GERN · Geron Corp

$1.53 +0.01 (+0.33%)

Market Cap

$976.56M

642.47M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Quarter:

Holders: 259

Total Value: $611,495,336

Total Shares: 477,730,836

Institutional Ownership Trend

26 quarters

Holder Concentration

Across 259 13F filers · Δ vs Mar 31, 2026

Top Holder

10.3%

−3.7 pts

Top 5

41.9%

−5.1 pts

Top 10

62.2%

−5.6 pts

HHI

490

Broadly held−114

Quarterly Activity

vs Mar 31, 2026

New

Increased

Decreased

Sold Out

Unchanged

Holder count: +10

Institutional value: -10.1%

Shares held: +4.6%

Top Buyers

vs Mar 31, 2026

Institution	Added	Now Holds
Mega CITADEL ADVISORS LLC	+7,443,359	29,554,028
Mega VANGUARD PORTFOLIO MANAGEMENT LLC	+6,143,815	10,812,598
Mega GOLDMAN SACHS GROUP INC	+5,456,777	16,633,368
Mega STATE STREET CORP	+5,359,488	35,856,671
Large TORONTO DOMINION BANK	+4,107,289	4,235,000

Top Sellers

vs Mar 31, 2026

Institution	Sold	Now Holds
Large RA CAPITAL MANAGEMENT, L.P.	−20,961,615	42,809,751
Large Point72 Asset Management, L.P.	−16,411,933	4,045,201
Mega DEUTSCHE BANK AG\	−1,763,540	3,439,976
Mega Nuveen, LLC	−1,385,013	1,546,476
Mega FMR LLC	−1,094,349	5,502,807

Top New Positions

vs Mar 31, 2026

Institution	Acquired	Value (USD)
Mega VANGUARD CAPITAL MANAGEMENT LLC	27,250,215	$34,880,275
Mid Vivo Capital, LLC	8,671,797	$11,099,900
Mid SEGALL BRYANT & HAMILL, LLC	4,485,554	$5,741,509
Mid MOORE CAPITAL MANAGEMENT, LP	1,191,941	$1,525,684
Mid SummitTX Capital, L.P.	657,024	$840,990

Top Sold-Out Positions

vs Mar 31, 2026

Institution	Previously Held	Last Value (USD)
Mid Deep Track Capital, LP	31,720,278	$47,263,214
Large MARSHALL WACE, LLP	2,960,047	$4,410,470
Mid Vestal Point Capital, LP	2,540,000	$3,784,600
Large HRT FINANCIAL LP	1,214,752	$1,809,980
Large CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM	737,143	$1,098,343

Filter: New 42 Increased 94 Reduced 61 Sold out 32 Unchanged 62

Showing 100 of 266

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

BlackRock, Inc.

$62,867,924

10.28%

49,115,571

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	497,898	$637,309	1.0%	Sole
BlackRock Investment Management (Australia) Ltd	1,867	$2,389	0.0%	Sole
BlackRock Asset Management Canada Ltd	4,268	$5,463	0.0%	Sole
BlackRock Investment Management, LLC	1,524,310	$1,951,116	3.1%	Sole
BLACKROCK ADVISORS LLC	1,255,314	$1,606,801	2.6%	Sole
BlackRock Fund Advisors	25,512,974	$32,656,606	51.9%	Sole
BlackRock Institutional Trust Company, N.A.	17,310,726	$22,157,729	35.2%	Sole
BlackRock Fund Managers Ltd.	110,285	$141,164	0.2%	Sole
BlackRock Investment Management (UK) Ltd.	244,668	$313,175	0.5%	Sole
BlackRock Asset Management Ireland Ltd	2,194,089	$2,808,433	4.5%	Sole
BlackRock Asset Management Schweiz AG	84,895	$108,665	0.2%	Sole
Aperio Group, LLC	374,277	$479,074	0.8%	Sole

RA CAPITAL MANAGEMENT, L.P.

$54,796,481

8.96%

42,809,751

Soleus Capital Management, L.P.

$54,795,083

8.96%

42,808,659

STATE STREET CORP

$45,896,538

7.51%

35,856,671

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	35,856,671	$45,896,538	100.0%	Defined

CITADEL ADVISORS LLC

$37,169,827

6.08%

29,038,928

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	5,830,547	$7,463,100	20.1%	Defined
Held directly	23,208,381	$29,706,727	79.9%	Defined

VANGUARD CAPITAL MANAGEMENT LLC

$34,880,275

5.70%

27,250,215

Clearbridge Investments, LLC

$25,150,531

4.11%

19,648,853

Eversept Partners, LP

$23,958,300

3.92%

18,717,422

GOLDMAN SACHS GROUP INC

$21,290,709

3.48%

16,633,368

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	15,936,224	$20,398,366	95.8%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	678,385	$868,332	4.1%	Defined
Goldman Sachs Trust Company, N.A.	18,759	$24,011	0.1%	Defined

GEODE CAPITAL MANAGEMENT, LLC

$18,631,621

3.05%

14,555,954

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	2,015,554	$2,579,909	13.8%	Defined
Held directly	12,540,400	$16,051,712	86.2%	Defined

MORGAN STANLEY

$16,570,149

2.71%

12,945,431

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	205	$262	0.0%	Defined
MORGAN STANLEY & CO. LLC	276,854	$354,373	2.1%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	12,476,749	$15,970,238	96.4%	Defined
MORGAN STANLEY SMITH BARNEY LLC	162,381	$207,847	1.3%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	29,242	$37,429	0.2%	Defined

Nantahala Capital Management, LLC

$15,648,000

2.56%

12,225,000

VANGUARD PORTFOLIO MANAGEMENT LLC

$13,840,125

2.26%

10,812,598

Vivo Capital, LLC

$11,099,900

1.82%

8,671,797

MILLENNIUM MANAGEMENT LLC

$10,383,701

1.70%

8,112,267

Kynam Capital Management, LP

$8,750,487

1.43%

6,836,318

Candriam S.C.A.

$7,154,152

1.17%

5,589,182

FMR LLC

$7,043,592

1.15%

5,502,807

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	5,500,115	$7,040,147	100.0%	Defined
STRATEGIC ADVISERS LLC	2,692	$3,445	0.0%	Defined

Pale Fire Capital SE

$6,793,221

1.11%

5,307,204

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Barta Jan joint	5,307,204	$6,793,221	100.0%	Defined

NORTHERN TRUST CORP

$6,441,229

1.05%

5,032,212

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	4,586,370	$5,870,553	91.1%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	357,947	$458,171	7.1%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	87,895	$112,505	1.7%	Other

UBS Group AG

$6,267,001

1.02%

4,896,096

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	28,955	$37,062	0.6%	Defined
UBS AG	4,434,569	$5,676,248	90.6%	Defined
UBS Switzerland AG	432,476	$553,569	8.8%	Defined
UBS Securities LLC	96	$122	0.0%	Defined

RENAISSANCE TECHNOLOGIES LLC

$5,861,120

0.96%

4,579,000

SEGALL BRYANT & HAMILL, LLC

$5,741,509

0.94%

4,485,554

TWO SIGMA INVESTMENTS, LP

$5,514,256

0.90%

4,308,013

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$5,484,367

0.90%

4,284,662

TORONTO DOMINION BANK

$5,420,800

0.89%

4,235,000

Point72 Asset Management, L.P.

$5,177,857

0.85%

4,045,201

VANGUARD FIDUCIARY TRUST CO

$5,073,360

0.83%

3,963,563

BANK OF AMERICA CORP /DE/

$4,523,818

0.74%

3,534,234

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	2,645,377	$3,386,082	74.9%	Defined
MERRILL LYNCH, PIERCE, FENNER & SMITH INC.	33,061	$42,318	0.9%	Defined
BOFA SECURITIES, INC.	412,608	$528,138	11.7%	Defined
MANAGED ACCOUNT ADVISORS LLC joint	443,188	$567,280	12.5%	Defined

DEUTSCHE BANK AG\

$4,403,168

0.72%

3,439,976

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DWS INVESTMENT MANAGEMENT AMERICAS, INC	10,017	$12,821	0.3%	Defined
DWS Investments UK Ltd	3,429,959	$4,390,347	99.7%	Defined

Qube Research & Technologies Ltd

$4,302,827

0.70%

3,361,584

PANAGORA ASSET MANAGEMENT INC

$3,762,520

0.62%

2,939,469

ExodusPoint Capital Management, LP

$3,233,465

0.53%

2,526,145

BNP PARIBAS ARBITRAGE, SA

$3,093,044

0.51%

2,416,441

EDMOND DE ROTHSCHILD HOLDING S.A.

$2,982,400

0.49%

2,330,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
EDMOND DE ROTHSCHILD ASSET MANAGEMENT (FRANCE)	2,330,000	$2,982,400	100.0%	Sole

NAN FUNG TRINITY (HK) LTD

$2,846,144

0.47%

2,223,550

Bank of New York Mellon Corp

$2,071,352

0.34%

1,618,246

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Investments Corporation	537,499	$687,998	33.2%	Defined
The Bank of New York Mellon	1,079,894	$1,382,264	66.7%	Defined
BNY Mellon, NA	632	$808	0.0%	Defined
BNY Mellon Advisors, Inc.	219	$280	0.0%	Defined
Held directly	2	$2	0.0%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$1,980,465

0.32%

1,547,239

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	1,547,239	$1,980,465	100.0%	Other

Nuveen, LLC

$1,979,488

0.32%

1,546,476

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	43,532	$55,720	2.8%	Defined
TEACHERS ADVISORS, LLC joint	1,502,944	$1,923,768	97.2%	Defined

MOORE CAPITAL MANAGEMENT, LP

$1,525,684

0.25%

1,191,941

Invesco Ltd.

$1,504,264

0.25%

1,175,208

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	86,066	$110,164	7.3%	Defined
Invesco Trust Co joint	222,639	$284,977	18.9%	Defined
Invesco Capital Management LLC	866,503	$1,109,123	73.7%	Defined

Seven Fleet Capital Management LP

$1,353,251

0.22%

1,057,228

World Investment Advisors

$1,323,237

0.22%

1,033,779

Virtus Investment Advisers, LLC

$1,294,337

0.21%

1,011,201

Voleon Capital Management LP

$1,286,384

0.21%

1,004,988

Hennion & Walsh Asset Management, Inc.

$1,281,164

0.21%

1,000,910

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$1,259,131

0.21%

983,698

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Asset Management Switzerland AG	790,209	$1,011,467	80.3%	Sole
UBS Asset Management (UK) Ltd.	140,207	$179,464	14.3%	Other
Held directly	53,282	$68,200	5.4%	Defined

Swiss National Bank

$1,233,920

0.20%

964,000

CITIGROUP INC

$1,057,556

0.17%

826,216

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	826,216	$1,057,556	100.0%	Defined

RHUMBLINE ADVISERS

$937,043

0.15%

732,065

J. Goldman & Co LP

$911,518

0.15%

712,124

Man Group plc

$891,736

0.15%

696,669

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Numeric Investors LLC	696,669	$891,736	100.0%	Defined

SummitTX Capital, L.P.

$840,990

0.14%

657,024

ALPS ADVISORS INC

$833,381

0.14%

651,079

Exome Asset Management LLC

$798,778

0.13%

624,046

FRANKLIN RESOURCES INC

$792,180

0.13%

618,891

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FRANKLIN ADVISERS INC	618,891	$792,180	100.0%	Defined

Rafferty Asset Management, LLC

$776,243

0.13%

606,440

Arax Advisory Partners

$774,781

0.13%

605,298

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$760,704

0.12%

594,300

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	594,300	$760,704	100.0%	Other

GSA CAPITAL PARTNERS LLP

$752,851

0.12%

588,165

AMERIPRISE FINANCIAL INC

$740,250

0.12%

578,321

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	578,311	$740,238	100.0%	Defined
Ameriprise Financial Services, LLC	10	$12	0.0%	Defined

Lion Point Capital, LP

$739,840

0.12%

578,000

ALLIANCEBERNSTEIN L.P.

$731,930

0.12%

571,821

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	571,821	$731,930	100.0%	Defined

WELLS FARGO & COMPANY/MN

$714,879

0.12%

558,500

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	510,406	$653,319	91.4%	Defined
WELLS FARGO CLEARING SERVICES, LLC	48,094	$61,560	8.6%	Defined

Squarepoint Ops LLC

$684,110

0.11%

534,461

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Squarepoint Capital LLP	534,461	$684,110	100.0%	Defined

Inspire Investing, LLC

$673,232

0.11%

525,963

Quadrature Capital Ltd

$670,801

0.11%

524,064

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Quadrature US Inc.	524,064	$670,801	100.0%	Other

Nvest Financial, LLC

$661,952

0.11%

517,150

ADAR1 Capital Management, LLC

$647,445

0.11%

505,817

Private Advisor Group, LLC

$643,683

0.11%

502,878

Verdad Advisers, LP

$632,577

0.10%

494,201

XTX Topco Ltd

$593,788

0.10%

463,897

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	463,897	$593,788	100.0%	Defined

PRICE T ROWE ASSOCIATES INC /MD/

$588,267

0.10%

459,584

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	459,584	$588,267	100.0%	Defined

CITADEL ADVISORS LLC

$549,760

0.09%

429,500

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	429,500	$549,760	100.0%	Defined

SG Americas Securities, LLC

$487,827

0.08%

381,115

RAYMOND JAMES FINANCIAL INC

$436,016

0.07%

340,638

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	327,126	$418,721	96.0%	Defined
Raymond James Financial Services Advisors, Inc.	13,512	$17,295	4.0%	Defined

MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.

$419,953

0.07%

328,089

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Global X Management CO LLC joint	257,642	$329,781	78.5%	Defined
Global X Management (AUS) Ltd joint	70,447	$90,172	21.5%	Defined

MetLife Investment Management, LLC

$419,818

0.07%

327,983

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
METLIFE INC	327,983	$419,818	100.0%	Defined

VANGUARD ASSET MANAGEMENT, Ltd

$400,789

0.07%

313,117

Legal & General Group Plc

$396,218

0.06%

309,547

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	54,246	$69,434	17.5%	Defined
Legal & General Investment Management Ltd joint	255,301	$326,784	82.5%	Defined

PROFUND ADVISORS LLC

$392,430

0.06%

306,586

MOODY NATIONAL BANK TRUST DIVISION

$380,421

0.06%

297,204

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MOODY BANCSHARES INC	297,204	$380,421	100.0%	Defined

JPMORGAN CHASE & CO

$370,590

0.06%

289,524

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	231,959	$296,907	80.1%	Defined
J.P. Morgan Investment Management Inc.	6,954	$8,901	2.4%	Defined
J.P. Morgan Securities plc	50,611	$64,782	17.5%	Defined

MML INVESTORS SERVICES, LLC

$317,319

0.05%

247,906

Jupiter Topco LLC

$306,376

0.05%

239,357

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	239,357	$306,376	100.0%	Defined

Mitsubishi UFJ Trust & Banking Corp

$288,757

0.05%

225,592

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
First Sentier Investors RQI Pty Ltd	225,592	$288,757	100.0%	Defined

Vanguard Global Advisers, LLC

$281,386

0.05%

219,833

Diversified Trust Co

$279,718

0.05%

218,530

ALLSTATE CORP

$269,036

0.04%

210,185

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ALLSTATE INVESTMENTS LLC	210,185	$269,036	100.0%	Sole

STATE OF WISCONSIN INVESTMENT BOARD

$268,279

0.04%

209,593

Mason & Associates Inc

$256,399

0.04%

200,312

Verition Fund Management LLC

$256,000

0.04%

200,000

Cresset Asset Management, LLC

$255,624

0.04%

199,707

JANE STREET GROUP, LLC

$246,912

0.04%

192,900

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	192,900	$246,912	100.0%	Defined

ProShare Advisors LLC

$241,397

0.04%

188,592

LPL Financial LLC

$233,493

0.04%

182,417

Verition Fund Management LLC

$222,321

0.04%

173,689

NEW YORK STATE COMMON RETIREMENT FUND

$219,459

0.04%

171,453

Creative Planning

$212,498

0.03%

166,015

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CREATIVE PLANNING TRUST Co LLC	4,931	$6,311	3.0%	Defined
Held directly	161,084	$206,187	97.0%	Sole

Headlands Technologies LLC

$204,202

0.03%

159,533

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC	153,940	$197,042	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Capital	153,371	$196,314	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MANULIFE FINANCIAL CORP joint	148,907	$190,600	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Amundi Asset Management	132,367	$169,429	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	65,830	$84,262	53.1%	Defined
D. E. Shaw Investment Management, L.L.C. joint	58,101	$74,369	46.9%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	100,445	$128,569	87.2%	Defined
BARCLAYS CAPITAL INC.	14,795	$18,937	12.8%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DRW Investments joint	111,450	$142,656	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
OSAIC WEALTH, INC.	83,154	$106,437	74.6%	Defined
CW Advisors, LLC	22,970	$29,401	20.6%	Sole
Osaic Advisory Services, LLC	5,300	$6,784	4.8%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	109,231	$139,815	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	106,400	$136,192	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PGIM Quantitative Solutions LLC	95,832	$122,664	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GKFO LLC	88,779	$113,637	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	85,600	$109,568	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Credit Agricole Corporate & Investment Bank joint	82,100	$105,088	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CIBC Private Wealth Advisors, Inc.	42,000	$53,760	57.6%	Defined
CIBC National Trust Co	80	$102	0.1%	Defined
CIBC WORLD MARKETS CORP	30,891	$39,540	42.3%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Virtu Americas LLC	64,470	$82,521	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
TEACHERS INSURANCE & ANNUITY ASSOCIATION OF AMERICA	58,250	$74,560	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RF&L WEALTH MANAGEMENT, LLC	82	$104	0.1%	Other
Arax Advisory Partners	1,565	$2,003	2.8%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	206	$263	0.4%	Other
Stokes Family Office, LLC	17	$21	0.0%	Other
Held directly	53,835	$68,908	96.6%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
EATON VANCE MANAGEMENT	45,637	$58,415	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Edward D. Jones & Co., L.P.	43,829	$56,101	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GUGGENHEIM FUNDS DISTRIBUTORS, LLC	32,104	$41,093	85.6%	Defined
SECURITY INVESTORS LLC	5,383	$6,890	14.4%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	34,600	$44,288	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI INVESTMENTS MANAGEMENT CORP	641	$820	1.9%	Sole
SEI TRUST CO	33,408	$42,762	98.1%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Latour Trading LLC (Latour)	31,774	$40,670	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Brevan Howard US Investment Management, LP	31,173	$39,901	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BMO Bank N.A.	25,972	$33,244	86.4%	Defined
BMO NESBITT BURNS INC.	1,100	$1,408	3.7%	Defined
Stoker Ostler Wealth Advisors, Inc.	3,000	$3,840	10.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ROCKEFELLER & CO. LLC	1,072	$1,372	4.3%	Defined
ROCKEFELLER TRUST COMPANY, N.A.	1,455	$1,862	5.8%	Defined
ROCKEFELLER FINANCIAL LLC	22,654	$28,997	90.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
HSBC Bank USA, N.A.	1,433	$1,834	7.4%	Defined
HSBC Global Asset Management (UK) Ltd	13,993	$17,911	72.3%	Defined
HSBC SECURITIES USA INC	3,939	$5,041	20.3%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Universal-Investment-Gesellschaft mbH	18,927	$24,226	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC CMA LTD.	17,027	$21,794	97.1%	Defined
RBC Dominion Securities Inc.	510	$652	2.9%	Defined
Held directly	4	$5	0.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNP PARIBAS ASSET MANAGEMENT Europe	8,376	$10,721	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
US BANK NATIONAL ASSOCIATION	8,000	$10,240	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nykredit Bank A/S	7,089	$9,073	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fifth Third Bank, National Association	3,813	$4,880	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PNC Bank, National Association	3,161	$4,046	96.0%	Defined
PNC Wealth Management LLC	133	$170	4.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MITSUBISHI UFJ FINANCIAL GROUP INC	1,441	$1,844	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Optiver VOF	545	$697	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marshall Wace North America L.P. joint	2,960,047	$4,410,470	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Hudson Bay Capital UK LLP	60,800	$90,592	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STIFEL NICOLAUS & CO INC \MO\	14,442	$21,518	97.3%	Defined
Held directly	394	$587	2.7%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Robert W. Baird & Co. Inc	11,500	$17,135	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wilmington Trust National Association joint	10,787	$16,071	100.0%	Defined

Key facts CIK 886744 CUSIP 374163103 13F (30d) 226 filings 220 filers Visit website Investor relations