 2028 and beyond.

•At each milestone date, we evaluate, on an asset-by-asset basis, whether to (i) proceed with additional pre-construction and/or construction activities based on leasing

demand and/or market conditions, (ii) pause future investments, or (iii) consider the potential dispositions of these real estate assets. If we cease activities necessary to

prepare a project for its intended use, costs related to such project, including interest, payroll, property taxes, insurance, and other costs directly related and essential to the

construction of Class A/A+ properties, are expensed as incurred. Annualized capitalized operating expenses and payroll represent approximately 2% and 1%, respectively,

of the total average real estate basis subject to capitalization for the three months ended March 31, 2026.

•We expect average real estate basis capitalized to decrease from $6.86 billion for the three months ended March 31, 2026 to a range from $3.8 billion to $5.3 billion for the

fourth quarter of 2026, driven by potential dispositions, deliveries of development and redevelopment, and pauses in construction/pre-construction activities. The estimated

range for the fourth quarter of 2026 represents a $200 million reduction (at the midpoint) in the projected range from $4.0 billion to $5.5 billion initially disclosed on

December 3, 2025. Refer to “Average real estate basis used for capitalization of interest” in Item 2 for additional details.

5)Key lease expirations

•We estimate 1.5 million RSF of leases expiring in 2027, with approximately $97.4 million of annual rental revenue, to have 6 to 24 months of downtime on a weighted-

average basis. These expirations have a weighted-average contractual lease expiration date of February 2027. 2027 expirations increased from the prior quarter primarily

due to one expiration of a 232,902 RSF single-tenant lease at our Alexandria Center® for Life Science – Waltham Megacampus with approximately $27.0 million of annual

rental revenue, for which we no longer expect the tenant to renew. While our initial acquisition underwriting assumed this tenant would eventually relocate to another

submarket, the relocation by the tenant is expected to occur earlier than previously anticipated. Refer to “Summary of contractual lease expirations” in the Item 2 for

additional details.

6)Construction spending

•We are currently evaluating our future construction spending estimates beyond 2026, and a number of factors could cause our preliminary estimates for the period beyond

2026 to change as we refine our estimates over the course of the year. We estimate our annual construction spending beyond 2026 could decline by approximately $500

million, subject to market conditions, and is expected to primarily focus on, i) construction spending required to complete our development and redevelopment projects that

are expected to stabilize through 2028 and are 77% leased, and ii) revenue- and non-revenue-enhancing capital expenditures, in order to secure leasing of vacant space

and renewals and re-leasing of space at our operating properties.

We expect to introduce 2027 guidance and related assumptions at our Investor Day on December 2, 2026, consistent with our historical practice.

89

Consolidated and unconsolidated real estate joint ventures

We present components of balance sheet and operating results information for the noncontrolling interest share of our

consolidated real estate joint ventures and for our share of investments in unconsolidated real estate joint ventures to help investors

estimate balance sheet and operating results information related to our partially owned entities. These amounts are estimated by

computing, for each joint venture that we consolidate in our financial statements, the noncontrolling interest percentage of each financial

item to arrive at the cumulative noncontrolling interest share of each component presented. In addition, for our real estate joint ventures

that we do not control and do not consolidate, we apply our economic ownership percentage to the unconsolidated real estate joint

ventures to arrive at our proportionate share of each component presented. Refer to Note 4 – “Consolidated and unconsolidated real

estate joint ventures” to our unaudited consolidated financial statements in Item 1 for further discussion.

Consolidated Real Estate Joint Ventures
Property/Market/Submarket Noncontrolling<br><br>Interest Share Operating RSF<br><br>at 100%
50 and 60 Binney Street/Greater Boston/Cambridge/Inner Suburbs 66.0% 532,395
75/125 Binney Street/Greater Boston/Cambridge/Inner Suburbs 60.0% 388,270
100 and 225 Binney Street and 300 Third Street/Greater Boston/Cambridge/Inner Suburbs 70.0% 870,641
15 Necco Street/Greater Boston/Seaport Innovation District 43.3% 345,996
Alexandria Center® for Science and Technology – Mission Bay/San Francisco Bay Area/<br><br>Mission Bay(1) 75.0% 550,851
211 and 213 East Grand Avenue/San Francisco Bay Area/South San Francisco 70.0% 300,930
500 Forbes Boulevard/San Francisco Bay Area/South San Francisco 90.0% 155,685
Alexandria Center® for Life Science – Millbrae/San Francisco Bay Area/South San Francisco 51.4% 285,346
3215 Merryfield Row/San Diego/Torrey Pines 70.0% 170,523
Campus Point by Alexandria/San Diego/University Town Center(2)(3) 42.8% (4) 1,159,770
5200 Illumina Way/San Diego/University Town Center 49.0% 792,687
9625 Towne Centre Drive/San Diego/University Town Center 70.0% 163,648
SD Tech by Alexandria/San Diego/Sorrento Mesa(2)(5) 50.0% 1,051,752
Summers Ridge Science Park/San Diego/Sorrento Mesa(6) 70.0% 316,531
1201 and 1208 Eastlake Avenue East/Seattle/Lake Union 70.0% 206,134
400 Dexter Avenue North/Seattle/Lake Union 70.0% 290,754
800 Mercer Street/Seattle/Lake Union 40.0% (2)
Unconsolidated Real Estate Joint Ventures
Property/Market/Submarket Our Ownership<br><br>Share Operating RSF<br><br>at 100%
1655 and 1725 Third Street/San Francisco Bay Area/Mission Bay 10.0% 586,208
101 West Dickman Street/Maryland/Beltsville 58.4% (7) 135,958

Refer to “Joint venture financial information” under “Definitions and reconciliations” in Item 2 for additional

(1)Includes 1450, 1500, and 1700 Owens Street and 455 Mission Bay Boulevard South.

(2)Includes properties currently under construction or in our future development and redevelopment pipeline. Refer to “New Class A/A+ development and redevelopment

properties” in Item 2 for additional details.

(3)Includes 10200, 10290, and 10300 Campus Point Drive and 4135, 4155, 4165, 4224, and 4242 Campus Point Court.

(4)The noncontrolling interest share of our real estate joint venture partner is anticipated to decrease to 25%, as we expect to fund the majority of future construction costs

at the campus until our ownership interest increases to 75%, after which future capital would be contributed pro rata with our partner. Refer to “New Class A/A+

development and redevelopment properties: under construction” in Item 2 for additional details.

(5)Includes 9605, 9645, 9675, 9725, 9735, 9805, 9808, 9855, and 9868 Scranton Road and 10055, 10065, and 10075 Barnes Canyon Road.

(6)Includes 9965, 9975, 9985, and 9995 Summers Ridge Road.

(7)Represents a joint venture with a local real estate operator in which our joint venture partner manages the day-to-day activities that significantly affect the economic

performance of the joint venture.

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The following table presents key terms related to our unconsolidated real estate joint ventures’ secured loans as of March 31,

2026 (dollars in thousands):

Maturity Date Stated Rate Interest<br><br>Rate(1) At 100%
Unconsolidated Joint Venture Aggregate<br><br>Commitment Debt Balance(2)
101 West Dickman Street 10/29/26 SOFR+1.95% (3) 5.68% $26,750 19,048
1655 and 1725 Third Street 2/10/35 6.37% 6.44% 500,000 496,967
$526,750 516,015

All values are in US Dollars.

(1)Includes interest expense and amortization of loan fees.

(2)Represents outstanding principal, net of unamortized deferred financing costs, as of March 31, 2026.

(3)This loan is subject to a fixed SOFR floor of 0.75%.

The following tables present information related to the operating results and financial positions of our consolidated and

unconsolidated real estate joint ventures as of and for the three months ended March 31, 2026 (in thousands):

Three Months Ended March 31, 2026
Noncontrolling Interest<br><br>Share of Consolidated Real<br><br>Estate Joint Ventures Our Share of<br><br>Unconsolidated<br><br>Real Estate Joint Ventures
Total revenues $97,212 $3,006
Rental operations (30,677) (1,191)
66,535 1,815
General and administrative (622) (22)
Interest (63) (1,026)
Depreciation and amortization of real estate assets (29,473) (914)
Fixed returns allocated to redeemable noncontrolling interest(1) 347
$36,724 $(147)
Straight-line rent and below-market lease revenue $2,981 $197
Funds from operations(2) $66,197 $767

Refer to “Joint venture financial information” under “Definitions and reconciliations” in Item 2 for additional details.

(1)Represents an allocation of joint venture earnings to redeemable noncontrolling interest for a property in the San Francisco Bay Area market. This redeemable

noncontrolling interest earns a fixed return on their investment and does not participate in the operating results of the property.

(2)Refer to “Funds from operations and funds from operations, as adjusted, attributable to Alexandria Real Estate Equities, Inc.’s common stockholders” under “Definitions

and reconciliations” in Item 2 for the definition and its reconciliation from the most directly comparable financial measure presented in accordance with GAAP.

As of March 31, 2026
Noncontrolling Interest<br><br>Share of Consolidated<br><br>Real Estate Joint Ventures Our Share of<br><br>Unconsolidated<br><br>Real Estate Joint Ventures
Investments in real estate $3,261,264 $88,793
Cash, cash equivalents, and restricted cash 111,509 1,648
Other assets 411,563 10,686
Secured notes payable (60,821)
Other liabilities (154,688) (9,786)
Redeemable noncontrolling interests (9,234)
$3,620,414 $30,520

During the three months ended March 31, 2026 and 2025, our consolidated real estate joint ventures distributed an aggregate

of $60.1 million and $66.0 million, respectively, to our joint venture partners. Refer to our consolidated statements of cash flows and

Note 4 – “Consolidated and unconsolidated real estate joint ventures” to our unaudited consolidated financial statements in Item 1 for

additional information.

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Investments

We hold investments in publicly traded companies and privately held entities primarily involved in the life science industry. The

tables below summarize components of our investment income (loss) and non-real estate investments (in thousands). Refer to Note 7 –

“Investments” to our unaudited consolidated financial statements in Item 1 for additional information.

Three Months Ended
March 31, 2026 March 31, 2025
Realized gains:
Realized gains 18,198 29,333
Impairment of non-real estate investments (12,448) (11,180)
5,750 18,153
Unrealized losses (10,332) (68,145)
Investment loss (4,582) (49,992)

All values are in US Dollars.

March 31, 2026
Investments Cost Unrealized<br><br>Gains Unrealized<br><br>Losses Carrying Amount
Publicly traded companies 83,916 $34,674 $(16,514) 102,076
Entities that report NAV 471,058 102,050 (38,132) 534,976
Entities that do not report NAV:
Entities with observable price changes 82,128 54,780 (10,991) 125,917
Entities without observable price changes 405,567 405,567
Investments accounted for under the equity method N/A N/A N/A 367,883
March 31, 2026 1,042,669 $191,504 $(65,637) 1,536,419
December 31, 2025 1,010,488 $184,434 $(51,056) 1,501,249

All values are in US Dollars.

Public/Private Mix (Cost) Tenant/Non-Tenant Mix (Cost)

1

13

6%

Public

17%

Tenant

94%

Private

83%

Non-Tenant

(1)Primarily related to two non-real estate investments in privately held entities that do not report NAV.

(2)Primarily relates to the accounting reclassifications of unrealized gains recognized in prior periods into realized gains upon our realization of investments during the three

months ended March 31, 2026.

(3)Primarily relates to the decrease in fair values of our investments in publicly traded entities and privately held entities that report NAV during the three months ended

March 31, 2025.

(4)Represents 2.6% of gross assets as of March 31, 2026. Refer to “Gross assets” under “Definitions and reconciliations” in Item 2 for additional details.

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Liquidity

Liquidity
4.2B
(In millions)
Availability under our unsecured senior line of credit, net of amounts outstanding under our commercial paper program
Cash, cash equivalents, and restricted cash
Investments in publicly traded companies
Liquidity as of March 31, 2026

All values are in US Dollars.

We expect to meet certain long-term liquidity requirements, such as requirements for development, redevelopment, other

construction projects, capital improvements, tenant improvements, property acquisitions, equity repurchases, leasing costs, revenue-

and non-revenue-enhancing capital expenditures, scheduled debt maturities, distributions to noncontrolling interests, and payment of

dividends through net cash provided by operating activities, as adjusted, periodic asset dispositions, strategic real estate joint ventures,

long-term secured and unsecured indebtedness, borrowings under our unsecured senior line of credit, issuances under our commercial

paper program, and issuances of additional debt and/or equity securities.

We also expect to continue meeting our short-term liquidity and capital requirements, as further detailed in this section,

generally through our working capital and net cash provided by operating activities, as adjusted. We believe that the net cash provided

by operating activities, as adjusted, will continue to be sufficient to enable us to make the distributions necessary to continue qualifying

as a REIT.

For additional information on our liquidity requirements related to our contractual obligations and commitments, refer to

Note 5 – “Leases” and Note 10 – “Secured and unsecured senior debt” to our unaudited consolidated financial statements in Item 1.

Over the next several years, our balance sheet, capital structure, and liquidity objectives are as follows:

•Retain net cash provided by operating activities, as adjusted, for investment in development and redevelopment projects

and/or acquisitions;

•Maintain significant balance sheet liquidity;

•Maintain a strong credit profile and relative long-term cost of capital;

•Maintain diverse sources of capital, including sources from net cash provided by operating activities, as adjusted,

unsecured debt, secured debt, selective real estate asset sales, strategic real estate joint ventures, non-real estate

investment sales, and common stock;

•Maintain commitment to long-term capital to fund growth;

•Maintain prudent laddering of debt maturities;

•Maintain solid credit metrics;

•Prudently manage variable-rate debt exposure;

•Maintain a large, unencumbered asset pool to provide financial flexibility;

•Fund common stock dividends and distributions to noncontrolling interests from net cash provided by operating activities,

as adjusted;

•Manage a disciplined level of development and redevelopment projects as a percentage of our gross real estate assets;

and

•Maintain high levels of pre-leasing and percentage leased in development and redevelopment projects.

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The following table presents the availability under our unsecured senior line of credit, net of amounts outstanding under our

commercial paper program; cash, cash equivalents, and restricted cash; and investments in publicly traded companies as of March 31,

2026 (in thousands):

Description Stated Rate Aggregate<br><br>Commitments Outstanding<br><br>Balance Remaining<br><br>Commitments/<br><br>Liquidity
Availability under our unsecured senior line of credit, net of<br><br>amounts outstanding under our commercial paper program SOFR+0.835% $5,000,000 $1,355,271 $3,644,729
Cash, cash equivalents, and restricted cash 423,385
Investments in publicly traded companies 102,076
Liquidity as of March 31, 2026 $4,170,190

Cash, cash equivalents, and restricted cash

As of March 31, 2026 and December 31, 2025, we had $423.4 million and $553.8 million, respectively, of cash, cash

equivalents, and restricted cash. We expect existing cash, cash equivalents, and restricted cash, net cash provided by operating

activities, as adjusted, proceeds from real estate asset sales, sales of partial interests, strategic real estate joint ventures, non-real

estate investment sales, borrowings under our unsecured senior line of credit, issuances under our commercial paper program,

issuances of unsecured senior notes payable, and issuances of common stock to continue to be sufficient to fund our operating

activities and cash commitments for investing and financing activities, such as regular quarterly dividends, distributions to noncontrolling

interests, scheduled debt repayments, acquisitions, and certain capital expenditures, including expenditures related to construction

activities and any common stock repurchases.

Cash flows

We report and analyze our cash flows based on operating activities, investing activities, and financing activities. The following

table summarizes changes in our cash flows for the three months ended March 31, 2026 and 2025 (in thousands):

Three Months Ended March 31,
2026 2025 Change
Net cash provided by operating activities $196,624 $207,949 $(11,325)
Net cash used in investing activities $(584,810) $(654,779) $69,969
Net cash provided by financing activities $258,003 $370,775 $(112,772)

Operating activities

Cash flows provided by operating activities are primarily dependent upon the occupancy level of our asset base, the rental

rates of our leases, the collectibility of rent and recovery of operating expenses from our tenants, the timing of completion of

development and redevelopment projects, and the timing of acquisitions and dispositions of operating properties. Net cash provided by

operating activities for the three months ended March 31, 2026 decreased by $11.3 million, or 5.4%, to $196.6 million, compared to

$207.9 million for the three months ended March 31, 2025, primarily reflecting the impact of real estate dispositions and sales of partial

interests completed since January 1, 2025.

94

Investing activities

Cash used in investing activities for the three months ended March 31, 2026 and 2025 consisted of the following (in

thousands):

Three Months Ended March 31, Change
2026 2025
Sources of cash from investing activities:
Proceeds from sales of real estate $— $68,182 $(68,182)
Sales of and distributions from non-real estate investments 35,000 12,691 22,309
Return of capital from unconsolidated real estate joint ventures 113 113
35,113 80,873 (45,760)
Uses of cash for investing activities:
Additions to real estate 545,999 645,841 (99,842)
Change in escrow deposits 9,506 (9,506)
Investments in unconsolidated real estate joint ventures 297 10,994 (10,697)
Additions to non-real estate investments 73,627 69,311 4,316
619,923 735,652 (115,729)
Net cash used in investing activities $584,810 $654,779 $(69,969)

The change in net cash used in investing activities for the three months ended March 31, 2026, compared to the three months

ended March 31, 2025, was primarily due to a decreased use of cash for additions to real estate and an increased source of cash from

sales of and distributions from non-real estate investments. Refer to Note 3 – “Investments in real estate” to our unaudited consolidated

financial statements in Item 1 for additional information.

Financing activities

Cash flows provided by financing activities for the three months ended March 31, 2026 and 2025 consisted of the following

(in thousands):

Three Months Ended March 31,
2026 2025 Change
Borrowings under secured notes payable $— $824 $(824)
Repayments of borrowings under secured notes payable (8,892) (8,892)
Proceeds from issuance of unsecured senior notes payable 747,592 548,532 199,060
Repayments of unsecured senior note payable (1,252,203) (1,252,203)
Proceeds from issuances under commercial paper program 12,319,811 2,700,000 9,619,811
Repayments of borrowings under commercial paper program (11,318,040) (2,400,000) (8,918,040)
Payments of loan fees (8,814) (5,406) (3,408)
Changes related to debt 479,454 843,950 (364,496)
Contributions from and sales of noncontrolling interests 18,065 54,409 (36,344)
Distributions to noncontrolling interests (60,111) (66,034) 5,923
Purchases and redemptions of noncontrolling interests (49,707) (17,818) (31,889)
Repurchase of common stock (208,187) 208,187
Dividends on common stock (123,752) (229,987) 106,235
Taxes paid related to net settlement of equity awards (5,946) (5,558) (388)
Net cash provided by financing activities $258,003 $370,775 $(112,772)

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Sources of capital

Net cash provided by operating activities, as adjusted

We expect to retain $475 million to $575 million of net cash provided by operating activities, as adjusted, for the year ending

December 31, 2026. Refer to “Net cash provided by operating activities, as adjusted” under “Definitions and reconciliations” in Item 2 for

the definition and reconciliation from the most directly comparable financial measure presented in accordance with GAAP. For the year

ending December 31, 2026, we expect our recently delivered projects, our development and redevelopment projects expected to be

delivered, and contributions from Same Properties to contribute to income from rentals, net operating income, and cash flows. We

anticipate contractual near-term growth in annual net operating income (cash basis) of $25 million related to the commencement of

contractual rents on the projects recently placed into service that are near the end of their initial free rent period. Refer to “Cash flows”

in Item 2 for a discussion of cash flows provided by operating activities for the three months ended March 31, 2026.

Debt

We expect to fund a portion of our capital needs for 2026 and beyond from issuances under our commercial paper program,

issuances of unsecured senior notes payable, and/or borrowings under our unsecured senior line of credit, and/or borrowings under

secured construction loans.

As of March 31, 2026, our unsecured senior line of credit, which matures in 2030, including extension options under our

control, had aggregate commitments of $5.0 billion and bore an interest rate of SOFR plus 0.835%. In addition to the cost of borrowing,

the unsecured senior line of credit is subject to an annual facility fee of 0.14% based on the aggregate commitments outstanding.

Based upon our ability to achieve certain annual sustainability targets, the interest rate and facility fee rate are also subject to upward or

downward adjustments of up to four basis points with respect to the interest rate and up to one basis point with respect to the facility fee

rate.

During the three months ended March 31, 2026, we achieved certain annual sustainability targets, as described in our

unsecured senior line of credit agreement, which reduced the borrowing rate by four basis points for a one-year period to SOFR plus

0.835%, from SOFR plus 0.875%, and reduced the facility fee by one basis point to 0.14% from 0.15%. As of March 31, 2026, we had

no outstanding balance on our unsecured senior line of credit.

Our commercial paper program provides us with the ability to issue up to $2.50 billion of commercial paper notes with a

maturity of generally 30 days or less and with a maximum maturity of 397 days from the date of issuance. Our commercial paper

program is back-stopped by our unsecured senior line of credit, and at all times we expect to retain a minimum undrawn amount of

borrowing capacity under our unsecured senior line of credit equal to any outstanding balance under our commercial paper program.

We use borrowings under the program to fund short-term capital needs. The notes issued under our commercial paper program are

sold under customary terms in the commercial paper market. They are typically issued at a discount to par, representing a yield to

maturity dictated by market conditions at the time of issuance. In the event we are unable to issue commercial paper notes or refinance

outstanding commercial paper notes under terms equal to or more favorable than those under the unsecured senior line of credit, we

expect to borrow under the unsecured senior line of credit. The commercial paper notes sold during the three months ended

March 31, 2026 were issued at a weighted-average yield to maturity of 4.08%. As of March 31, 2026, we had $1.35 billion of

commercial paper notes outstanding.

In January 2026 and April 2026, we repaid, upon maturity,  $300.0 million of 4.30% unsecured senior notes payable and

$350.0 million of 3.80% unsecured senior notes payable, respectively. These repayments were funded temporarily with borrowings

under our commercial paper program, which will be repaid through planned dispositions and sales of partial interests included in our

2026 guidance. No gain or loss was incurred in connection with these repayments.

In February 2026, we completed tender offers to repurchase an aggregate debt principal amount of $1.33 billion across a

portion of our outstanding 4.00% Senior Notes due 2050, 3.00% Senior Notes due 2051, and 3.55% Senior Notes due 2052. Cash

consideration paid was $952.2 million. The repurchase was primarily funded through the issuance of $750.0 million of 5.25% unsecured

senior notes due 2036, and approximately $200 million of short-term borrowings under our commercial paper program, which we expect

to repay through planned 2026 dispositions and sales of partial interests. In connection with the debt repurchase, we recognized a gain

on early extinguishment of debt aggregating $366.4 million, including the write-off of unamortized debt issuance costs and other

transaction-related costs.

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The following table presents our average debt outstanding and weighted-average interest rates during the three months ended

March 31, 2026 (dollars in thousands):

Three Months Ended March 31, 2026
Average Debt<br><br>Outstanding Weighted-Average<br><br>Interest Rate
Long-term fixed-rate debt $11,432,675 3.94%
Short-term variable-rate unsecured senior line of credit and commercial paper<br><br>program debt 1,736,226 4.05
Blended average interest rate 13,168,901 3.95
Loan fee amortization and annual facility fee related to unsecured senior line of credit N/A 0.13
Total/weighted average $13,168,901 4.08%

Real estate dispositions and sales of partial interests

We expect to continue to focus on the disciplined execution of select sales of real estate. Future sales will provide an important

source of capital to fund our development and redevelopment projects and potential opportunistic share repurchases, capital for growth,

and to reduce debt. We may also consider additional sales of partial interests in core Class A/A+ properties, development projects, and/

or land. For the year ending December 31, 2026, we expect real estate dispositions and sales of partial interests in real estate assets to

range from $2.10 billion to $3.70 billion. The amount of asset sales necessary to meet our forecasted sources of capital will vary

depending upon the amount of EBITDA associated with the assets sold.

Refer to Note 3 – “Investments in real estate” and Note 4 – “Consolidated and unconsolidated real estate joint ventures,” and

Note 14 – “Stockholders’ equity” to our unaudited consolidated financial statements in Item 1 and to “Dispositions and sales of partial

interests” in Item 2 for additional information on our real estate dispositions.

As a REIT, we are generally subject to a 100% tax on the net income from real estate asset sales that the IRS characterizes as

“prohibited transactions.” We do not expect our sales will be categorized as prohibited transactions. However, unless we meet certain

“safe harbor” requirements, whether a real estate asset sale is a “prohibited transaction” will be based on the facts and circumstances

of the sale. Our real estate asset sales may not always meet such “safe harbor” requirements. Refer to “Item 1A. Risk factors” in our

annual report on Form 10-K for the year ended December 31, 2025 for additional information about the “prohibited transaction” tax.

Common equity transactions

During the three months ended March 31, 2026, we did not issue any common stock under our ATM program. As of March 31,

2026, the remaining aggregate amount available under our ATM program for future sales of common stock was $1.47 billion.

Other sources

As a well-known seasoned issuer, we may, from time to time, issue securities at our discretion based on our needs and market

conditions, including, as necessary, to balance our use of incremental debt capital.

Additionally, we, together with joint venture partners, hold interests in real estate joint ventures that we consolidate in our

financial statements. These existing joint ventures provide significant equity capital to fund a portion of our future construction spending,

and our joint venture partners may also contribute equity into these entities for financing-related activities. From April 1, 2026 through

December 31, 2027 and beyond, we expect to receive capital contributions aggregating $120.0 million from existing consolidated real

estate joint venture partners to fund construction. During the year ending December 31, 2026, contributions from noncontrolling

interests from existing joint venture partners are expected to aggregate to up to $100.0 million at the midpoint of our guidance range for

2026 construction spending.

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Uses of capital

Summary of capital expenditures

One of our primary uses of capital relates to the development, redevelopment, pre-construction, and construction of properties.

We currently have projects in our development and redevelopment pipeline aggregating 3.4 million RSF of Class A/A+ properties

undergoing construction. We incur capitalized construction costs related to development, redevelopment, pre-construction, and other

construction activities. We also incur additional capitalized project costs, including interest, property taxes, insurance, and other costs

directly related and essential to the development, redevelopment, pre-construction, or construction of a project, during periods when

activities necessary to prepare an asset for its intended use are in progress. Refer to “New Class A/A+ development and redevelopment

properties: under construction” and “Summary of capital expenditures” in Item 2 for additional information on our capital expenditures.

We capitalize interest cost as a cost of the project only during the period in which activities necessary to prepare an asset for

its intended use are ongoing, provided that expenditures for the asset have been made and interest cost has been incurred. Capitalized

interest, classified in investments in real estate in our consolidated balance sheets, aggregated $70.0 million for the three months

ended March 31, 2026, a decrease from $80.1 million capitalized during the three months ended March 31, 2025. This reflects a lower

weighted-average capitalized cost basis of $6.86 billion for the three months ended March 31, 2026, as compared to $8.03 billion for

the three months ended March 31, 2025.

Property taxes, insurance on real estate, and indirect project costs, such as construction, administration, legal fees, and office

costs that clearly relate to projects under development or construction, are capitalized as incurred during the period an asset is

undergoing activities to prepare it for its intended use. We capitalized payroll and other indirect costs related to development,

redevelopment, pre-construction, and construction projects aggregating $17.6 million and $24.8 million, and property taxes, insurance

on real estate, and indirect project costs aggregating $36.9 million and $36.2 million during the three months ended March 31, 2026 and

2025, respectively.

Pre-construction activities include entitlements, permitting, design, site work, and other activities preceding commencement of

construction of aboveground building improvements. The advancement of pre-construction efforts is focused on reducing the time

required to deliver projects to prospective tenants. These critical activities add significant value for future ground-up development and

are required for the vertical construction of buildings. Should we cease activities necessary to prepare an asset for its intended use, the

interest, taxes, insurance, and certain other direct and indirect project costs related to the asset would be expensed as incurred.

Expenditures for repairs and maintenance are expensed as incurred.

Fluctuations in our development, redevelopment, and construction activities could result in significant changes to total

expenses and net income. For example, had we experienced a 10% reduction in development, redevelopment, and construction

activities without a corresponding decrease in indirect project costs, including interest and payroll, total expenses would have increased

by approximately $12.5 million for the three months ended March 31, 2026.

We use third-party brokers to assist in our leasing activity, who are paid on a contingent basis upon successful leasing. We are

required to capitalize initial direct costs related to successful leasing transactions that result directly from and are essential to the lease

transaction and would not have been incurred had that lease transaction not been successfully executed. During the three months

ended March 31, 2026, we capitalized total initial direct leasing costs of $18.8 million. Costs that we incur to negotiate or arrange a

lease regardless of its outcome, such as fixed employee compensation, tax, or legal advice to negotiate lease terms, and other costs,

are expensed as incurred.

Real estate acquisitions and common stock repurchase program

On December 8, 2025, we announced that our Board of Directors authorized a new common stock repurchase program that

allows for the repurchase of up to $500.0 million of our common stock through December 31, 2026. This new program replaced our

prior stock repurchase program. As of the date of this report, no repurchases have been made under the new program and

$500.0 million remains available for future share repurchases.

We did not make any real estate acquisitions during the three months ended March 31, 2026.

98

Dividends

During the three months ended March 31, 2026 and 2025, we paid common stock dividends of $123.8 million and

$230.0 million, respectively. The decrease of $106.2 million in dividends paid on our common stock for the three months ended

March 31, 2026, compared to the three months ended March 31, 2025, was primarily due to a decrease in the related dividends to

$0.72 per common share paid for the three months ended March 31, 2026 from $1.32 per common share paid during the three months

ended March 31, 2025.

We have historically funded the payment of our common stock dividends using net cash provided by operating activities, as

adjusted. We expect to continue funding future quarterly common stock dividends from net cash provided by operating activities, as

adjusted, which may be supplemented by proceeds from periodic asset dispositions, issuances of additional debt and/or equity

securities, and borrowings under our unsecured senior line of credit and/or our commercial paper program. Future dividends are at the

discretion of our Board and subject to various considerations, including net income, cash flows, capital requirements, debt covenants,

market conditions, dividend yield, taxable income, payout ratios, and other factors. There can be no assurance that we will maintain our

dividends at the current level or increase dividends in the future.

Unsecured senior notes payable and unsecured senior line of credit

The requirements of, and our actual performance with respect to, the key financial covenants under our unsecured senior

notes payable as of March 31, 2026 were as follows:

Covenant Ratios(1) Requirement March 31, 2026
Total Debt to Total Assets Less than or equal to 60% 32%
Secured Debt to Total Assets Less than or equal to 40% —%
Consolidated EBITDA(2) to Interest Expense Greater than or equal to 1.5x 7.7x
Unencumbered Total Asset Value to Unsecured Debt Greater than or equal to 150% 302%

(1)All covenant ratio titles utilize terms as defined in the respective debt agreements.

(2)The calculation of consolidated EBITDA is based on the definitions contained in our loan agreements and is not directly comparable to the computation of EBITDA as

described in Exchange Act Release No. 47226.

In addition, the terms of the indentures, among other things, limit the ability of the Company, Alexandria Real Estate Equities,

L.P., and the Company’s subsidiaries to (i) consummate a merger, or consolidate, or sell all or substantially all of the Company’s assets

and (ii) incur certain secured or unsecured indebtedness.

The requirements of, and our actual performance with respect to, the key financial covenants under our unsecured senior line

of credit as of March 31, 2026 were as follows:

Covenant Ratios(1) Requirement March 31, 2026
Leverage Ratio Less than or equal to 60.0% 34.5%
Secured Debt Ratio Less than or equal to 45.0% —%
Fixed-Charge Coverage Ratio Greater than or equal to 1.50x 3.23x
Unsecured Interest Coverage Ratio Greater than or equal to 1.75x 6.93x

(1)All covenant ratio titles utilize terms as defined in the credit agreement.

In managing our liquidity, we also consider the contractual interest payment obligations associated with our outstanding debt.

Interest payments on our fixed-rate debt are determined based on contractual interest rates, including interest payment dates and

scheduled maturity dates. As of March 31, 2026, 89.2% of our debt was fixed-rate debt. For additional information regarding our debt,

refer to Note 10 – “Secured and unsecured senior debt” to our unaudited consolidated financial statements in Item 1.

99

Ground lease obligations

Ground lease obligations as of March 31, 2026 included leases for 31 of our properties and accounted for approximately 9% of

our total number of properties. Among these 31 properties, 17 properties are subject to ground leases with a weighted-average

remaining lease term of 53 years, including extension options that we are reasonably certain to exercise. These leases are with a single

lessor in our Greater Stanford submarket with whom we have extended three ground leases over the past 10 years.

Our remaining 14 properties subject to ground leases are located across multiple submarkets and have remaining lease terms

ranging from approximately 45 to 81 years. The weighted-average remaining lease term of these ground leases is 73 years, including

extension options that we are reasonably certain to exercise.

In many cases, we seek to extend our ground leases well ahead of their scheduled contractual expirations. If we are

successful in extending ground leases, we could see significant up-front or increased recurring future payments to the ground lessor

and/or increased ground lease expense, which may require us to increase our capital funding needs.

Operating lease agreements

As of March 31, 2026, the remaining contractual payments under ground and office lease agreements in which we are the

lessee aggregated $748.1 million and $19.2 million, respectively. As of March 31, 2026, our operating lease liability, calculated as the

present value of the remaining payments aggregating $767.2 million under our operating lease agreements, including our extension

options that we are reasonably certain to exercise, was $358.6 million and was classified in accounts payable, accrued expenses, and

other liabilities in our consolidated balance sheet. As of March 31, 2026, the weighted-average remaining lease term of operating leases

in which we are the lessee was approximately 60 years, including extension options that we are reasonably certain to exercise, and the

weighted-average discount rate was 4.7%. Our corresponding operating lease right-of-use assets, adjusted for initial direct leasing

costs and other consideration exchanged with the landlord prior to the commencement of the lease, aggregated $693.8 million. We

classify the right-of-use asset in other assets in our consolidated balance sheets. Refer to “Lease accounting” in Note 2 – “Summary of

significant accounting policies” to our unaudited consolidated financial statements in Item 1 for additional information.

Commitments

As of March 31, 2026, remaining aggregate costs under contract for the construction of properties undergoing development,

redevelopment, and improvements under the terms of leases approximated $938.4 million. We expect payments for these obligations to

occur over one to three years, subject to capital planning adjustments from time to time. We may have the ability to cease the

construction of certain projects, which would result in the reduction of our commitments. In addition, we have letters of credit and

performance obligations aggregating $5.3 million.

We are committed to funding approximately $353.0 million related to our non-real estate investments. These funding

commitments are primarily associated with our investments in privately held entities that report NAV and expire at various dates over

the next 12 years, with a weighted-average expiration of 8.0 years as of March 31, 2026.

Our former joint venture partner in the Greater Boston market has an option, subject to certain conditions, to obtain a

$50 million secured loan from us, which, if the option is exercised, will bear interest at 6.5%, with a floor of 9.0% and a term not to

exceed five years. As of March 31, 2026, the option has not been exercised and is set to expire in July 2027.

In connection with the sale of a property in our San Diego market, we entered into a loan agreement with the buyer under

which we committed to provide up to $165.7 million of financing through December 30, 2029. As of March 31, 2026, $47.6 million of the

commitment remained available to be drawn by the borrower.

Exposure to environmental liabilities

In connection with the acquisition of all of our properties, we have obtained Phase I environmental assessments to ascertain

the existence of any environmental liabilities or other issues. The Phase I environmental assessments of our properties have not

revealed any environmental liabilities that we believe would have a material adverse effect on our financial condition or results of

operations taken as a whole, nor are we aware of any material environmental liabilities that have occurred since the Phase I

environmental assessments were completed. In addition, we carry a policy of pollution legal liability insurance covering exposure to

certain environmental losses at substantially all of our properties.

100

Foreign currency translation gains and losses

The following table presents the change in accumulated other comprehensive loss attributable to Alexandria Real Estate

Equities, Inc.’s stockholders during the three months ended March 31, 2026 primarily due to the changes in the foreign exchange rates

for our real estate investments in Canada (in thousands). We reclassify unrealized foreign currency translation gains and losses into net

income as we dispose of these holdings.

Total
Balance as of December 31, 2025 $(29,395)
Other comprehensive loss before reclassifications (1,518)
Reclassification adjustment for loss included in net income (23)
Net other comprehensive loss (1,541)
Balance as of March 31, 2026 $(30,936)

Inflation

As of March 31, 2026, approximately 91% of our leases (on an annual rental revenue basis) were triple net leases, which

require tenants to pay substantially all real estate taxes, insurance, utilities, repairs and maintenance, common area expenses, and

other operating expenses (including increases thereto) in addition to base rent. Approximately 97% of our leases (on an annual rental

revenue basis) contained effective annual rent escalations approximating 3% that were either fixed or indexed based on a consumer

price index or other indices. Accordingly, we do not believe that our cash flows or earnings from real estate operations are subject to

significant risks from inflation. A period of inflation, however, could cause an increase in the cost of issuing new unsecured senior notes

payable and our variable-rate borrowings, including borrowings under our unsecured senior line of credit and commercial paper

program, and secured loans held by our unconsolidated real estate joint ventures.

101

Issuer and guarantor subsidiary summarized financial information

Alexandria Real Estate Equities, Inc. (the “Issuer”) has sold certain debt securities registered under the Securities Act of 1933,

as amended, that are fully and unconditionally guaranteed by Alexandria Real Estate Equities, L.P. (the “LP” or the “Guarantor

Subsidiary”), an indirectly 100% owned subsidiary of the Issuer. The Issuer’s other subsidiaries, including, but not limited to, the

subsidiaries that own substantially all of its real estate (collectively, the “Combined Non-Guarantor Subsidiaries”), will not provide a

guarantee of such securities, including the subsidiaries that are partially or 100% owned by the LP. The following summarized financial

information presents, on a combined basis, balance sheet information as of March 31, 2026 and December 31, 2025, and results of

operations and comprehensive income for the three months ended March 31, 2026 and year ended December 31, 2025 for the Issuer

and the Guarantor Subsidiary. The information presented below excludes eliminations necessary to arrive at the information on a

consolidated basis. In presenting the summarized financial statements, the equity method of accounting has been applied to (i) the

Issuer’s interests in the Guarantor Subsidiary, (ii) the Guarantor Subsidiary’s interests in the Combined Non-Guarantor Subsidiaries,

and (iii) the Combined Non-Guarantor Subsidiaries’ interests in the Guarantor Subsidiary, where applicable, even though all such

subsidiaries meet the requirements to be consolidated under GAAP. All assets and liabilities have been allocated to the Issuer and the

Guarantor Subsidiary generally based on legal entity ownership.

The following tables present combined summarized financial information as of March 31, 2026 and December 31, 2025 and for

the three months ended March 31, 2026 and year ended December 31, 2025 for the Issuer and Guarantor Subsidiary. Amounts

provided do not represent our total consolidated amounts (in thousands):

March 31, 2026 December 31, 2025
Assets:
Cash, cash equivalents, and restricted cash $71,915 $127,100
Other assets 182,017 173,303
Total assets $253,932 $300,403
Liabilities:
Unsecured senior notes payable $11,166,009 $12,047,394
Unsecured senior line of credit and commercial paper 1,353,986 353,161
Other liabilities 420,800 433,707
Total liabilities $12,940,795 $12,834,262 Three Months Ended<br><br>March 31, 2026 Year Ended<br><br>December 31, 2025
--- --- ---
Total revenues $14,508 $48,748
Total expenses (100,208) (350,655)
Gain on early extinguishment of debt 366,435
Net income (loss) 280,735 (301,907)
Net income attributable to unvested restricted stock awards (2,779) (8,417)
Net income (loss) attributable to Alexandria Real Estate Equities, Inc.’s common<br><br>stockholders $277,956 $(310,324)

As of March 31, 2026, 327 of our 339 properties were held indirectly by the REIT’s wholly owned consolidated subsidiary,

Alexandria Real Estate Equities, L.P.

Critical accounting estimates

Refer to our annual report on Form 10-K for the year ended December 31, 2025 for a discussion of our critical accounting

estimates related to recognition of real estate acquired, impairment of long-lived assets, impairment of non-real estate investments, and

monitoring of tenant credit quality.

102

Definitions and reconciliations

This section contains additional information on certain non-GAAP financial measures, including reconciliations from the most

directly comparable financial measure calculated and presented in accordance with GAAP and the reasons why we use these

supplemental measures of performance and believe they provide useful information to investors, as well as the definitions of other

terms used in this report.

Funds from operations and funds from operations, as adjusted, attributable to Alexandria Real Estate Equities, Inc.’s common

stockholders

GAAP-basis accounting for real estate assets utilizes historical cost accounting and assumes that real estate values diminish

over time. In an effort to overcome the difference between real estate values and historical cost accounting for real estate assets, the

Nareit Board of Governors established funds from operations as an improved measurement tool. Since its introduction, funds from

operations has become a widely used non-GAAP financial measure among equity REITs. We believe that funds from operations is

helpful to investors as an additional measure of the performance of an equity REIT. Moreover, we believe that funds from operations, as

adjusted, allows investors to compare our performance to the performance of other real estate companies on a consistent basis, without

having to account for differences recognized because of real estate acquisition and disposition decisions, financing decisions, capital

structure, capital market transactions, variances resulting from the volatility of market conditions outside of our control, or other

corporate activities that may not be representative of the operating performance of our properties.

The 2018 White Paper published by the Nareit Board of Governors (the “Nareit White Paper”) defines funds from operations as

net income (computed in accordance with GAAP), excluding gains or losses on sales of real estate, and impairments of real estate, plus

depreciation and amortization of operating real estate assets, and after adjustments for our share of consolidated and unconsolidated

partnerships and real estate joint ventures. Impairments represent the write-down of assets when fair value over the recoverability

period is less than the carrying value due to changes in general market conditions and do not necessarily reflect the operating

performance of the properties during the corresponding period.

We compute funds from operations, as adjusted, as funds from operations calculated in accordance with the Nareit White

Paper, excluding significant gains, losses, and impairments realized on non-real estate investments, unrealized gains or losses on non-

real estate investments, impairments of real estate primarily consisting of right-of-use assets and pre-acquisition costs related to

projects that we decided to no longer pursue, gains or losses on early extinguishment of debt, changes in the provision for expected

credit losses on financial instruments, significant termination fees, acceleration of stock compensation expense due to the resignations

of executive officers, deal costs, the income tax effect related to such items, and the amount of such items that is allocable to our

unvested restricted stock awards. We compute the amount that is allocable to our unvested restricted stock awards with nonforfeitable

dividends using the two-class method. Under the two-class method, we allocate net income (after amounts attributable to noncontrolling

interests) to common stockholders and to unvested restricted stock awards with nonforfeitable dividends by applying the respective

weighted-average shares outstanding during each quarter-to-date and year-to-date period. This may result in a difference of the

summation of the quarter-to-date and year-to-date amounts. Neither funds from operations nor funds from operations, as adjusted,

should be considered as alternatives to net income (determined in accordance with GAAP) as indications of financial performance, or to

cash flows from operating activities (determined in accordance with GAAP) as measures of liquidity, nor are they indicative of the

availability of funds for our cash needs, including our ability to make distributions.

We are not able to forecast the net income of future periods without unreasonable effort, and therefore do not provide a

reconciliation for funds from operations on a forward-looking basis. This is due to the inherent difficulty of forecasting the timing and/or

amount of items that depend on market conditions outside of our control, including the timing of dispositions, capital events, and

financing decisions, as well as components such as gain on sales of real estate, unrealized gains or losses on non-real estate

investments, impairments of real estate, impairments of non-real estate investments, and changes in provision for expected credit

losses on financial instruments. Our attempt to predict these amounts may produce significant but inaccurate estimates, which would

potentially be misleading for our investors.

103

The following tables present a reconciliation of net income (loss) attributable to Alexandria Real Estate Equities, Inc.’s common

stockholders, the most directly comparable financial measure presented in accordance with GAAP, including our share of amounts from

consolidated and unconsolidated real estate joint ventures, to funds from operations attributable to Alexandria Real Estate Equities,

Inc.’s common stockholders – diluted, and funds from operations attributable to Alexandria Real Estate Equities, Inc.’s common

stockholders – diluted, as adjusted, and the related per share amounts for the three months ended March 31, 2026 and 2025 (in

thousands, except per share amounts). Per share amounts may not add due to rounding.

Three Months Ended March 31,
2026 2025
Net income (loss) attributable to Alexandria Real Estate Equities, Inc.’s common stockholders –<br><br>basic and diluted 358,874 $(11,599)
Depreciation and amortization of real estate assets 303,296 339,381
Noncontrolling share of depreciation and amortization from consolidated real estate JVs (29,473) (33,411)
Our share of depreciation and amortization from unconsolidated real estate JVs 914 1,054
Gain on sales of real estate (13,165)
Impairment of real estate – rental properties and land 5,499
Allocation to unvested restricted stock awards (2,181) (686)
Funds from operations attributable to Alexandria Real Estate Equities, Inc.’s common stockholders<br><br>– diluted(2) 636,929 281,574
Unrealized losses on non-real estate investments 10,332 68,145
Impairment of non-real estate investments 12,448 11,180
Impairment of real estate 32,154
Gain on early extinguishment of debt (366,435)
Increase in provision for expected credit losses on financial instruments 285
Allocation to unvested restricted stock awards 2,674 (1,329)
Funds from operations attributable to Alexandria Real Estate Equities, Inc.’s common stockholders<br><br>– diluted, as adjusted 295,948 $392,009

All values are in US Dollars.

(1)Primarily represents an incremental impairment charge recognized during the three months ended March 31, 2026 in connection with the amendment of the sales

agreement for our Canada portfolio, which was classified as held for sale as of December 31, 2025. Refer to Note 3 – “Sales of real estate assets and impairment of real

estate” under Item 1 to our unaudited consolidated financial statements for additional details.

(2)Calculated in accordance with standards established by the Nareit Board of Governors.

(3)Primarily related to two non-real estate investments in privately held entities that do not report NAV.

(4)In February 2026, we completed tender offers to repurchase debt principal aggregating $1.33 billion across a portion of our outstanding 4.00% Senior Notes due 2050,

3.00% Senior Notes due 2051, and 3.55% Senior Notes due 2052 for $952.2 million. The gain includes the write-off of unamortized debt issuance costs and other

transaction-related costs. Refer to Note 10 – “Secured and unsecured senior debt” under Item 1 to our unaudited consolidated financial statements for additional details.

104

Three Months Ended March 31,
(Per share) 2026 2025
Net income (loss) per share attributable to Alexandria Real Estate Equities, Inc.’s common<br><br>stockholders – diluted $2.10 $(0.07)
Depreciation and amortization of real estate assets 1.61 1.80
Gain on sales of real estate (0.08)
Impairment of real estate – rental properties and land 0.03
Allocation to unvested restricted stock awards (0.01)
Funds from operations per share attributable to Alexandria Real Estate Equities, Inc.’s common<br><br>stockholders – diluted 3.73 1.65
Unrealized losses on non-real estate investments 0.06 0.40
Impairment of non-real estate investments 0.07 0.07
Impairment of real estate 0.19
Gain on early extinguishment of debt (2.14)
Allocation to unvested restricted stock awards 0.01 (0.01)
Funds from operations per share attributable to Alexandria Real Estate Equities, Inc.’s common<br><br>stockholders – diluted, as adjusted $1.73 $2.30
Weighted-average shares of common stock outstanding – diluted(1)
Earnings per share – diluted 170,867 170,522
Funds from operations – diluted, per share 170,867 170,599
Funds from operations – diluted, as adjusted, per share 170,867 170,599

(1)Refer to “Weighted-average shares of common stock outstanding – diluted” in this section for additional information.

The following table reconciles net income (loss) to funds from operations for the share of consolidated real estate joint

ventures attributable to noncontrolling interests and our share of unconsolidated real estate joint ventures for the three months ended

March 31, 2026 (in thousands):

Three Months Ended March 31, 2026
Noncontrolling Interest<br><br>Share of Consolidated<br><br>Real Estate Joint Ventures Our Share of<br><br>Unconsolidated Real<br><br>Estate Joint Ventures
Net income (loss) $36,724 $(147)
Depreciation and amortization of real estate assets 29,473 914
Funds from operations $66,197 $767

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Adjusted EBITDA and Adjusted EBITDA margin

We use Adjusted EBITDA as a supplemental performance measure of our operations, for financial and operational decision-

making, and as a supplemental means of evaluating period-to-period comparisons on a consistent basis. Adjusted EBITDA is calculated

as earnings before interest, taxes, depreciation, and amortization (“EBITDA”), excluding stock compensation expense, gains or losses

on early extinguishment of debt, gains or losses on sales of real estate, impairments of real estate, changes in provision for expected

credit losses on financial instruments, and significant termination fees. Adjusted EBITDA also excludes unrealized gains or losses and

significant realized gains or losses and impairments that result from our non-real estate investments. These non-real estate investment

amounts are classified in our consolidated statements of operations outside of total revenues.

We believe Adjusted EBITDA provides investors with relevant and useful information as it allows investors to evaluate the

operating performance of our business activities without having to account for differences recognized because of investing and

financing decisions related to our real estate and non-real estate investments, our capital structure, capital market transactions, and

variances resulting from the volatility of market conditions outside of our control. For example, we exclude gains or losses on the early

extinguishment of debt to allow investors to measure our performance independent of our indebtedness and capital structure. We

believe that adjusting for the effects of impairments and gains or losses on sales of real estate, significant impairments and realized

gains or losses on non-real estate investments, changes in provision for expected credit losses on financial instruments, and significant

termination fees allows investors to evaluate performance from period to period on a consistent basis without having to account for

differences recognized because of investing and financing decisions related to our real estate and non-real estate investments or other

corporate activities that may not be representative of the operating performance of our properties.

In addition, we believe that excluding charges related to stock compensation and unrealized gains or losses facilitates for

investors a comparison of our business activities across periods without the volatility resulting from market forces outside of our control.

Adjusted EBITDA has limitations as a measure of our performance. Adjusted EBITDA does not reflect our historical expenditures or

future requirements for capital expenditures or contractual commitments. While Adjusted EBITDA is a relevant measure of performance,

it does not represent net income (loss) or cash flows from operations calculated and presented in accordance with GAAP, and it should

not be considered as an alternative to those indicators in evaluating performance or liquidity.

In order to calculate the Adjusted EBITDA margin, we divide Adjusted EBITDA by total revenues as presented in our

consolidated statements of operations. We believe that this supplemental performance measure provides investors with additional

useful information regarding the profitability of our operating activities.

We are not able to forecast the net income of future periods without unreasonable effort, and therefore do not provide a

reconciliation for Adjusted EBITDA on a forward-looking basis. This is due to the inherent difficulty of forecasting the timing and/or

amount of items that depend on market conditions outside of our control, including the timing of dispositions, capital events, and

financing decisions, as well as quarterly components such as gain on sales of real estate, unrealized gains or losses on non-real estate

investments, impairments of real estate, impairments of non-real estate investments, and changes in provision for expected credit

losses on financial instruments. Our attempt to predict these amounts may produce significant but inaccurate estimates, which would

potentially be misleading for our investors.

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The following table reconciles net income (loss), the most directly comparable financial measure calculated and presented in

accordance with GAAP, to Adjusted EBITDA and calculates the Adjusted EBITDA margin for the three months ended March 31, 2026

and 2025 (dollars in thousands):

Three Months Ended March 31,
2026 2025
Net income 398,377 $38,662
Interest expense 64,584 50,876
Income taxes 3,225 1,145
Depreciation and amortization 305,441 342,062
Stock compensation expense 11,032 10,064
Gain on early extinguishment of debt (366,435)
Gain on sales of real estate (13,165)
Unrealized losses on non-real estate investments 10,332 68,145
Impairment of real estate 5,499 32,154
Impairment of non-real estate investments 12,448 11,180
Decrease in provision for expected credit losses on financial instruments 285
Adjusted EBITDA 444,503 $541,408
Total revenues 671,022 $758,158
Adjusted EBITDA margin 66% 71%

All values are in US Dollars.

(1)In February 2026, we completed tender offers to repurchase debt principal aggregating $1.33 billion across a portion of our outstanding 4.00% Senior Notes due 2050,

3.00% Senior Notes due 2051, and 3.55% Senior Notes due 2052 for $952.2 million. The gain includes the write-off of unamortized debt issuance costs and other

transaction-related costs. Refer to Note 10 – “Secured and unsecured senior debt” under Item 1 to our unaudited consolidated financial statements.

Annual rental revenue

Annual rental revenue represents the annualized fixed base rental obligations, calculated in accordance with GAAP. It includes

the amortization of deferred revenue related to tenant-funded and tenant-built landlord improvements for leases in effect as of the end

of the period, related to our operating RSF. Annual rental revenue is presented using 100% of the annual rental revenue from our

consolidated properties and our share of annual rental revenue for our unconsolidated real estate joint ventures. Annual rental revenue

per RSF is computed by dividing annual rental revenue by the sum of 100% of the RSF of our consolidated properties and our share of

the RSF of properties held in unconsolidated real estate joint ventures. As of March 31, 2026, approximately 91% of our leases (on an

annual rental revenue basis) were triple net leases, which require tenants to pay substantially all real estate taxes, insurance, utilities,

repairs and maintenance, common area expenses, and other operating expenses (including increases thereto) in addition to base rent.

Annual rental revenue excludes these operating expenses recovered from our tenants. Amounts recovered from our tenants related to

these operating expenses, along with base rent, are classified in income from rentals in our consolidated statements of operations.

Capitalization rates

Capitalization rates are calculated based on net operating income and net operating income (cash basis) annualized,

excluding lease termination fees, on stabilized operating assets for the quarter preceding the date on which the property is sold, or

near-term prospective net operating income.

Capitalized interest

We capitalize interest cost as a cost of a project during periods for which activities necessary to develop, redevelop, or

reposition a project for its intended use are ongoing, provided that expenditures for the asset have been made and interest cost has

been incurred. Activities necessary to develop, redevelop, or reposition a project include pre-construction activities such as

entitlements, permitting, design, site work, and other activities preceding commencement of construction of aboveground building

improvements. The advancement of pre-construction efforts is focused on reducing the time required to deliver projects to prospective

tenants. These critical activities add significant value for future ground-up development and are required for the vertical construction of

buildings. If we cease activities necessary to prepare a project for its intended use, interest costs related to such project are expensed

as incurred.

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Cash interest

Cash interest is equal to interest expense calculated in accordance with GAAP plus capitalized interest, less amortization of

loan fees and debt premiums (discounts). Refer to “Fixed-charge coverage ratio” in this section for a reconciliation of interest expense,

the most directly comparable financial measure calculated and presented in accordance with GAAP, to cash interest.

Class A/A+ properties and AAA locations

Class A/A+ properties are properties clustered in AAA locations that provide innovative tenants with highly dynamic and

collaborative environments that enhance their ability to successfully recruit and retain world-class talent and inspire productivity,

efficiency, creativity, and success. These properties are typically well-located, professionally managed, and well-maintained, offering a

wide range of amenities and featuring premium construction materials and finishes. Class A/A+ properties are generally newer or have

undergone substantial redevelopment and are generally expected to command higher annual rental rates compared to other classes of

similar properties. AAA locations are in close proximity to concentrations of specialized skills, knowledge, institutions, and related

businesses. It is important to note that our definition of property classification may not be directly comparable to other equity REITs.

Credit rating

Represents the credit ratings assigned by S&P Global Ratings or Moody’s Ratings as of March 31, 2026. A credit rating is not

a recommendation to buy, sell, or hold securities and may be subject to revision or withdrawal at any time.

Development, redevelopment, and pre-construction

A key component of our business model is our disciplined allocation of capital to the development and redevelopment of new

Class A/A+ properties, as well as property enhancements identified during the underwriting of certain acquired properties. These efforts

are primarily concentrated in collaborative Megacampus ecosystems within AAA life science innovation clusters, as well as other

strategic locations that support innovation and growth. These projects are generally focused on providing high-quality, generic, and

reusable spaces that meet the real estate requirements of a wide range of tenants. Upon completion, each development or

redevelopment project is expected to generate increases in rental income, net operating income, and cash flows. Our development and

redevelopment projects are generally in locations that are highly desirable to high-quality entities, which we believe results in higher

occupancy levels, longer lease terms, higher rental income, higher returns, and greater long-term asset value.

Development projects generally consist of the ground-up development of generic and reusable laboratory facilities.

Redevelopment projects generally consist of the permanent change in use of acquired office, warehouse, or shell space into facilities

designed for life science innovation or advanced technology. We generally will not commence new development projects for

aboveground construction of new Class A/A+ laboratory space without first securing significant pre-leasing for such space, except when

there is solid market demand for high-quality Class A/A+ properties.

Pre-construction activities include entitlements, permitting, design, site work, and other activities preceding commencement of

construction of aboveground building improvements. The advancement of pre-construction efforts is focused on reducing the time

required to deliver projects to prospective tenants. These critical activities add significant value for future ground-up development and

are required for the vertical construction of buildings. Ultimately, these projects will provide high-quality facilities and are expected to

generate significant revenue and cash flows.

Development, redevelopment, and pre-construction spending also includes the following costs: (i) amounts to bring certain

acquired properties up to market standard and/or other costs identified during the acquisition process (generally within two years of

acquisition) and (ii) permanent conversion of space for highly flexible, move-in-ready laboratory space to foster the growth of promising

early- and growth-stage life science companies.

Revenue-enhancing and repositioning capital expenditures represent spending to reposition or significantly change the use of

a property, including through improvement in the asset quality from Class B to Class A/A+.

Non-revenue-enhancing capital expenditures represent costs required to maintain the current revenues of a stabilized

property, including the associated costs for renewed and re-leased space.

Dividend payout ratio (common stock)

Dividend payout ratio (common stock) is the ratio of the absolute dollar amount of dividends on our common stock (shares of

common stock outstanding on the respective record dates multiplied by the related dividend per share) to funds from operations

attributable to Alexandria’s common stockholders – diluted, as adjusted.

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Dividend yield

Dividend yield for the quarter represents the annualized quarter dividend divided by the closing common stock price at the end

of the quarter.

Fixed-charge coverage ratio

Fixed-charge coverage ratio is a non-GAAP financial measure representing the ratio of Adjusted EBITDA to cash interest and

fixed charges. We believe that this ratio is useful to investors as a supplemental measure of our ability to satisfy fixed financing

obligations and preferred stock dividends. Cash interest is equal to interest expense calculated in accordance with GAAP plus

capitalized interest, less amortization of loan fees and debt premiums (discounts).

The following table reconciles interest expense, the most directly comparable financial measure calculated and presented in

accordance with GAAP, to cash interest and computes fixed-charge coverage ratio for the three months ended March 31, 2026 and

2025 (dollars in thousands):

Three Months Ended March 31,
2026 2025
Adjusted EBITDA $444,503 $541,408
Interest expense $64,584 $50,876
Capitalized interest 69,973 80,065
Amortization of loan fees (4,428) (4,691)
Amortization of debt discounts (320) (349)
Cash interest and fixed charges $129,809 $125,901
Fixed-charge coverage ratio:
– quarter annualized 3.4x 4.3x
– trailing 12 months 3.8x 4.4x

We are not able to forecast the net income of future periods without unreasonable effort, and therefore do not provide a

reconciliation for fixed-charge coverage ratio on a forward-looking basis. This is due to the inherent difficulty of forecasting the timing

and/or amount of items that depend on market conditions outside of our control, including the timing of dispositions, capital events, and

financing decisions, as well as quarterly components such as gain on sales of real estate, unrealized gains or losses on non-real estate

investments, impairments of real estate, impairments of non-real estate investments, and changes in provision for expected credit

losses on financial instruments. Our attempt to predict these amounts may produce significant but inaccurate estimates, which would

potentially be misleading for our investors.

Gross assets

Gross assets are calculated as total assets plus accumulated depreciation as of March 31, 2026 and December 31, 2025 (in

thousands):

March 31, 2026 December 31, 2025
Total assets $34,167,397 $34,081,835
Accumulated depreciation 6,393,658 6,127,525
Gross assets $40,561,055 $40,209,360

Incremental annual net operating income on development and redevelopment projects

Incremental annual net operating income represents the amount of net operating income, on an annual basis, expected to be

realized upon a project being placed into service and achieving full occupancy. Incremental annual net operating income is calculated

as the initial stabilized yield multiplied by the project’s total cost at completion.

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Initial stabilized yield (unlevered)

Initial stabilized yield is calculated as the estimated amounts of net operating income at stabilization divided by our investment

in the property. For this calculation, we exclude any tenant-funded and tenant-built landlord improvements from our investment in the

property. Our initial stabilized yield excludes the benefit of leverage. Our cash rents related to our development and redevelopment

projects are generally expected to increase over time due to contractual annual rent escalations. Our estimates for initial stabilized

yields, initial stabilized yields (cash basis), and total costs at completion represent our initial estimates at the commencement of the

project. We expect to update this information upon completion of the project, or sooner if there are significant changes to the expected

project yields or costs.

•Initial stabilized yield reflects rental income, including contractual rent escalations and any rent concessions over the

term(s) of the lease(s), calculated on a straight-line basis, and any amortization of deferred revenue related to tenant-

funded and tenant-built landlord improvements.

•Initial stabilized yield (cash basis) reflects cash rents at the stabilization date after initial rental concessions, if any, have

elapsed and our total cash investment in the property.

Investment-grade or publicly traded large cap tenants

Investment-grade or publicly traded large cap tenants represent tenants that are investment-grade rated or publicly traded

companies with an average daily market capitalization greater than $10 billion for the twelve months ended March 31, 2026, as reported

by Bloomberg Professional Services. Credit ratings from Moody’s Ratings and S&P Global Ratings reflect credit ratings of the tenant’s

parent entity, and there can be no assurance that a tenant’s parent entity will satisfy the tenant’s lease obligation upon such tenant’s

default. We monitor the credit quality and related material changes of our tenants. Material changes that cause a tenant’s market

capitalization to decrease below $10 billion, which are not immediately reflected in the twelve-month average, may result in their

exclusion from this measure.

Investments in real estate

The following table presents our new Class A/A+ development and redevelopment pipeline, excluding properties held for sale,

as a percentage of gross assets and as a percentage of annual rental revenue as of March 31, 2026 (dollars in thousands):

Book Value Percentage of<br><br>Gross Assets
Projects under active construction $3,117,332 8%
Future development projects(1) and land parcels primarily located in Megacampuses 3,740,237 9
Total Class A/A+ development and redevelopment pipeline, excluding properties held for<br><br>sale 6,857,569 17
Properties held for sale – land parcels 230,905 1
Total Class A/A+ development and redevelopment pipeline $7,088,474 18%

(1)Includes projects with existing buildings that are generating or can generate operating cash flows. Also includes development rights associated with existing operating

campuses.

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The square footage presented in the table below is classified as operating as of March 31, 2026. These lease expirations or

vacant space at recently acquired properties represent future opportunities for which we intend, subject to market conditions and

leasing, to commence first-time conversion from non-laboratory space to laboratory space, or to commence future ground-up

development:

Dev/Redev RSF of Lease Expirations Targeted for<br><br>Development and Redevelopment
Property/Submarket 2026 2027 Thereafter(1) Total
Future projects:
446, 458, and 500 Arsenal Street/Cambridge/Inner Suburbs Dev 116,623 116,623
1122 and 1150 El Camino Real/South San Francisco Dev 375,232 375,232
2100 Geng Road/Greater Stanford Dev 12,125 12,125
960 Industrial Road/Greater Stanford Dev 112,590 112,590
Campus Point by Alexandria/University Town Center Dev 96,805 96,805
Sequence District by Alexandria/Sorrento Mesa Dev/Redev 555,754 555,754
Canada Redev 247,743 247,743
Total 1,516,872 1,516,872

(1)Includes vacant square footage as of March 31, 2026.

Joint venture financial information

We present components of balance sheet and operating results information related to our real estate joint ventures, which are

not presented, or intended to be presented, in accordance with GAAP. We present the proportionate share of certain financial line items

as follows: (i) for each real estate joint venture that we consolidate in our financial statements, which are controlled by us through

contractual rights or majority voting rights, but of which we own less than 100%, we apply the noncontrolling interest economic

ownership percentage to each financial item to arrive at the amount of such cumulative noncontrolling interest share of each component

presented; and (ii) for each real estate joint venture that we do not control and do not consolidate, which are instead controlled jointly or

by our joint venture partners through contractual rights or majority voting rights, we apply our economic ownership percentage to each

financial item to arrive at our proportionate share of each component presented.

The components of balance sheet and operating results information related to our real estate joint ventures do not represent

our legal claim to those items. For each entity that we do not wholly own, the joint venture agreement generally determines what equity

holders can receive upon capital events, such as sales or refinancing, or in the event of a liquidation. Equity holders are normally

entitled to their respective legal ownership of any residual cash from a joint venture only after all liabilities, priority distributions, and

claims have been repaid or satisfied.

We believe that this information can help investors estimate the balance sheet and operating results information related to our

partially owned entities. Presenting this information provides a perspective not immediately available from consolidated financial

statements and one that can supplement an understanding of the joint venture assets, liabilities, revenues, and expenses included in

our consolidated results.

The components of balance sheet and operating results information related to our real estate joint ventures are limited as an

analytical tool as the overall economic ownership interest does not represent our legal claim to each of our joint ventures’ assets,

liabilities, or results of operations. In addition, joint venture financial information may include financial information related to the

unconsolidated real estate joint ventures that we do not control. We believe that, to facilitate investors’ clear understanding of our

operating results and our total assets and liabilities, joint venture financial information should be examined in conjunction with our

consolidated statements of operations and balance sheets. Joint venture financial information should not be considered an alternative

to our consolidated financial statements, which are presented and prepared in accordance with GAAP.

Megacampus™

A Megacampus ecosystem is a cluster campus that consists of approximately 1 million RSF or greater, including operating,

active development/redevelopment, and land RSF less operating RSF expected to be demolished.

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The following table reconciles our annual rental revenue and development and redevelopment pipeline RSF, excluding

properties classified as held for sale, as of March 31, 2026 (dollars in thousands):

Annual Rental<br><br>Revenue Development and<br><br>Redevelopment<br><br>Pipeline RSF
Megacampus $1,414,438 16,919,119
Core and non-core 388,856 4,990,866
Total $1,803,294 21,909,985
Megacampus as a percentage of annual rental revenue and of total development and<br><br>redevelopment pipeline RSF 78% 77%

Net cash provided by operating activities, as adjusted

We use net cash provided by operating activities, as adjusted, as a supplemental measure for financial and operational

decision-making, and as a supplemental means of evaluating period-to-period comparisons on a consistent basis. Net cash provided by

operating activities, as adjusted, is calculated as net cash provided by operating activities as shown in our consolidated statements of

cash flows, adjusted for changes in operating assets and liabilities (as they represent timing differences), and reduced by dividends and

distributions to noncontrolling interests (excludes liquidating distributions from asset sales).

We believe net cash provided by operating activities, as adjusted, provides investors with relevant and useful information as it

allows investors to evaluate our operating cash flows on a more consistent basis that excludes period-to-period timing differences in

operating assets and liabilities (working capital) and reflects cash dividends and distributions paid quarterly.

The following table reconciles net cash flows from operating activities, the most directly comparable financial measure

presented in accordance with GAAP, to net cash provided by operating activities, as adjusted:

Three Months Ended March 31,
(in thousands) 2026 2025
Net cash provided by operating activities $196,624 207,949
Decreases in operating assets and liabilities 143,523 220,294
Common stock dividends paid (123,752) (229,987)
Distributions to noncontrolling interests (60,111) (66,034)
Net cash provided by operating activities, as adjusted $156,284 $132,222

Net debt and preferred stock to Adjusted EBITDA

Net debt and preferred stock to Adjusted EBITDA is a non-GAAP financial measure that we believe is useful to investors as a

supplemental measure of evaluating our balance sheet leverage. Net debt and preferred stock is equal to the sum of total consolidated

debt less cash, cash equivalents, and restricted cash, plus preferred stock outstanding as of the end of the period. Refer to “Adjusted

EBITDA and Adjusted EBITDA margin” in this section for further information on the calculation of Adjusted EBITDA.

We are not able to forecast the net income of future periods without unreasonable effort, and therefore do not provide a

reconciliation for net debt and preferred stock to Adjusted EBITDA on a forward-looking basis. This is due to the inherent difficulty of

forecasting the timing and/or amount of items that depend on market conditions outside of our control, including the timing of

dispositions, capital events, and financing decisions, as well as quarterly components such as gain on sales of real estate, unrealized

gains or losses on non-real estate investments, impairments of real estate, impairments of non-real estate investments, and changes in

provision for expected credit losses on financial instruments. Our attempt to predict these amounts may produce significant but

inaccurate estimates, which would potentially be misleading for our investors.

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The following table reconciles debt to net debt and preferred stock and computes the ratio to Adjusted EBITDA as of March 31,

2026 and December 31, 2025 (dollars in thousands):

March 31, 2026 December 31, 2025
Unsecured senior notes payable $11,166,009 $12,047,394
Unsecured senior line of credit and commercial paper 1,353,986 353,161
Unamortized deferred financing costs 69,071 74,314
Cash and cash equivalents (418,720) (549,062)
Restricted cash (4,665) (4,693)
Preferred stock
Net debt and preferred stock $12,165,681 $11,921,114
Adjusted EBITDA:
– quarter annualized $1,778,012 $2,097,444
– trailing 12 months $2,044,906 $2,141,811
Net debt and preferred stock to Adjusted EBITDA:
– quarter annualized 6.8x 5.7x
– trailing 12 months 5.9x 5.6x

Net operating income, net operating income (cash basis), and operating margin

The following table reconciles net income (loss) to net operating income and net operating income (cash basis) and computes

operating margin for the three months ended March 31, 2026 and 2025 (dollars in thousands):

Three Months Ended March 31,
2026 2025
Net income $398,377 $38,662
Equity in losses of unconsolidated real estate joint ventures 147 507
General and administrative expenses 34,685 30,675
Interest expense 64,584 50,876
Depreciation and amortization 305,441 342,062
Impairment of real estate 5,499 32,154
Gain on early extinguishment of debt (366,435)
Gain on sales of real estate (13,165)
Investment loss 4,582 49,992
Net operating income 446,880 531,763
Straight-line rent revenue (17,862) (22,023)
Amortization of deferred revenue related to tenant-funded and -built landlord improvements (5,405) (1,651)
Amortization of acquired below-market leases (5,615) (15,222)
Provision for expected credit losses on financial instruments 285
Net operating income (cash basis) $417,998 $493,152
Net operating income (from above) $446,880 $531,763
Total revenues $671,022 $758,158
Operating margin 67% 70%

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Net operating income is a non-GAAP financial measure calculated as net income (loss), the most directly comparable financial

measure calculated and presented in accordance with GAAP, excluding equity in the earnings of our unconsolidated real estate joint

ventures, general and administrative expenses, interest expense, depreciation and amortization, impairments of real estate, gains or

losses on early extinguishment of debt, gains or losses on sales of real estate, and investment income or loss. We believe net operating

income provides useful information to investors regarding our financial condition and results of operations because it primarily reflects

those income and expense items that are incurred at the property level. Therefore, we believe net operating income is a useful measure

for investors to evaluate the operating performance of our consolidated real estate assets. Net operating income on a cash basis is net

operating income adjusted to exclude the effect of straight-line rent, amortization of acquired above- and below-market lease revenue,

amortization of deferred revenue related to tenant-funded and tenant-built landlord improvements, and changes in the provision for

expected credit losses on financial instruments required by GAAP. We believe that net operating income on a cash basis is helpful to

investors as an additional measure of operating performance because it eliminates straight-line rent revenue and the amortization of

acquired above- and below-market leases and tenant-funded and tenant-built landlord improvements.

Furthermore, we believe net operating income is useful to investors as a performance measure of our consolidated properties

because, when compared across periods, net operating income reflects trends in occupancy rates, rental rates, and operating costs,

which provide a perspective not immediately apparent from net income or loss. Net operating income can be used to measure the initial

stabilized yields of our properties by calculating net operating income generated by a property divided by our investment in the property.

Net operating income excludes certain components from net income in order to provide results that are more closely related to the

results of operations of our properties. For example, interest expense is not necessarily linked to the operating performance of a real

estate asset and is often incurred at the corporate level rather than at the property level. In addition, depreciation and amortization,

because of historical cost accounting and useful life estimates, may distort comparability of operating performance at the property level.

Impairments of real estate have been excluded in deriving net operating income because we do not consider impairments of real estate

to be property-level operating expenses. Impairments of real estate relate to changes in the values of our assets and do not reflect the

current operating performance with respect to related revenues or expenses. Our impairments of real estate represent the write-down in

the value of the assets to the estimated fair value less cost to sell. These impairments result from investing decisions or a deterioration

in market conditions. We also exclude realized and unrealized investment gain or loss, which results from investment decisions that

occur at the corporate level related to non-real estate investments in publicly traded companies and certain privately held entities.

Therefore, we do not consider these activities to be an indication of operating performance of our real estate assets at the property

level. Our calculation of net operating income also excludes charges incurred from changes in certain financing decisions, such as

losses on early extinguishment of debt and changes in provision for expected credit losses on financial instruments, as these charges

often relate to corporate strategy. Property operating expenses included in determining net operating income primarily consist of costs

that are related to our operating properties, such as utilities, repairs, and maintenance; rental expense related to ground leases;

contracted services, such as janitorial, engineering, and landscaping; property taxes and insurance; and property-level salaries.

General and administrative expenses consist primarily of accounting and corporate compensation, corporate insurance, professional

fees, rent, and supplies that are incurred as part of corporate office management. We calculate operating margin as net operating

income divided by total revenues.

We believe that, to facilitate investors’ clear understanding of our operating results, net operating income should be examined

in conjunction with net income or loss as presented in our consolidated statements of operations. Net operating income should not be

considered as an alternative to net income or loss as an indication of our performance, nor as an alternative to cash flows as a measure

of our liquidity or our ability to make distributions.

We are not able to forecast the net income of future periods without unreasonable effort, and therefore do not provide a

reconciliation for net operating income on a forward-looking basis. This is due to the inherent difficulty of forecasting the timing and/or

amount of items that depend on market conditions outside of our control, including the timing of dispositions, capital events, and

financing decisions, as well as components such as gain on sales of real estate, unrealized gains or losses on non-real estate

investments, impairments of real estate, impairments of non-real estate investments, and changes in provision for expected credit

losses on financial instruments. Our attempt to predict these amounts may produce significant but inaccurate estimates, which would

potentially be misleading for our investors.

Operating statistics

We present certain operating statistics related to our properties, including number of properties, RSF, occupancy percentage,

leasing activity, and contractual lease expirations as of the end of the period. We believe these measures are useful to investors

because they facilitate an understanding of certain trends for our properties. We compute the number of properties, RSF, occupancy

percentage, leasing activity, and contractual lease expirations at 100%, excluding RSF at properties classified as held for sale, for all

properties in which we have an investment, including properties owned by our consolidated and unconsolidated real estate joint

ventures. For operating metrics based on annual rental revenue, refer to “Annual rental revenue” in this section.

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Same property comparisons

As a result of changes within our total property portfolio during the comparative periods presented, including changes from

assets acquired or sold, properties placed into development or redevelopment, and development or redevelopment properties recently

placed into service, the consolidated total income from rentals, as well as rental operating expenses in our operating results, can show

significant changes from period to period. In order to supplement an evaluation of our results of operations over a given quarterly or

annual period, we analyze the operating performance for all consolidated properties that were fully operating for the entirety of the

comparative periods presented, referred to as same properties. We separately present quarterly and year-to-date same property results

to align with the interim financial information required by the SEC in our management’s discussion and analysis of our financial

condition and results of operations. These same properties are analyzed separately from properties acquired subsequent to the first day

in the earliest comparable quarterly or year-to-date period presented, properties that underwent development or redevelopment at any

time during the comparative periods, unconsolidated real estate joint ventures, properties classified as held for sale, and corporate

entities (legal entities performing general and administrative functions), which are excluded from same property results. Additionally,

termination fees, if any, are excluded from the results of same properties. Refer to “Same properties” in Item 2 for additional information.

Stabilized occupancy date

The stabilized occupancy date represents the estimated date on which a development or redevelopment project is expected to

reach occupancy of 95% or greater.

Tenant collections

Tenant collections represent the percentage of recognized rental income billed during the respective quarter that has been

collected as of the date of this report. Rental income from tenants for whom collection is considered not probable is recognized only

upon receipt of cash and, accordingly, is included in this calculation only to the extent recognized and collected.

Tenant recoveries

Tenant recoveries represent revenues comprising reimbursement of real estate taxes, insurance, utilities, repairs and

maintenance, common area expenses, and other operating expenses and earned in the period during which the applicable expenses

are incurred and the tenant’s obligation to reimburse us arises.

We classify rental revenues and tenant recoveries generated through the leasing of real estate assets within revenues in

income from rentals in our consolidated statements of operations. We provide investors with a separate presentation of rental revenues

and tenant recoveries in “Results of operations” in Item 2 because we believe it promotes investors’ understanding of our operating

results. We believe that the presentation of tenant recoveries is useful to investors as a supplemental measure of our ability to recover

operating expenses under our triple net leases, including recoveries of utilities, repairs and maintenance, insurance, property taxes,

common area expenses, and other operating expenses, and of our ability to mitigate the effect to net income for any significant

variability to components of our operating expenses.

The following table reconciles income from rentals to tenant recoveries for the three months ended March 31, 2026 and 2025

(in thousands):

Three Months Ended March 31,
2026 2025
Income from rentals $653,013 $743,175
Rental revenues (474,786) (552,112)
Tenant recoveries $178,227 $191,063

Total equity capitalization

Total equity capitalization is equal to the outstanding shares of common stock multiplied by the closing price on the last trading

day at the end of each period presented.

Total market capitalization

Total market capitalization is equal to the sum of total equity capitalization and total debt.

Unencumbered net operating income as a percentage of total net operating income

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Unencumbered net operating income as a percentage of total net operating income is a non-GAAP financial measure that we

believe is useful to investors as a performance measure of the results of operations of our unencumbered real estate assets as it

reflects those income and expense items that are incurred at the unencumbered property level. Unencumbered net operating income is

derived from assets classified in continuing operations, which are not subject to any mortgage, deed of trust, lien, or other security

interest, as of the period for which income is presented.

The following table summarizes unencumbered net operating income as a percentage of total net operating income for the

three months ended March 31, 2026 and 2025 (dollars in thousands):

Three Months Ended March 31,
2026 2025
Unencumbered net operating income $446,880 $530,691
Encumbered net operating income 1,072
Total net operating income $446,880 $531,763
Unencumbered net operating income as a percentage of total net operating income 100.0% 99.8%

Weighted-average shares of common stock outstanding – diluted

From time to time, we enter into capital market transactions, including forward equity sales agreements (“Forward

Agreements”), to fund acquisitions, to fund construction of our development and redevelopment projects, and for general working

capital purposes. While the Forward Agreements are outstanding, we are required to consider the potential dilutive effect of our Forward

Agreements under the treasury stock method. Under this method, we also include the dilutive effect of unvested restricted stock awards

(“RSAs”) with forfeitable dividends in the calculation of diluted shares. Refer to Note 13 – “Earnings per share” and Note 14 –

“Stockholders’ equity” to our unaudited consolidated financial statements in Item 1 for additional information.

The weighted-average shares of common stock outstanding used in calculating EPS – diluted, funds from operations per

share – diluted, and funds from operations per share – diluted, as adjusted, for the three months ended March 31, 2026 and 2025 are

calculated as follows. Also shown are the weighted-average unvested RSAs with nonforfeitable dividends used in calculating the

amounts allocable to these awards pursuant to the two-class method for each of the respective periods presented below (in thousands):

Three Months Ended March 31,
2026 2025
Basic shares for earnings per share 170,598 170,522
Unvested RSAs with forfeitable dividends 269
Diluted shares for earnings per share 170,867 170,522
Basic shares for funds from operations per share and funds from operations per share, as<br><br>adjusted 170,598 170,522
Unvested RSAs with forfeitable dividends 269 77
Diluted shares for funds from operations per share and funds from operations per share, as<br><br>adjusted 170,867 170,599
Weighted-average unvested RSAs with nonforfeitable dividends used in the allocations of net<br><br>income, funds from operations, and funds from operations, as adjusted 1,340 2,053

116

ITEM 3. QUANTITATIVE AND QUALITATIVE DISCLOSURES ABOUT MARKET RISK

Interest rate risk

The primary market risk to which we believe we may be exposed is interest rate risk, which may result from many factors,

including government monetary and tax policies, domestic and international economic and political considerations, and other factors

that are beyond our control.

In order to modify and manage the interest rate characteristics of our outstanding debt and to limit the effects of interest rate

risks on our operations, we may utilize a variety of financial instruments, including interest rate hedge agreements, caps, floors, and

other interest rate exchange contracts. The use of these types of instruments to hedge a portion of our exposure to changes in interest

rates may carry additional risks, such as counterparty credit risk and the legal enforceability of hedge agreements. As of March 31,

2026, we did not have any outstanding interest rate hedge agreements.

Our future earnings and fair values relating to our outstanding debt are primarily dependent upon prevalent market rates of

interest. The following tables illustrate the effect of a 1% change in interest rates, assuming a zero percent interest rate floor, on our

fixed- and variable-rate debt as of March 31, 2026 (in thousands):

As of
March 31, 2026 December 31, 2025
Annualized effect on future earnings due to variable-rate debt:
Rate increase of 1% $(4,706) $(1,259)
Rate decrease of 1% $4,706 $1,259
Effect on fair value of total consolidated debt:
Rate increase of 1% $(658,167) $(746,058)
Rate decrease of 1% $742,060 $852,698

These amounts are determined by considering the effect of the hypothetical interest rates on our borrowings as of March 31,

2026 and December 31, 2025. These analyses do not consider the effects of the reduced level of overall economic activity that could

exist in such an environment. Furthermore, in the event of a change of such magnitude, we would consider taking actions to further

mitigate our exposure to the change. Because of the uncertainty of the specific actions that would be taken and their possible effects,

the sensitivity analyses assume no changes in our capital structure.

Equity price risk

We have exposure to equity price market risk because we hold equity investments in publicly traded companies and privately

held entities. All of our investments in actively traded public companies are reflected in our consolidated balance sheets at fair value.

Our investments in privately held entities that report NAV per share are measured at fair value using NAV as a practical expedient to fair

value. Our equity investments in privately held entities that do not report NAV per share are measured at cost less impairments,

adjusted for observable price changes during the period. Changes in fair value of public investments, changes in NAV per share

reported by privately held entities, and observable price changes of privately held entities that do not report NAV per share are

classified as investment income (loss) in our consolidated statements of operations. There is no assurance that future declines in value

will not have a material adverse effect on our future results of operations. The following table illustrates the effect that a 10% change in

the value of our equity investments would have on earnings as of March 31, 2026 and December 31, 2025 (in thousands):

As of
March 31, 2026 December 31, 2025
Equity price risk:
Fair value increase of 10% $116,854 $114,387
Fair value decrease of 10% $(116,854) $(114,387)

117

Foreign currency exchange rate risk

We have exposure to foreign currency exchange rate risk related to our operations in Canada. The functional currency of our

Canadian subsidiaries is the Canadian dollar. Gains or losses resulting from the translation of these subsidiaries’ balance sheets and

statements of operations are classified in accumulated other comprehensive income (loss) as a separate component of total equity and

are excluded from net income (loss). Gains or losses will be reflected in our consolidated statements of operations when there is a sale

or partial sale of our investment in these operations or upon a complete or substantially complete liquidation of the investment. The

following tables illustrate the effect that a 10% change in Canadian dollar exchange rates relative to the USD would have on our

potential future earnings and on the fair value of our net investment in Canadian subsidiaries, based on our current operating assets

outside the U.S. as of March 31, 2026 and December 31, 2025 (in thousands):

As of
March 31, 2026 December 31, 2025
Effect on potential future earnings due to foreign currency exchange rate:
Rate increase of 10% $290 $182
Rate decrease of 10% $(290) $(182)
Effect on the fair value of net investment in foreign subsidiaries due to foreign currency<br><br>exchange rate:
Rate increase of 10% $35,164 $35,306
Rate decrease of 10% $(35,164) $(35,306)
Change in the fair value of cross-currency swap agreements designated as a net<br><br>investment hedge(1):
Rate increase of 10% (USD weakening) $24,400 $(24,600)
Rate decrease of 10% (USD strengthening) $(24,400) $24,600

(1)Refer to Note 11 – “Hedge agreements” to our unaudited consolidated financial statements for additional information.

The sensitivity analyses assume a parallel shift of all foreign currency exchange rates with respect to the U.S. dollar; however,

foreign currency exchange rates do not typically move in such a manner, and actual results may differ materially.

Our exposure to market risk elements for the three months ended March 31, 2026 was consistent with the risk elements

presented above, including the effects of changes in interest rates, equity prices, and foreign currency exchange rates.

ITEM 4. CONTROLS AND PROCEDURES

Evaluation of disclosure controls and procedures

As of March 31, 2026, we had performed an evaluation, under the supervision of our principal executive officers and principal

financial officer, of the effectiveness of the design and operation of our disclosure controls and procedures. These controls and

procedures have been designed to ensure that information required for disclosure is recorded, processed, summarized, and reported

within the requisite time periods. Based on our evaluation, the principal executive officers and principal financial officer concluded that

our disclosure controls and procedures were effective as of March 31, 2026.

Changes in internal control over financial reporting

There has not been any change in our internal control over financial reporting during the three months ended March 31, 2026

that has materially affected, or is reasonably likely to materially affect, our internal control over financial reporting.

118

PART II – OTHER INFORMATION

ITEM 1. LEGAL PROCEEDINGS

Stockholder Matters

On November 25, 2025, a securities class action was filed against the Company and certain of its officers and directors in the

United States District Court for the Central District of California. On April 15, 2026, the lead plaintiffs filed an amended complaint

alleging violations of Sections 10(b) and 20(a) of the Securities Exchange Act of 1934 and Rule 10b-5 thereunder, based on alleged

material misrepresentations and omissions related to the Company’s business performance and real estate impairment charges

(captioned Hern v. Alexandria Real Estate Equities, Inc., et al.). The amended complaint seeks damages and other relief on behalf of

investors who acquired the Company’s securities between January 30, 2024 and December 5, 2025. The Company does not believe

the amended complaint states any meritorious claims and intends to defend this case vigorously.

On February 3, 2026 and March 25, 2026, stockholder derivative actions were filed against certain officers and directors of the

Company, with the Company named as a nominal defendant, in the United States District Court for the District of Maryland (captioned

De Albuquerque Torres v. Alexandria Real Estate Equities, Inc., et al.) and the United States District Court for the Central District of

California (captioned Tabone v. Moglia, et al.). The derivative complaints allege violations of federal securities laws and breaches of

fiduciary duty based on allegations similar to those in the securities class action and seek damages and other relief on behalf of the

Company. On April 8, 2026, the Maryland derivative action was stayed pending resolution of any motion to dismiss in the securities

class action. The Company does not believe the derivative complaints state any meritorious claims and intends to defend these cases

vigorously.

At this time, we cannot predict the outcome of these matters or reasonably estimate the amount or range of any possible loss,

if any, and therefore we have not recorded an accrual related to these

matters.

Option Parcel Development at Alexandria Center® for Life Science – New York City Campus

Refer to “Other” in Note 3 – “Investments in real estate” to our unaudited consolidated financial statements for information

regarding litigation involving our subsidiary in connection with an option and ground lease for a development parcel at the Alexandria

Center® for Life Science – New York City campus.

ITEM 1A. RISK FACTORS

In addition to the information set forth in this quarterly report on Form 10-Q, one should also carefully review and consider the

information contained in the other reports and periodic filings that we make with the SEC, including, without limitation, the information

contained under the caption “Item 1A. Risk factors” in our annual report on Form 10-K for the year ended December 31, 2025. Those

risk factors could materially affect our business, financial condition, and results of operations. The risks that we describe in our public

filings are not the only risks that we face. Additional risks and uncertainties not currently known to us, or that we presently deem to be

immaterial, also may materially adversely affect our business, financial condition, and results of operations.

There have been no material changes in our risk factors from those disclosed under the caption “Item 1A. Risk factors” in our

annual report on Form 10-K for the year ended December 31, 2025.

ITEM 2. UNREGISTERED SALES OF EQUITY SECURITIES AND USE OF PROCEEDS

Purchases of equity securities

On December 8, 2025, we announced that our Board of Directors authorized a new share repurchase program that allows the

repurchase of shares with an aggregate value of up to $500.0 million through December 31, 2026 in the open market, through privately

negotiated transactions, or otherwise, in accordance with all applicable securities laws and regulations, including Rule 10b-18 of the

Exchange Act. This new program replaced our prior stock repurchase program. As of the date of this report, no repurchases have been

made under the new program and $500.0 million remains available for future share repurchases.

ITEM 5. OTHER INFORMATION

Disclosure of 10b5-1 plans

During the three months ended March 31, 2026, none of our officers or directors adopted or terminated any contract,

instruction, or written plan for the purchase or sale of our securities that was intended to satisfy the affirmative defense conditions of

Rule 10b5-1(c) or any “non-Rule 10b5-1 trading arrangement.”

119

ITEM 6. EXHIBITS

Exhibit<br><br>Number Exhibit Title Incorporated by<br><br>Reference to: Date Filed
3.1* Articles of Amendment and Restatement of the Company, dated May 21, 1997 Form 10-Q August 14, 1997
3.2* Certificate of Correction of the Company, dated June 20, 1997 Form 10-Q August 14, 1997
3.3* Articles of Amendment of the Company, effective as of May 10, 2017 Form 8-K May 12, 2017
3.4* Articles of Amendment of the Company, effective as of May 18, 2022 Form 8-K May 19, 2022
3.5* Articles Supplementary, dated June 9, 1999, relating to the 9.50% Series A<br><br>Cumulative Redeemable Preferred Stock Form 10-Q August 13, 1999
3.6* Articles Supplementary, dated February 10, 2000, relating to the election to be<br><br>subject to Subtitle 8 of Title 3 of the Maryland General Corporation Law Form 8-K February 10, 2000
3.7* Articles Supplementary, dated February 10, 2000, relating to the Series A<br><br>Junior Participating Preferred Stock Form 8-K February 10, 2000
3.8* Articles Supplementary, dated January 18, 2002, relating to the 9.10%<br><br>Series B Cumulative Redeemable Preferred Stock Form 8-A January 18, 2002
3.9* Articles Supplementary, dated June 22, 2004, relating to the 8.375% Series C<br><br>Cumulative Redeemable Preferred Stock Form 8-A June 28, 2004
3.10* Articles Supplementary, dated March 25, 2008, relating to the 7.00% Series D<br><br>Cumulative Convertible Preferred Stock Form 8-K March 25, 2008
3.11* Articles Supplementary, dated March 12, 2012, relating to the 6.45% Series E<br><br>Cumulative Redeemable Preferred Stock Form 8-K March 14, 2012
3.12* Articles Supplementary, effective as of May 10, 2017, relating to Reclassified<br><br>Preferred Stock Form 8-K May 12, 2017
3.13* Articles Supplementary, effective as of March 31, 2026, relating to Subtitle 8 of<br><br>Title 3 of the Maryland General Corporation Law Form 8-K March 31, 2026
3.14* Amended and Restated Bylaws of the Company (Amended December 6,<br><br>2024) Form 8-K December 9, 2024
4.1* Indenture, dated as of February 13, 2025, among Alexandria Real Estate<br><br>Equities, Inc., as issuer, Alexandria Real Estate Equities, L.P., as Guarantor,<br><br>and U.S. Bank Trust Company, National Association, as Trustee Form 8-K February 13, 2025
4.2* Supplemental Indenture No. 2, dated as of February 25, 2026, by and among<br><br>Alexandria Real Estate Equities, Inc., as Issuer, Alexandria Real Estate<br><br>Equities, L.P., as Guarantor, and U.S. Bank Trust Company, National<br><br>Association, as trustee Form 8-K February 25, 2026
4.3* Form of 5.25% Senior Note due 2036 (included in Exhibit 4.2 above) Form 8-K February 25, 2026
22.1 List of Guarantor Subsidiaries of the Company N/A Filed herewith
31.1 Certification of Principal Executive Officer Pursuant to Section 302 of the<br><br>Sarbanes-Oxley Act of 2002 N/A Filed herewith
31.2 Certification of Principal Executive Officer Pursuant to Section 302 of the<br><br>Sarbanes-Oxley Act of 2002 N/A Filed herewith
31.3 Certification of Principal Financial Officer Pursuant to Section 302 of the<br><br>Sarbanes-Oxley Act of 2002 N/A Filed herewith
32.0 Certification of Principal Executive Officers and Principal Financial Officer<br><br>Pursuant to 18 U.S.C. Section 1350, as Adopted Pursuant to Section 906 of<br><br>the Sarbanes-Oxley Act of 2002 N/A Filed herewith
101.1 The following materials from the Company’s quarterly report on Form 10-Q for<br><br>the quarterly period ended March 31, 2026, formatted in iXBRL (Inline<br><br>eXtensible Business Reporting Language): (i) Consolidated Balance Sheets<br><br>as of March 31, 2026 and December 31, 2025 (unaudited), (ii) Consolidated<br><br>Statements of Operations for the three months ended March 31, 2026 and<br><br>2025 (unaudited), (iii) Consolidated Statements of Comprehensive Income for<br><br>the three months ended March 31, 2026 and 2025 (unaudited), (iv)<br><br>Consolidated Statements of Changes in Stockholders’ Equity and<br><br>Noncontrolling Interests for the three months ended March 31, 2026 and 2025<br><br>(unaudited), (v) Consolidated Statements of Cash Flows for the three months<br><br>ended March 31, 2026 and 2025 (unaudited), and (vi) Notes to Consolidated<br><br>Financial Statements (unaudited) N/A Filed herewith
104 Cover Page Interactive Data File – the cover page from this Quarterly Report<br><br>on Form 10-Q for the quarter ended March 31, 2026 is formatted in Inline<br><br>XBRL and contained in Exhibit 101.1 N/A Filed herewith

(*) Incorporated by reference.

(1) Management contract or compensatory arrangement.

120

SIGNATURES

Pursuant to the requirements of the Securities Exchange Act of 1934, the registrant has duly caused this report to be signed

on its behalf by the undersigned, thereunto duly authorized, on April 27, 2026.

ALEXANDRIA REAL ESTATE EQUITIES, INC.
/s/ Joel S. Marcus
Joel S. Marcus<br><br>Executive Chairman<br><br>(Principal Executive Officer)
/s/ Peter M. Moglia
Peter M. Moglia<br><br>Chief Executive Officer and Chief Investment Officer<br><br>(Principal Executive Officer)
/s/ Marc E. Binda
Marc E. Binda<br><br>Chief Financial Officer and Treasurer<br><br>(Principal Financial Officer)

Document

EXHIBIT 22.1

List of Guarantor Subsidiaries of Alexandria Real Estate Equities, Inc.

The following subsidiary was, as of March 31, 2026, a guarantor of the registrant's 3.80% Senior Notes due 2026, 3.95% Senior Notes due 2027, 3.95% Senior Notes due 2028, 4.50% Senior Notes due 2029, 2.75% Senior Notes due 2029, 4.70% Senior Notes due 2030, 4.90% Senior Notes due 2030, 3.375% Senior Notes due 2031, 2.00% Senior Notes due 2032, 1.875% Senior Notes due 2033, 2.95% Senior Notes due 2034, 4.75% Senior Notes due 2035, 5.50% Senior Notes due 2035, 5.25% Senior Notes due 2036, 5.25% Senior Notes due 2036, 4.85% Senior Notes due 2049, 4.00% Senior Notes due 2050, 3.00% Senior Notes due 2051, 3.55% Senior Notes due 2052, 5.15% Senior Notes due 2053, and 5.625% Senior Notes due 2054.

Name of Subsidiary Jurisdiction of Organization
Alexandria Real Estate Equities, L.P. Delaware

Document

EXHIBIT 31.1

CERTIFICATION OF PRINCIPAL EXECUTIVE OFFICER

PURSUANT TO

SECTION 302 OF THE SARBANES-OXLEY ACT OF 2002

I, Joel S. Marcus, certify that:

1.    I have reviewed this Quarterly Report on Form 10-Q of Alexandria Real Estate Equities, Inc.;

2.    Based on my knowledge, this report does not contain any untrue statement of a material fact or omit to state a material fact necessary to make the statements made, in light of the circumstances under which such statements were made, not misleading with respect to the period covered by this report;

3.    Based on my knowledge, the financial statements, and other financial information included in this report, fairly present in all material respects the financial condition, results of operations, and cash flows of the registrant as of, and for, the periods presented in this report;

4.    The registrant’s other certifying officers and I are responsible for establishing and maintaining disclosure controls and procedures (as defined in Exchange Act Rules 13a-15(e) and 15d-15(e)) and internal control over financial reporting (as defined in Exchange Act Rules 13a-15(f) and 15d-15(f)) for the registrant and have:

a.    Designed such disclosure controls and procedures, or caused such disclosure controls and procedures to be designed under our supervision, to ensure that material information relating to the registrant, including its consolidated subsidiaries, is made known to us by others within those entities, particularly during the period in which this report is being prepared;

b.    Designed such internal control over financial reporting, or caused such internal control over financial reporting to be designed under our supervision, to provide reasonable assurance regarding the reliability of financial reporting and the preparation of financial statements for external purposes in accordance with generally accepted accounting principles;

c.    Evaluated the effectiveness of the registrant’s disclosure controls and procedures and presented in this report our conclusions about the effectiveness of the disclosure controls and procedures, as of the end of the period covered by this report based on such evaluation; and

d.    Disclosed in this report any change in the registrant’s internal control over financial reporting that occurred during the registrant’s most recent fiscal quarter (the registrant’s fourth fiscal quarter in the case of an annual report) that has materially affected, or is reasonably likely to materially affect, the registrant’s internal control over financial reporting; and

5.    The registrant’s other certifying officers and I have disclosed, based on our most recent evaluation of internal control over financial reporting, to the registrant’s auditors and the audit committee of the registrant’s board of directors (or persons performing the equivalent functions):

a.    All significant deficiencies and material weaknesses in the design or operation of internal control over financial reporting which are reasonably likely to adversely affect the registrant’s ability to record, process, summarize, and report financial information; and

b.    Any fraud, whether or not material, that involves management or other employees who have a significant role in the registrant’s internal control over financial reporting.

Date: April 27, 2026

/s/ Joel S. Marcus
Joel S. Marcus
Executive Chairman

Document

EXHIBIT 31.2

CERTIFICATION OF PRINCIPAL EXECUTIVE OFFICER

PURSUANT TO

SECTION 302 OF THE SARBANES-OXLEY ACT OF 2002

I, Peter M. Moglia, certify that:

1.    I have reviewed this Quarterly Report on Form 10-Q of Alexandria Real Estate Equities, Inc.;

2.    Based on my knowledge, this report does not contain any untrue statement of a material fact or omit to state a material fact necessary to make the statements made, in light of the circumstances under which such statements were made, not misleading with respect to the period covered by this report;

3.    Based on my knowledge, the financial statements, and other financial information included in this report, fairly present in all material respects the financial condition, results of operations, and cash flows of the registrant as of, and for, the periods presented in this report;

4.    The registrant’s other certifying officers and I are responsible for establishing and maintaining disclosure controls and procedures (as defined in Exchange Act Rules 13a-15(e) and 15d-15(e)) and internal control over financial reporting (as defined in Exchange Act Rules 13a-15(f) and 15d-15(f)) for the registrant and have:

a.    Designed such disclosure controls and procedures, or caused such disclosure controls and procedures to be designed under our supervision, to ensure that material information relating to the registrant, including its consolidated subsidiaries, is made known to us by others within those entities, particularly during the period in which this report is being prepared;

b.    Designed such internal control over financial reporting, or caused such internal control over financial reporting to be designed under our supervision, to provide reasonable assurance regarding the reliability of financial reporting and the preparation of financial statements for external purposes in accordance with generally accepted accounting principles;

c.    Evaluated the effectiveness of the registrant’s disclosure controls and procedures and presented in this report our conclusions about the effectiveness of the disclosure controls and procedures, as of the end of the period covered by this report based on such evaluation; and

d.    Disclosed in this report any change in the registrant’s internal control over financial reporting that occurred during the registrant’s most recent fiscal quarter (the registrant’s fourth fiscal quarter in the case of an annual report) that has materially affected, or is reasonably likely to materially affect, the registrant’s internal control over financial reporting; and

5.    The registrant’s other certifying officers and I have disclosed, based on our most recent evaluation of internal control over financial reporting, to the registrant’s auditors and the audit committee of the registrant’s board of directors (or persons performing the equivalent functions):

a.    All significant deficiencies and material weaknesses in the design or operation of internal control over financial reporting which are reasonably likely to adversely affect the registrant’s ability to record, process, summarize, and report financial information; and

b.    Any fraud, whether or not material, that involves management or other employees who have a significant role in the registrant’s internal control over financial reporting.

Date: April 27, 2026

/s/ Peter M. Moglia
Peter M. Moglia
Chief Executive Officer and Chief Investment Officer

Document

EXHIBIT 31.3

CERTIFICATION OF PRINCIPAL FINANCIAL OFFICER

PURSUANT TO

SECTION 302 OF THE SARBANES-OXLEY ACT OF 2002

I, Marc E. Binda, certify that:

1.    I have reviewed this Quarterly Report on Form 10-Q of Alexandria Real Estate Equities, Inc.;

2.    Based on my knowledge, this report does not contain any untrue statement of a material fact or omit to state a material fact necessary to make the statements made, in light of the circumstances under which such statements were made, not misleading with respect to the period covered by this report;

3.    Based on my knowledge, the financial statements, and other financial information included in this report, fairly present in all material respects the financial condition, results of operations, and cash flows of the registrant as of, and for, the periods presented in this report;

4.    The registrant’s other certifying officers and I are responsible for establishing and maintaining disclosure controls and procedures (as defined in Exchange Act Rules 13a-15(e) and 15d-15(e)) and internal control over financial reporting (as defined in Exchange Act Rules 13a-15(f) and 15d-15(f)) for the registrant and have:

a.    Designed such disclosure controls and procedures, or caused such disclosure controls and procedures to be designed under our supervision, to ensure that material information relating to the registrant, including its consolidated subsidiaries, is made known to us by others within those entities, particularly during the period in which this report is being prepared;

b.    Designed such internal control over financial reporting, or caused such internal control over financial reporting to be designed under our supervision, to provide reasonable assurance regarding the reliability of financial reporting and the preparation of financial statements for external purposes in accordance with generally accepted accounting principles;

c.    Evaluated the effectiveness of the registrant’s disclosure controls and procedures and presented in this report our conclusions about the effectiveness of the disclosure controls and procedures, as of the end of the period covered by this report based on such evaluation; and

d.    Disclosed in this report any change in the registrant’s internal control over financial reporting that occurred during the registrant’s most recent fiscal quarter (the registrant’s fourth fiscal quarter in the case of an annual report) that has materially affected, or is reasonably likely to materially affect, the registrant’s internal control over financial reporting; and

5.    The registrant’s other certifying officers and I have disclosed, based on our most recent evaluation of internal control over financial reporting, to the registrant’s auditors and the audit committee of the registrant’s board of directors (or persons performing the equivalent functions):

a.    All significant deficiencies and material weaknesses in the design or operation of internal control over financial reporting which are reasonably likely to adversely affect the registrant’s ability to record, process, summarize, and report financial information; and

b.    Any fraud, whether or not material, that involves management or other employees who have a significant role in the registrant’s internal control over financial reporting.

Date: April 27, 2026

/s/ Marc E. Binda
Marc E. Binda
Chief Financial Officer and Treasurer

Document

EXHIBIT 32.0

CERTIFICATION OF PRINCIPAL EXECUTIVE OFFICERS AND PRINCIPAL FINANCIAL OFFICER

PURSUANT TO

18 U.S.C. SECTION 1350.

AS ADOPTED PURSUANT TO

SECTION 906 OF THE SARBANES-OXLEY ACT OF 2002

I, Joel S. Marcus, certify, pursuant to 18 U.S.C. Section 1350, as adopted pursuant to Section 906 of the Sarbanes-Oxley Act of 2002, that the Quarterly Report on Form 10-Q of Alexandria Real Estate Equities, Inc. for the quarter ended March 31, 2026, fully complies with the requirements of Section 13(a) or 15(d) of the Securities Exchange Act of 1934, as amended, and that the information contained in such report fairly presents, in all material respects, the financial condition and results of operations of Alexandria Real Estate Equities, Inc.

Date: April 27, 2026

/s/ Joel S. Marcus
Joel S. Marcus
Executive Chairman

I, Peter M. Moglia, certify, pursuant to 18 U.S.C. Section 1350, as adopted pursuant to Section 906 of the Sarbanes-Oxley Act of 2002, that the Quarterly Report on Form 10-Q of Alexandria Real Estate Equities, Inc. for the quarter ended March 31, 2026, fully complies with the requirements of Section 13(a) or 15(d) of the Securities Exchange Act of 1934, as amended, and that the information contained in such report fairly presents, in all material respects, the financial condition and results of operations of Alexandria Real Estate Equities, Inc.

Date: April 27, 2026

/s/ Peter M. Moglia
Peter M. Moglia
Chief Executive Officer and Chief Investment Officer

I, Marc E. Binda, certify, pursuant to 18 U.S.C. Section 1350, as adopted pursuant to Section 906 of the Sarbanes-Oxley Act of 2002, that the Quarterly Report on Form 10-Q of Alexandria Real Estate Equities, Inc. for the quarter ended March 31, 2026, fully complies with the requirements of Section 13(a) or 15(d) of the Securities Exchange Act of 1934, as amended, and that the information contained in such report fairly presents, in all material respects, the financial condition and results of operations of Alexandria Real Estate Equities, Inc.

Date: April 27, 2026

/s/ Marc E. Binda
Marc E. Binda
Chief Financial Officer and Treasurer