CSTL · Castle Biosciences Inc · Holdings

$34.51 +1.21 (+3.63%) At close · Oct 27 Q2 2026 earnings call transcript

Russell 2000 member Rebalance outlook locked

Market Cap

$1.05B

30.55M

Volume · Oct 27 295.28K Avg daily vol (3M) 397.34K

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Use institutional holdings through the 13F API or MCP → See every holder of CSTL: 13F, 13D and 13G, insiders and funds →

Institutional holdings are being reconciled with SEC filings. Holder totals and changes may be incomplete until reconciliation finishes.

The Sep 30, 2026 filing window is still open — showing the latest known positions. Funds that have not filed yet carry their prior-quarter positions; a fund that filed without this stock counts as sold out. Each fund's Change is measured against its own previous reported quarter, shown in the Reported column. Pick a completed quarter in the Quarter selector below to put every fund on one quarter and see the stock-wide activity breakdown.

Quarter:

Holders: 179

Total Value: $658,504,915

Total Shares: 27,373,315

Showing 1–100 of 179 holdings by value · page 1 of 2

Institutional Ownership Trend

27 quarters

Holder Concentration

Across 177 13F filers

Top Holder

12.0%

of institutional value

Top 5

36.3%

of institutional value

Top 10

51.5%

of institutional value

HHI

423

Broadly held

Who Holds CSTL

Last 8 quarters through Sep 30, 2026 (filing window open)

Each kind of 13F filer's share of the common shares 13F filers report. Holder type comes from regulatory records, never the filer's name; style, turnover and horizon are measured from each filer's own 13F book. The quarter selector does not change this card.

Jun 30, 2026, the newest closed quarter: 176 filers, like-for-like change -0.9%; actively managed books -2.1%; index-like books +1.8%.

Holder type	Share	Filers	Change	New	Exited
Registered adviser	50.1%	88	-2.6%	13	17
Exempt adviser	2.3%	2	+3.3%	0	1
Adviser	0.6%	3	0.0%	0	0
Bank	5.8%	15	+12.8%	1	0
Insurer	6.3%	2	-1.9%	1	0
Broker-dealer	0.1%	1	+42.3%	0	1
Benefit plan	0.2%	2	+30.1%	0	0
Non-US	0.7%	7	-70.2%	0	2
Holding company	18.4%	8	+25.2%	1	1
Unknown	15.5%	48	-15.7%	3	8

Style	Share	Filers	Change	New	Exited
Index-like	30.1%	24	+1.8%	0	0
Systematic-like	7.3%	16	-7.6%	3	5
Concentrated	2.4%	18	-7.4%	3	5
Diversified active	60.2%	118	-1.1%	13	20
Not measured	0.0%	0	—	0	0
Unknown	0.0%	0	—	0	0

Turnover	Share	Filers	Change	New	Exited
Low turnover	42.1%	87	-6.0%	4	13
Medium turnover	35.3%	46	+11.3%	6	7
High turnover	13.9%	28	-12.4%	8	9
Unknown	8.8%	15	+1.6%	1	1
Not measured	0.0%	0	—	0	0

Holding horizon	Share	Filers	Change	New	Exited
Short horizon	11.2%	24	-6.0%	5	6
Medium horizon	20.4%	26	+13.7%	5	5
Long horizon	61.8%	114	+1.0%	8	15
Unknown	6.6%	12	-33.6%	1	4
Not measured	0.0%	0	—	0	0

13F book size	Share	Filers	Change	New	Exited
Under $1B	0.8%	17	-26.3%	4	10
$1B to $10B	12.7%	46	-31.9%	4	10
$10B to $100B	33.3%	56	+26.5%	9	6
$100B or more	53.3%	57	-3.1%	2	4
Not measured	0.0%	0	—	0	0

Client base	Share	Filers	Change	New	Exited
Individuals and high net worth	2.5%	34	-34.8%	5	8
Pooled investment vehicles	22.2%	18	+6.5%	3	6
Registered funds and BDCs	17.7%	16	-6.7%	3	2
Pension and profit sharing plans	0.0%	1	0.0%	0	0
Insurance companies	0.1%	2	+3.3%	0	0
Governments and sovereign funds	2.2%	4	+44.8%	0	0
Other investment advisers	0.0%	2	-50.0%	0	0
Diversified	5.4%	11	-12.7%	2	1
Other	0.0%	0	—	0	0
Unknown	49.9%	88	+0.8%	6	13

Private funds	Share	Filers	Change	New	Exited
Hedge funds	27.7%	31	-5.7%	4	7
Private equity funds	0.1%	3	+5.1%	1	1
Venture capital funds	0.0%	1	-100.0%	0	1
Real estate funds	0.0%	3	+2053.5%	1	1
Securitized asset funds	0.2%	1	-1.0%	0	0
Other private funds	8.4%	11	+16.4%	1	1
Mixed private funds	0.1%	1	+7.2%	0	0
Private funds, type not stated	1.3%	1	+395776.1%	1	1
No private funds	14.6%	38	-4.2%	5	6
Unknown	47.6%	86	+0.7%	6	12

Location	Share	Filers	Change	New	Exited
United States	88.3%	142	-2.9%	15	23
Outside the US	11.7%	34	+17.4%	4	7

The Sep 30, 2026 filing window is still open, so its bar carries the Jun 30, 2026 positions of filers that have not filed yet.

How the mix is measured

Shares are 13F common shares restated to today's split basis; options and principal amounts are left out. The change compares only filers with a 13F in both quarters and keeps each filer in its bucket of the later quarter on both sides; a filer that continues under a new CIK is compared through its successor. Holder type comes from each filer's current regulatory profile; client base and private funds come from the Form ADV roster current at each quarter end (rosters from December 2019, so an earlier quarter reads Unknown). Style, turnover, horizon and book size are measured from each filer's 13F book in the newest measured quarter, and Not measured means none was. Actively managed books are those measured systematic-like, concentrated or diversified active.

Excluded 3 holders with no observed 2026-03-31 13F and 2 holders with no observed 2026-06-30 13F. Compared across CIK changes: JANUS HENDERSON GROUP PLC continues as Jupiter Topco LLC.

Conviction and Crowding

176 13F filers, quarter ended Jun 30, 2026

Measured across every 13F filer's common-share book once the quarter's filing window closed. Crowding counts filers with high turnover or that advise private funds on Form ADV.

Conviction holders

0.000% of filers hold it in their top 10

Mar 31, 2026: 1

Holders above 5%

median weight 0.001%

Mar 31, 2026: 0

Crowding

40.4%

of shares outstanding, 92nd percentile

Mar 31, 2026: 43.2%

Private-fund advisers

34.2%

of shares outstanding

Institutional ownership

90.4%

92.8% of the estimated free float

Top 10 holders

51.5%

of 13F shares, Herfindahl 0.042

Active holders

-2.1%

like-for-like share change of active filers

5% stakes on record

0 13D, 4 13G (4 institutional, 0 passive, 0 exempt)

How long holders have held it value-weighted 11.8 quarters

Under 1 year 26.0% 1 to 3 years 37.0% 3 years or more 37.0% Since coverage start 0.000%

Managers in the top fifth by current three-year FundScore alpha were net buyers of $0 (0 buying, 0 selling).

Estimated Holder Cost Basis

Quarter ended Jun 30, 2026, on that quarter's share basis

Estimated, never filed: each 13F buy is costed at its quarter's average traded price and each sale keeps the average cost, so this is what holders paid on average if they traded evenly through each quarter.

Estimated avg cost

$26.70

estimated per share

Mar 31, 2026: $27.91

Quarter-end price

$23.85

implied by 13F values

Estimated gain or loss

-10.7%

price against the estimated cost

Estimated at a loss

66.4%

of costed 13F value

Estimated cost covers 62.5% of the 13F shares; positions already held when a filer's 13F history starts, or bought in a quarter without a trusted average price, have no estimate.

Shared Holders

Companies CSTL's active holders also own, quarter ended Jun 30, 2026

#	Company	Shared holders	Similarity
1	ELUT Elutia Inc.	3	0.354
2	BDSX Biodesix Inc	3	0.316
3	LUNG Pulmonx Corp	3	0.316
4	CADL Candel Therapeutics, Inc.	3	0.277
5	SOPH SOPHiA GENETICS SA	3	0.277
6	FDMT 4D Molecular Therapeutics, Inc.	3	0.218
7	PSNL Personalis, Inc.	3	0.209
8	STOK Stoke Therapeutics, Inc.	3	0.204
9	URGN UroGen Pharma Ltd.	3	0.183
10	GPCR Structure Therapeutics Inc.	3	0.144

Shareholder Targets

13F filers that own its peers but not CSTL, quarter ended Jun 30, 2026

Found through its holder-overlap peers: ELUT, BDSX, LUNG, CADL, SOPH, FDMT, PSNL, STOK, URGN, GPCR, ORKA, ALKS, QURE, ZETA, ABVX, TWLO, AEIS, GH, UTHR, CIEN. 2,049 filers in all.

#	Filer	In peers	Of book	Largest peers
1	Avoro Capital Advisors LLC Registered adviser, Concentrated	$2,166,800,955 3 held	19.5%	UTHR, GPCR, QURE
2	WELLINGTON MANAGEMENT GROUP LLP Holding company, Diversified active	$1,830,297,818 14 held	0.320%	UTHR, GPCR, AEIS
3	Capital World Investors Adviser, Diversified active	$1,714,948,844 4 held	0.200%	CIEN, TWLO, ORKA
4	AMERIPRISE FINANCIAL INC Adviser, Diversified active	$1,607,856,962 15 held	0.330%	AEIS, GH, CIEN
5	Deep Track Capital, LP Adviser, Diversified active	$1,276,974,540 6 held	18.8%	GH, GPCR, ALKS
6	Pictet Asset Management Holding SA Unknown, Diversified active	$1,000,169,119 12 held	0.950%	TWLO, CIEN, UTHR
7	TCG Crossover Management, LLC Unknown, Diversified active	$800,603,958 1 held	19.2%	ABVX
8	Jericho Capital Asset Management L.P. Registered adviser, Diversified active	$727,825,057 2 held	6.02%	TWLO, CIEN
9	BAKER BROS. ADVISORS LP Registered adviser, Concentrated	$710,703,346 3 held	3.57%	ALKS, STOK, GPCR
10	Amundi Holding company, Index-like	$703,442,650 6 held	0.170%	CIEN, TWLO, UTHR

Fund Ownership of CSTL by Fund Type

Registered funds' Form N-PORT reports for the quarter ended Jun 30, 2026

Index funds and index ETFs hold 15.05% of shares outstanding.

Fund type	Of shares outstanding	Change	Shares	Funds
Index funds	3.28%	+0.41 pp	1,003,057	57
Index ETFs	11.77%	+1.14 pp	3,594,940	48
Active ETFs	2.00%	-0.14 pp	609,933	19
Active funds	12.98%	+0.67 pp	3,965,637	90
Unclassified	0.26%	+0.26 pp	79,259	8
All registered funds	30.29%	+2.35 pp	9,252,826	222

How fund types are measured

Each fund series counts once, with its latest Form N-PORT report dated inside the quarter; only long share positions count, restated to today's split basis. The fund type is the series' latest Form N-CEN answer on or before the quarter end (the index-fund box and the exchange-traded flag), never its name. Unclassified funds have no usable N-CEN; unit investment trusts are among them, so index ownership is a floor. Funds sit inside their advisers' 13F reports, so these shares overlap the 13F figures and are never added to them.

Holder type: Registered adviser 89 Exempt adviser 1 Adviser 3 Bank 15 Insurer 2 Broker-dealer 1 Benefit plan 2 Non-US 8 Holding company 8 Unknown 50

Showing 100 of 179

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

BlackRock, Inc.

$78,513,308

11.92%

3,291,963

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	105,233	$2,509,807	3.2%	Sole
BlackRock Investment Management (Australia) Ltd	67	$1,597	0.0%	Sole
BlackRock Asset Management Canada Ltd	23,688	$564,958	0.7%	Sole
BlackRock Investment Management, LLC	107,381	$2,561,036	3.3%	Sole
BLACKROCK ADVISORS LLC	283,647	$6,764,980	8.6%	Sole
BlackRock Fund Advisors	1,602,886	$38,228,831	48.7%	Sole
BlackRock Institutional Trust Company, N.A.	995,488	$23,742,388	30.2%	Sole
BlackRock Japan Co. Ltd	2,138	$50,991	0.1%	Sole
BlackRock Fund Managers Ltd.	5,948	$141,859	0.2%	Sole
BlackRock Investment Management (UK) Ltd.	23,912	$570,301	0.7%	Sole
BlackRock Asset Management Ireland Ltd	119,955	$2,860,926	3.6%	Sole
BlackRock (Luxembourg) S.A.	1,202	$28,667	0.0%	Sole
BlackRock Life Ltd	428	$10,207	0.0%	Sole
BlackRock Asset Management Schweiz AG	4,734	$112,905	0.1%	Sole
Aperio Group, LLC	15,256	$363,855	0.5%	Sole

RTW INVESTMENTS, LP

$66,263,289

10.06%

2,778,335

PRINCIPAL FINANCIAL GROUP INC

$41,180,554

6.25%

1,726,648

VANGUARD CAPITAL MANAGEMENT LLC

$29,251,953

4.44%

1,226,497

ACADIAN ASSET MANAGEMENT LLC

$21,910,231

3.33%

918,668

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Acadian Inc	918,668	$21,910,231	100.0%	Defined

Allianz Asset Management GmbH

$20,970,302

3.18%

879,258

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Allianz Global Investors GmbH	855,446	$20,402,387	97.3%	Defined
Allianz Global Investors Japan Co. Ltd.	753	$17,959	0.1%	Defined
Allianz Global Investors UK Ltd	22,748	$542,539	2.6%	Defined
Held directly	311	$7,417	0.0%	Defined

VANGUARD PORTFOLIO MANAGEMENT LLC

$20,052,722

3.05%

840,785

SUMMIT PARTNERS PUBLIC ASSET MANAGEMENT, LLC

$19,673,197

2.99%

824,872

STATE STREET CORP

$19,606,584

2.98%

822,079

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	822,079	$19,606,584	100.0%	Defined

GEODE CAPITAL MANAGEMENT, LLC

$18,982,501

2.88%

795,912

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	115,420	$2,752,767	14.5%	Defined
Held directly	680,492	$16,229,734	85.5%	Defined

MACKENZIE FINANCIAL CORP

$18,925,141

2.87%

793,507

DIMENSIONAL FUND ADVISORS LP

$18,867,375

2.87%

791,085

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	5,316	$126,786	0.7%	Defined
DFA Australia Ltd.	12,666	$302,084	1.6%	Defined
Dimensional Fund Advisors Pte. Ltd.	579	$13,809	0.1%	Defined
Dimensional Ireland Ltd	27,877	$664,866	3.5%	Defined
Held directly	744,647	$17,759,830	94.1%	Sole

Jefferies Financial Group Inc.

$15,440,275

2.34%

647,391

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jefferies Financial Services, Inc.	647,391	$15,440,275	100.0%	Defined

Qube Research & Technologies Ltd

$14,054,805

2.13%

589,300

AIGH Capital Management LLC

$13,000,658

1.97%

545,101

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Worth Venture Partners, LLC	132,755	$3,166,206	24.4%	Other
Held directly	412,346	$9,834,452	75.6%	Sole

WASATCH ADVISORS LP

$9,925,056

1.51%

279,343

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 1	279,343	$9,925,056	100.0%	Sole

RENAISSANCE TECHNOLOGIES LLC

$8,853,120

1.34%

371,200

GOLDMAN SACHS GROUP INC

$8,547,768

1.30%

358,397

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	105,493	$2,516,008	29.4%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	252,904	$6,031,760	70.6%	Defined

Point72 Asset Management, L.P.

$8,522,010

1.29%

357,317

KENNEDY CAPITAL MANAGEMENT LLC

$8,308,648

1.26%

348,371

Connor, Clark & Lunn Investment Management Ltd.

$7,911,688

1.20%

331,727

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 1	331,269	$7,900,765	99.9%	Sole
Held directly	458	$10,923	0.1%	Sole

MORGAN STANLEY

$7,862,960

1.19%

329,684

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	1,640	$39,114	0.5%	Defined
MORGAN STANLEY & CO. LLC	69,602	$1,660,007	21.1%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	244,933	$5,841,652	74.3%	Defined
MORGAN STANLEY SMITH BARNEY LLC	2,413	$57,549	0.7%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	11,096	$264,638	3.4%	Defined

Portolan Capital Management, LLC

$7,842,309

1.19%

328,818

PANAGORA ASSET MANAGEMENT INC

$7,179,804

1.09%

301,040

Allspring Global Investments Holdings, LLC

$6,975,409

1.06%

292,470

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Allspring Global Investments, LLC reporting on behalf of UBS ASSET MANAGEMENT AMERICAS INC	292,470	$6,975,409	100.0%	Defined

NORTHERN TRUST CORP

$6,801,016

1.03%

285,158

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	257,481	$6,140,921	90.3%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	22,967	$547,762	8.1%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	4,710	$112,333	1.7%	Other

CITADEL ADVISORS LLC

$6,328,668

0.96%

265,353

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	240,444	$5,734,589	90.6%	Defined
Held directly	24,909	$594,079	9.4%	Defined

KORNITZER CAPITAL MANAGEMENT INC /KS

$6,302,004

0.96%

264,235

SILVERCREST ASSET MANAGEMENT GROUP LLC

$6,107,865

0.93%

256,095

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$6,070,659

0.92%

254,535

Advisors Capital Management, LLC

$5,681,165

0.86%

238,204

NEW YORK STATE COMMON RETIREMENT FUND

$5,598,692

0.85%

234,746

PALISADE CAPITAL MANAGEMENT, LP

$5,531,554

0.84%

231,931

Nuveen, LLC

$5,526,663

0.84%

231,726

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	16,272	$388,086	7.0%	Defined
TEACHERS ADVISORS, LLC joint	215,454	$5,138,577	93.0%	Defined

BRIDGEWAY CAPITAL MANAGEMENT, LLC

$5,278,911

0.80%

221,338

Trexquant Investment LP

$4,915,747

0.75%

206,111

MILLENNIUM MANAGEMENT LLC

$4,898,050

0.74%

205,369

VANGUARD FIDUCIARY TRUST CO

$4,710,231

0.72%

197,494

AQR CAPITAL MANAGEMENT LLC

$3,972,621

0.60%

166,567

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Compound Planning, Inc.	6	$143	0.0%	Other
Arax Advisory Partners	524	$12,497	0.3%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	391	$9,325	0.2%	Other
Stokes Family Office, LLC	56	$1,335	0.0%	Other
Held directly	165,590	$3,949,321	99.4%	Defined

FIRST TRUST ADVISORS LP

$3,459,776

0.53%

145,064

Parkman Healthcare Partners LLC

$3,453,265

0.52%

144,791

Empowered Funds, LLC

$3,394,761

0.52%

142,338

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Bridgeway Capital Management, LLC	142,338	$3,394,761	100.0%	Other

HRT FINANCIAL LP

$2,763,094

0.42%

115,853

D. E. Shaw & Co., Inc.

$2,690,017

0.41%

112,789

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	112,789	$2,690,017	100.0%	Defined

Man Group plc

$2,593,949

0.39%

108,761

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Numeric Investors LLC	108,761	$2,593,949	100.0%	Defined

TWO SIGMA INVESTMENTS, LP

$2,393,919

0.36%

100,374

BANK OF AMERICA CORP /DE/

$2,355,067

0.36%

98,745

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	53,044	$1,265,099	53.7%	Defined
BOFA SECURITIES, INC.	45,701	$1,089,968	46.3%	Defined

Bank of New York Mellon Corp

$2,337,727

0.36%

98,018

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Investments Corporation	39,498	$942,027	40.3%	Defined
The Bank of New York Mellon	58,084	$1,385,303	59.3%	Defined
BNY Mellon, NA joint	434	$10,350	0.4%	Defined
Held directly	2	$47	0.0%	Defined

FRANKLIN RESOURCES INC

$2,330,931

0.35%

97,733

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FIDUCIARY TRUST INTERNATIONAL LLC	39	$930	0.0%	Defined
FRANKLIN ADVISERS INC	94,589	$2,255,947	96.8%	Defined
PUTNAM ADVISORY CO LLC	3,105	$74,054	3.2%	Defined

ARROWSTREET CAPITAL, LIMITED PARTNERSHIP

$2,211,753

0.34%

92,736

UBS Group AG

$2,131,425

0.32%

89,368

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	327	$7,798	0.4%	Defined
UBS AG	72,343	$1,725,380	80.9%	Defined
UBS Europe SE	16,698	$398,247	18.7%	Defined

JPMORGAN CHASE & CO

$2,022,573

0.31%

84,804

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	16,561	$394,979	19.5%	Defined
J.P. Morgan Investment Management Inc.	65,791	$1,569,115	77.6%	Defined
J.P. Morgan Securities LLC	49	$1,168	0.1%	Defined
J.P. Morgan Securities plc	2,403	$57,311	2.8%	Defined

SummitTX Capital, L.P.

$1,884,579

0.29%

79,018

SEI INVESTMENTS CO

$1,775,226

0.27%

74,433

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI INVESTMENTS MANAGEMENT CORP	166	$3,959	0.2%	Sole
SEI TRUST CO	74,267	$1,771,267	99.8%	Sole

ALGERT GLOBAL LLC

$1,680,900

0.26%

70,478

Orion Porfolio Solutions, LLC

$1,661,247

0.25%

69,654

BNP PARIBAS ARBITRAGE, SA

$1,659,268

0.25%

69,571

AMERICAN CENTURY COMPANIES INC

$1,560,100

0.24%

65,413

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AMERICAN CENTURY INVESTMENT MANAGEMENT INC	65,413	$1,560,100	100.0%	Defined

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$1,534,649

0.23%

64,346

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	404	$9,635	0.6%	Defined
UBS Asset Management Switzerland AG	49,504	$1,180,669	76.9%	Defined
UBS Asset Management (UK) Ltd.	14,438	$344,345	22.4%	Other

NORGES BANK

$1,433,385

0.22%

60,100

FMR LLC

$1,332,522

0.20%

55,871

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	55,517	$1,324,080	99.4%	Defined
STRATEGIC ADVISERS LLC	130	$3,100	0.2%	Defined
FIAM LLC	224	$5,342	0.4%	Defined

ENVESTNET ASSET MANAGEMENT INC

$1,315,494

0.20%

55,157

Invesco Ltd.

$1,309,076

0.20%

54,888

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	3,944	$94,063	7.2%	Defined
Invesco Trust Co joint	11,495	$274,155	20.9%	Defined
Invesco Capital Management LLC	39,449	$940,858	71.9%	Defined

Swiss National Bank

$1,280,745

0.19%

53,700

KBC Group NV

$1,280,572

0.19%

36,042

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
KBC Bank NV joint	36,042	$1,280,572	100.0%	Defined

PRICE T ROWE ASSOCIATES INC /MD/

$1,269,821

0.19%

53,242

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	53,242	$1,269,821	100.0%	Defined

JANE STREET GROUP, LLC

$1,068,312

0.16%

44,793

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	39,695	$946,725	88.6%	Defined
Jane Street Global Trading, LLC	5,098	$121,587	11.4%	Defined

XTX Topco Ltd

$952,163

0.14%

39,923

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	39,923	$952,163	100.0%	Defined

RHUMBLINE ADVISERS

$930,388

0.14%

39,010

HARBOR CAPITAL ADVISORS, INC.

$926,338

0.14%

26,072

CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM

$901,577

0.14%

37,802

Russell Investments Group, Ltd.

$755,543

0.11%

31,679

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC	15,218	$362,949	48.0%	Defined
Russell Investments Capital, LLC	3,906	$93,158	12.3%	Defined
Russell Investments Limited joint	12,555	$299,436	39.6%	Other

STRS OHIO

$727,425

0.11%

30,500

BARCLAYS PLC

$723,370

0.11%

30,330

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	29,071	$693,343	95.8%	Defined
BARCLAYS CAPITAL INC.	1,259	$30,027	4.2%	Defined

DEUTSCHE BANK AG\

$718,815

0.11%

30,139

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DWS Investments UK Ltd	30,139	$718,815	100.0%	Defined

WELLS FARGO & COMPANY/MN

$704,598

0.11%

29,543

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	25,094	$598,491	84.9%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	76	$1,812	0.3%	Defined
WELLS FARGO CLEARING SERVICES, LLC	4,323	$103,103	14.6%	Defined
WELLS FARGO INVESTMENT INSTITUTE, INC.	50	$1,192	0.2%	Defined

Engineers Gate Manager LP

$703,098

0.11%

29,480

First Bank & Trust

$652,899

0.10%

18,376

ALLIANCEBERNSTEIN L.P.

$646,335

0.10%

27,100

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	27,100	$646,335	100.0%	Defined

Inspire Investing, LLC

$638,758

0.10%

17,978

Universal- Beteiligungs- und Servicegesellschaft mbH

$601,520

0.09%

25,221

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Universal-Investment-Gesellschaft mbH	25,221	$601,520	100.0%	Defined

TRUIST FINANCIAL CORP

$567,296

0.09%

23,786

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Truist Advisory Services, Inc.	23,786	$567,296	100.0%	Defined

DekaBank Deutsche Girozentrale

$560,272

0.09%

15,769

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Deka Investment GmbH	15,769	$560,272	100.0%	Defined

DIVERSIFY WEALTH MANAGEMENT, LLC

$551,923

0.08%

15,534

Diversify Advisory Services, LLC

$551,923

0.08%

15,534

Tidal Investments LLC

$529,136

0.08%

22,186

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Amplify Investments, LLC	22,186	$529,136	100.0%	Defined

Green Alpha Advisors, LLC

$491,308

0.07%

13,828

Legal & General Group Plc

$490,068

0.07%

20,548

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	2,905	$69,284	14.1%	Defined
Legal & General Investment Management Ltd joint	17,643	$420,784	85.9%	Defined

Oak Harvest Investment Services

$479,122

0.07%

13,485

Schonfeld Strategic Advisors LLC

$475,330

0.07%

19,930

VANGUARD ASSET MANAGEMENT, Ltd

$421,024

0.06%

17,653

MetLife Investment Management, LLC

$405,640

0.06%

17,008

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
METLIFE INC	17,008	$405,640	100.0%	Defined

Campbell & CO Investment Adviser LLC

$392,952

0.06%

16,476

Quantinno Capital Management LP

$391,807

0.06%

16,428

CITIGROUP INC

$385,033

0.06%

16,144

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	16,144	$385,033	100.0%	Defined

PERKINS CAPITAL MANAGEMENT INC

$384,939

0.06%

16,140

Zurcher Kantonalbank (Zurich Cantonalbank)

$371,559

0.06%

15,579

Jupiter Topco LLC

$358,632

0.05%

15,037

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	15,037	$358,632	100.0%	Defined

TUDOR INVESTMENT CORP ET AL

$344,918

0.05%

14,462

LPL Financial LLC

$339,147

0.05%

14,220

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Key facts CIK 1447362 CUSIP 14843C105 13F (30d) 30 filings 30 filers Visit website Investor relations