PRTO · Protara Therapeutics, Inc.

$28.15 -0.38 (-1.32%) At close · Aug 28

Russell 2000 member Rebalance outlook locked

Market Cap

$226.04M

59.10M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Quarter:

Holders: 120

Total Value: $176,085,219

Total Shares: 46,095,626

Showing 1–100 of 121 holdings by value · page 1 of 2

Institutional Ownership Trend

25 quarters

Holder Concentration

Across 120 13F filers · Δ vs Mar 31, 2026

Top Holder

10.2%

−1.2 pts

Top 5

36.0%

−0.1 pts

Top 10

57.0%

+0.7 pts

HHI

435

Broadly held−10

Quarterly Activity

vs Mar 31, 2026

New

Increased

Decreased

Sold Out

Unchanged

Holder count: +5

Institutional value: -26.2%

Shares held: +0.7%

Top Buyers

vs Mar 31, 2026

Institution	Added	Now Holds
Mega Qube Research & Technologies Ltd	+868,512	1,045,516
Large MARSHALL WACE, LLP	+650,290	701,021
Mega MILLENNIUM MANAGEMENT LLC	+269,958	3,101,166
Mega BlackRock, Inc.	+268,531	3,663,499
Large JACOBS LEVY EQUITY MANAGEMENT, INC	+245,310	295,565

Top Sellers

vs Mar 31, 2026

Institution	Sold	Now Holds
Mega UBS Group AG	−945,640	456,773
Mid J. Goldman & Co LP	−721,247	135,000
Large Point72 Asset Management, L.P.	−603,782	548,932
Large Assenagon Asset Management S.A.	−507,667	59,954
Large Alyeska Investment Group, L.P.	−310,274	39,726

Top New Positions

vs Mar 31, 2026

Institution	Acquired	Value (USD)
Mega Jupiter Topco LLC	4,696,709	$17,941,428
Mid ORBIMED ADVISORS LLC	2,220,000	$8,480,400
Large Balyasny Asset Management L.P.	1,026,691	$3,921,959
Small Verdad Advisers, LP	312,249	$1,192,791
Mega Fisher Asset Management, LLC	99,566	$380,342

Top Sold-Out Positions

vs Mar 31, 2026

Institution	Previously Held	Last Value (USD)
Mega JANUS HENDERSON GROUP PLC	5,207,011	$27,128,527
Mid AIGH Capital Management LLC	1,622,733	$8,454,438
Small Patient Square Capital LP	1,030,846	$5,370,707
Mid Burkehill Global Management, LP	450,000	$2,344,500
Large ExodusPoint Capital Management, LP	116,305	$605,949

Filter: New 26 Increased 50 Reduced 29 Sold out 21 Unchanged 15

Showing 100 of 121

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

Jupiter Topco LLC

$17,941,428

10.19%

4,696,709

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS US LLC	4,696,709	$17,941,428	100.0%	Defined

BlackRock, Inc.

$13,994,564

7.95%

3,663,499

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	15,432	$58,950	0.4%	Sole
BlackRock Investment Management, LLC	145,096	$554,266	4.0%	Sole
BLACKROCK ADVISORS LLC	99,892	$381,587	2.7%	Sole
BlackRock Fund Advisors	2,020,359	$7,717,771	55.1%	Sole
BlackRock Institutional Trust Company, N.A.	1,382,720	$5,281,990	37.7%	Sole

MILLENNIUM MANAGEMENT LLC

$11,846,454

6.73%

3,101,166

Velan Capital Investment Management LP

$10,866,570

6.17%

2,844,652

VANGUARD CAPITAL MANAGEMENT LLC

$8,786,508

4.99%

2,300,133

Blackstone Inc.

$8,654,263

4.91%

2,265,514

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Blackstone Alternative Asset Management L.P.	2,265,514	$8,654,263	100.0%	Other

ORBIMED ADVISORS LLC

$8,480,400

4.82%

2,220,000

Sio Capital Management, LLC

$7,341,551

4.17%

1,921,872

ACORN CAPITAL ADVISORS, LLC

$6,693,079

3.80%

1,752,115

Integral Health Asset Management, LLC

$5,730,000

3.25%

1,500,000

5AM Venture Management, LLC

$5,232,567

2.97%

1,369,782

GEODE CAPITAL MANAGEMENT, LLC

$5,066,259

2.88%

1,326,246

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	184,536	$704,927	13.9%	Defined
Held directly	1,141,710	$4,361,332	86.1%	Defined

CITADEL ADVISORS LLC

$4,707,633

2.67%

1,232,365

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	338,636	$1,293,589	27.5%	Defined
Held directly	893,729	$3,414,044	72.5%	Defined

STATE STREET CORP

$4,080,886

2.32%

1,068,295

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	1,068,295	$4,080,886	100.0%	Defined

Qube Research & Technologies Ltd

$3,993,871

2.27%

1,045,516

Balyasny Asset Management L.P.

$3,921,959

2.23%

1,026,691

BAKER BROS. ADVISORS LP

$3,711,565

2.11%

971,614

ADAR1 Capital Management, LLC

$3,592,755

2.04%

940,512

BANK OF AMERICA CORP /DE/

$3,369,923

1.91%

882,179

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	860,649	$3,287,679	97.6%	Defined
BOFA SECURITIES, INC.	21,530	$82,244	2.4%	Defined

MARSHALL WACE, LLP

$2,677,899

1.52%

701,021

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marshall Wace North America L.P. joint	701,021	$2,677,899	100.0%	Other

DIMENSIONAL FUND ADVISORS LP

$2,609,937

1.48%

683,230

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	5,637	$21,533	0.8%	Defined
DFA Australia Ltd.	12,723	$48,601	1.9%	Defined
Dimensional Ireland Ltd	39,237	$149,885	5.7%	Defined
Held directly	625,633	$2,389,918	91.6%	Sole

NORTHERN TRUST CORP

$2,109,807

1.20%

552,306

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	543,913	$2,077,746	98.5%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	8,393	$32,061	1.5%	Other

Point72 Asset Management, L.P.

$2,096,920

1.19%

548,932

GOLDMAN SACHS GROUP INC

$2,086,586

1.18%

546,227

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	506,834	$1,936,105	92.8%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	39,393	$150,481	7.2%	Defined

Catalio Capital Management, LP

$1,986,965

1.13%

520,148

MORGAN STANLEY

$1,942,217

1.10%

508,434

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	280	$1,069	0.1%	Defined
MORGAN STANLEY & CO. LLC	16,050	$61,311	3.2%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	480,083	$1,833,917	94.4%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	12,021	$45,920	2.4%	Defined

Walleye Capital LLC

$1,930,750

1.10%

505,432

VANGUARD PORTFOLIO MANAGEMENT LLC

$1,878,129

1.07%

491,657

UBS Group AG

$1,744,871

0.99%

456,773

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS AG	451,188	$1,723,538	98.8%	Defined
UBS Switzerland AG	1,746	$6,669	0.4%	Defined
UBS Securities LLC	3,839	$14,664	0.8%	Defined

VANGUARD FIDUCIARY TRUST CO

$1,278,489

0.73%

334,683

Verdad Advisers, LP

$1,192,791

0.68%

312,249

JACOBS LEVY EQUITY MANAGEMENT, INC

$1,129,058

0.64%

295,565

RENAISSANCE TECHNOLOGIES LLC

$960,730

0.55%

251,500

OPPENHEIMER & CO INC

$910,542

0.52%

238,362

AQR CAPITAL MANAGEMENT LLC

$798,259

0.45%

208,969

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RF&L WEALTH MANAGEMENT, LLC	5	$19	0.0%	Other
Prism Planning Partners LLC	47	$179	0.0%	Other
Arax Advisory Partners	1,358	$5,187	0.6%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	788	$3,010	0.4%	Other
Stokes Family Office, LLC	545	$2,081	0.3%	Other
Held directly	206,226	$787,783	98.7%	Defined

Jump Financial, LLC

$775,842

0.44%

203,100

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Leap GP LLC joint	203,100	$775,842	100.0%	Defined

CIBC WORLD MARKET INC.

$636,010

0.36%

166,495

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$603,449

0.34%

157,971

Bank of New York Mellon Corp

$525,969

0.30%

137,689

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Investments Corporation	47,162	$180,158	34.3%	Defined
The Bank of New York Mellon	79,039	$301,928	57.4%	Defined
Newton Investment Management North America, LLC	11,486	$43,876	8.3%	Defined
Held directly	2	$7	0.0%	Defined

J. Goldman & Co LP

$515,700

0.29%

135,000

SUPERSTRING CAPITAL MANAGEMENT LP

$463,965

0.26%

121,457

Fisher Asset Management, LLC

$380,342

0.22%

99,566

AMERICAN CENTURY COMPANIES INC

$376,220

0.21%

98,487

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AMERICAN CENTURY INVESTMENT MANAGEMENT INC	98,487	$376,220	100.0%	Defined

BRIDGEWAY CAPITAL MANAGEMENT, LLC

$325,960

0.19%

85,330

Nuveen, LLC

$277,369

0.16%

72,610

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
TIAA CREF INVESTMENT MANAGEMENT LLC joint	2,524	$9,641	3.5%	Defined
TEACHERS ADVISORS, LLC joint	70,086	$267,728	96.5%	Defined

TWO SIGMA INVESTMENTS, LP

$266,070

0.15%

69,652

FMR LLC

$260,523

0.15%

68,200

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	68,066	$260,012	99.8%	Defined
STRATEGIC ADVISERS LLC	134	$511	0.2%	Defined

XTX Topco Ltd

$244,281

0.14%

63,948

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	63,948	$244,281	100.0%	Defined

RHUMBLINE ADVISERS

$241,595

0.14%

63,245

TRUIST FINANCIAL CORP

$229,200

0.13%

60,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
TRUIST BANK /NC/	60,000	$229,200	100.0%	Defined

Assenagon Asset Management S.A.

$229,024

0.13%

59,954

Trexquant Investment LP

$227,733

0.13%

59,616

DEUTSCHE BANK AG\

$220,463

0.13%

57,713

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DWS Investments UK Ltd	57,713	$220,463	100.0%	Defined

ALPS ADVISORS INC

$217,896

0.12%

57,041

AMERIPRISE FINANCIAL INC

$214,939

0.12%

56,267

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	56,247	$214,863	100.0%	Defined
Ameriprise Financial Services, LLC	20	$76	0.0%	Defined

WELLS FARGO & COMPANY/MN

$208,556

0.12%

54,596

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	43,372	$165,681	79.4%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	10,775	$41,160	19.7%	Defined
Wells Fargo Securities, LLC	449	$1,715	0.8%	Defined

PRUDENTIAL FINANCIAL INC

$193,788

0.11%

50,730

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PGIM Quantitative Solutions LLC	50,730	$193,788	100.0%	Defined

NANO CAP NEW MILLENNIUM GROWTH FUND L P

$191,000

0.11%

50,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AZG TANGIBLE ASSETS FUND LLC	50,000	$191,000	100.0%	Sole

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$154,251

0.09%

40,380

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	40,380	$154,251	100.0%	Other

Alyeska Investment Group, L.P.

$151,753

0.09%

39,726

Quadrature Capital Ltd

$150,198

0.09%

39,319

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Quadrature US Inc.	39,319	$150,198	100.0%	Other

PRICE T ROWE ASSOCIATES INC /MD/

$147,005

0.08%

38,483

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	38,483	$147,005	100.0%	Defined

Marex Group plc

$134,655

0.08%

35,250

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marex Financial	35,250	$134,655	100.0%	Defined

WINTON GROUP Ltd

$125,296

0.07%

32,800

PUBLIC EMPLOYEES RETIREMENT SYSTEM OF OHIO

$113,454

0.06%

29,700

Russell Investments Group, Ltd.

$108,751

0.06%

28,469

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC	26,876	$102,666	94.4%	Defined
Russell Investments Trust Company joint	1,593	$6,085	5.6%	Other

MetLife Investment Management, LLC

$108,403

0.06%

28,378

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
METLIFE INC	28,378	$108,403	100.0%	Defined

Invesco Ltd.

$102,589

0.06%

26,856

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	6,360	$24,295	23.7%	Defined
Invesco Trust Co joint	20,496	$78,294	76.3%	Defined

CITIGROUP INC

$92,233

0.05%

24,145

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	24,145	$92,233	100.0%	Defined

MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.

$92,042

0.05%

24,095

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Global X Management CO LLC joint	24,095	$92,042	100.0%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$87,478

0.05%

22,900

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	22,900	$87,478	100.0%	Other

JPMORGAN CHASE & CO

$86,361

0.05%

22,608

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	22,144	$84,589	97.9%	Defined
J.P. Morgan Investment Management Inc.	4	$15	0.0%	Defined
J.P. Morgan Securities plc	460	$1,757	2.0%	Defined

Jain Global LLC

$80,220

0.05%

21,000

Vanguard Global Advisers, LLC

$76,885

0.04%

20,127

PRELUDE CAPITAL MANAGEMENT, LLC

$74,921

0.04%

19,613

EntryPoint Capital, LLC

$73,672

0.04%

19,286

LPL Financial LLC

$73,076

0.04%

19,130

BARCLAYS PLC

$71,078

0.04%

18,607

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	15,881	$60,665	85.3%	Defined
BARCLAYS CAPITAL INC.	2,726	$10,413	14.7%	Defined

Squarepoint Ops LLC

$69,233

0.04%

18,124

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Squarepoint Capital LLP	18,124	$69,233	100.0%	Defined

ALLIANCEBERNSTEIN L.P.

$68,912

0.04%

18,040

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	18,040	$68,912	100.0%	Defined

IEQ CAPITAL, LLC

$68,488

0.04%

17,929

Police & Firemen's Retirement System of New Jersey

$67,113

0.04%

17,569

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Capital	17,569	$67,113	100.0%	Defined

MANUFACTURERS LIFE INSURANCE COMPANY, THE

$56,833

0.03%

14,878

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MANULIFE FINANCIAL CORP joint	14,878	$56,833	100.0%	Defined

INTECH INVESTMENT MANAGEMENT LLC

$51,806

0.03%

13,562

Mariner, LLC

$45,454

0.03%

11,899

VOYA INVESTMENT MANAGEMENT LLC

$43,811

0.02%

11,469

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	11,469	$43,811	100.0%	Defined

D. E. Shaw & Co., Inc.

$42,451

0.02%

11,113

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	11,113	$42,451	100.0%	Defined

COMMONWEALTH EQUITY SERVICES, LLC

$42,020

0.02%

11,000

Corient Private Wealth LLC

$40,835

0.02%

10,690

Essential Partners LLC

$37,443

0.02%

9,802

NEW YORK STATE COMMON RETIREMENT FUND

$30,178

0.02%

7,900

ROYAL BANK OF CANADA

$29,654

0.02%

7,763

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC CMA LTD.	1,577	$6,024	20.3%	Defined
RBC Capital Markets, LLC	2,686	$10,260	34.6%	Defined
RBC Dominion Securities Inc.	3,500	$13,370	45.1%	Defined

BNP PARIBAS ARBITRAGE, SA

$28,787

0.02%

7,536

Tower Research Capital LLC (TRC)

$19,061

0.01%

4,990

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Latour Trading LLC (Latour)	4,990	$19,061	100.0%	Defined

Legal & General Group Plc

$17,151

0.01%

4,490

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	4,490	$17,151	100.0%	Defined

Ameritas Investment Partners, Inc.

$12,598

0.01%

3,298

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
EATON VANCE MANAGEMENT	3,298	$12,598	100.0%	Other

Integrated Wealth Concepts LLC

$6,685

0.00%

1,750

Arax Advisory Partners

$6,383

0.00%

1,671

Sterling Capital Management LLC

$4,030

0.00%

1,055

State of Wyoming

$3,667

0.00%

960

Key facts CIK 1359931 CUSIP 74365U107 13F (30d) 106 filings 106 filers Visit website Investor relations