CDNA · CareDx, Inc.

$46.99 +0.48 (+1.03%) At close · Aug 20

Market Cap

$2.43B

51.73M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Quarter:

Holders: 225

Total Value: $1,504,015,230

Total Shares: 52,772,485

Institutional Ownership Trend

26 quarters

Superinvestors Holding CDNA

All superinvestors →

Joel Greenblatt $2,097,001

Holder Concentration

Across 225 13F filers · Δ vs Mar 31, 2026

Top Holder

9.1%

+0.5 pts

Top 5

25.5%

−3.2 pts

Top 10

41.4%

−5.3 pts

HHI

277

Broadly held−37

Quarterly Activity

vs Mar 31, 2026

New

Increased

Decreased

Sold Out

Unchanged

Holder count: +35

Institutional value: +64.9%

Shares held: +0.4%

Top Buyers

vs Mar 31, 2026

Institution	Added	Now Holds
Mega MORGAN STANLEY	+348,950	774,299
Mid Portolan Capital Management, LLC	+344,453	684,805
Mega BlackRock, Inc.	+292,613	4,814,709
Mid Ophir Asset Management Pty Ltd	+276,758	1,264,809
Mega D. E. Shaw & Co., Inc.	+262,752	1,137,660

Top Sellers

vs Mar 31, 2026

Institution	Sold	Now Holds
Large BAMCO INC /NY/	−1,399,083	1,937,326
Mid BRAIDWELL LP	−952,544	1,220,602
Mega STATE STREET CORP	−291,527	2,468,858
Mid Amova Asset Management Americas, Inc.	−287,051	1,188,075
Mega Sumitomo Mitsui Trust Group, Inc.	−283,496	1,191,630

Top New Positions

vs Mar 31, 2026

Institution	Acquired	Value (USD)
Mid Divisadero Street Capital Management, LP	700,938	$19,976,733
Mega ARROWSTREET CAPITAL, LIMITED PARTNERSHIP	372,111	$10,605,163
Mega Qube Research & Technologies Ltd	369,837	$10,540,354
Mega Jupiter Topco LLC	339,418	$9,673,413
Large ExodusPoint Capital Management, LP	214,342	$6,108,747

Top Sold-Out Positions

vs Mar 31, 2026

Institution	Previously Held	Last Value (USD)
Micro PARAGON ASSOCIATES & PARAGON ASSOCIATES II JOINT VENTURE	500,000	$8,680,000
Mega JANUS HENDERSON GROUP PLC	316,519	$5,494,769
Large CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM	45,903	$796,876
Mid ALGERT GLOBAL LLC	44,510	$772,693
Mid Brevan Howard Capital Management LP	17,604	$305,605

Filter: New 54 Increased 75 Reduced 68 Sold out 19 Unchanged 28

Showing 100 of 228

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

BlackRock, Inc.

$137,219,202

9.12%

4,814,709

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	60,903	$1,735,735	1.3%	Sole
BlackRock Investment Management (Australia) Ltd	208	$5,928	0.0%	Sole
BlackRock Asset Management Canada Ltd	22,746	$648,261	0.5%	Sole
BlackRock Investment Management, LLC	154,866	$4,413,681	3.2%	Sole
BLACKROCK ADVISORS LLC	115,438	$3,289,983	2.4%	Sole
BlackRock Fund Advisors	2,589,051	$73,787,953	53.8%	Sole
BlackRock Institutional Trust Company, N.A.	1,600,106	$45,603,021	33.2%	Sole
BlackRock Fund Managers Ltd.	7,905	$225,292	0.2%	Sole
BlackRock Investment Management (UK) Ltd.	1,282	$36,537	0.0%	Sole
BlackRock (Netherlands) B.V.	851	$24,253	0.0%	Sole
BlackRock Asset Management Ireland Ltd	166,097	$4,733,764	3.4%	Sole
BlackRock (Luxembourg) S.A.	6,567	$187,159	0.1%	Sole
BlackRock Life Ltd	781	$22,258	0.0%	Sole
BlackRock Asset Management Schweiz AG	10,750	$306,375	0.2%	Sole
Aperio Group, LLC	70,891	$2,020,393	1.5%	Sole
SpiderRock Advisors, LLC	6,267	$178,609	0.1%	Sole

STATE STREET CORP

$70,362,453

4.68%

2,468,858

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	2,468,858	$70,362,453	100.0%	Defined

VANGUARD CAPITAL MANAGEMENT LLC

$62,628,037

4.16%

2,197,475

DEERFIELD MANAGEMENT COMPANY, L.P.

$56,932,113

3.79%

1,997,618

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Deerfield UGP, LLC	1,997,618	$56,932,113	100.0%	Defined

ARK Investment Management LLC

$55,675,633

3.70%

1,953,531

Boston Partners

$55,221,628

3.67%

1,937,601

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Boston Partners Trust Co	56,372	$1,606,602	2.9%	Defined
Held directly	1,881,229	$53,615,026	97.1%	Defined

BAMCO INC /NY/

$55,213,791

3.67%

1,937,326

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARON CAPITAL MANAGEMENT INC /NY/ joint	1,937,326	$55,213,791	100.0%	Defined

GAGNON SECURITIES LLC

$50,465,947

3.36%

1,770,735

Chevy Chase Trust Holdings, LLC

$40,966,356

2.72%

1,437,416

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CHEVY CHASE TRUST Co	1,437,416	$40,966,356	100.0%	Defined

GEODE CAPITAL MANAGEMENT, LLC

$37,849,054

2.52%

1,328,037

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	189,871	$5,411,323	14.3%	Defined
Held directly	1,138,166	$32,437,731	85.7%	Defined

Ophir Asset Management Pty Ltd

$36,047,056

2.40%

1,264,809

RENAISSANCE TECHNOLOGIES LLC

$35,424,331

2.36%

1,242,959

BRAIDWELL LP

$34,787,157

2.31%

1,220,602

Sumitomo Mitsui Trust Group, Inc.

$33,961,455

2.26%

1,191,630

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Amova Asset Management Co., Ltd.	1,191,630	$33,961,455	100.0%	Defined

Blue Water Life Science Advisors, LP

$33,915,000

2.25%

1,190,000

Amova Asset Management Americas, Inc.

$33,860,137

2.25%

1,188,075

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Sumitomo Mitsui Trust Group, Inc. joint	1,188,075	$33,860,137	100.0%	Sole

DIMENSIONAL FUND ADVISORS LP

$33,660,693

2.24%

1,181,077

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	6,230	$177,555	0.5%	Defined
DFA Australia Ltd.	18,043	$514,225	1.5%	Defined
Dimensional Fund Advisors Pte. Ltd.	1,024	$29,184	0.1%	Defined
Dimensional Ireland Ltd	54,915	$1,565,077	4.6%	Defined
Held directly	1,100,865	$31,374,652	93.2%	Sole

D. E. Shaw & Co., Inc.

$32,423,309

2.16%

1,137,660

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	1,061,375	$30,249,187	93.3%	Defined
D. E. Shaw Investment Management, L.L.C. joint	76,285	$2,174,122	6.7%	Defined

JACOBS LEVY EQUITY MANAGEMENT, INC

$28,376,566

1.89%

995,669

VANGUARD PORTFOLIO MANAGEMENT LLC

$28,201,576

1.88%

989,529

MORGAN STANLEY

$22,067,520

1.47%

774,299

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	116	$3,306	0.0%	Defined
MORGAN STANLEY & CO. LLC	17,218	$490,713	2.2%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	614,667	$17,518,009	79.4%	Defined
MORGAN STANLEY INVESTMENT MANAGEMENT INC	1,698	$48,393	0.2%	Defined
MORGAN STANLEY SMITH BARNEY LLC	12,056	$343,595	1.6%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	128,544	$3,663,504	16.6%	Defined

Divisadero Street Capital Management, LP

$19,976,733

1.33%

700,938

GOLDMAN SACHS GROUP INC

$19,949,800

1.33%

699,993

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	415,737	$11,848,504	59.4%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	284,256	$8,101,296	40.6%	Defined

Portolan Capital Management, LLC

$19,516,942

1.30%

684,805

First Eagle Investment Management, LLC

$17,057,164

1.13%

598,497

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FIRST EAGLE SEPARATE ACCOUNT MANAGEMENT, LLC	276	$7,866	0.0%	Sole
Held directly	598,221	$17,049,298	100.0%	Sole

ROYCE & ASSOCIATES LP

$16,415,173

1.09%

575,971

BANK OF AMERICA CORP /DE/

$16,296,528

1.08%

571,808

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	260,884	$7,435,194	45.6%	Defined
BOFA SECURITIES, INC.	49,770	$1,418,445	8.7%	Defined
MANAGED ACCOUNT ADVISORS LLC joint	261,154	$7,442,889	45.7%	Defined

Gagnon Advisors, LLC

$15,586,450

1.04%

546,893

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$15,336,562

1.02%

538,125

Palo Alto Investors LP

$15,158,295

1.01%

531,870

Russell Investments Group, Ltd.

$14,636,173

0.97%

513,550

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC joint	261,329	$7,447,876	50.9%	Defined
Russell Investments Canada Limited joint	12,303	$350,635	2.4%	Other
Russell Investments Capital, LLC joint	63,371	$1,806,073	12.3%	Defined
Russell Investments France SAS	35,539	$1,012,861	6.9%	Defined
Russell Investments Limited	2,866	$81,681	0.6%	Defined
Russell Investments Trust Company joint	138,142	$3,937,047	26.9%	Other

NORTHERN TRUST CORP

$13,645,741

0.91%

478,798

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	443,314	$12,634,448	92.6%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	27,606	$786,770	5.8%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	7,878	$224,523	1.6%	Other

JANE STREET GROUP, LLC

$13,402,780

0.89%

470,273

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	346,046	$9,862,311	73.6%	Defined
Jane Street Global Trading, LLC	124,227	$3,540,469	26.4%	Defined

Meridiem Capital Partners LP

$12,867,550

0.86%

451,493

Invesco Ltd.

$12,745,341

0.85%

447,205

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	7,924	$225,833	1.8%	Defined
Invesco Trust Co joint	20,371	$580,573	4.6%	Defined
Invesco Capital Management LLC	418,910	$11,938,935	93.7%	Defined

TWO SIGMA INVESTMENTS, LP

$11,456,629

0.76%

401,987

Zweig-DiMenna Associates LLC

$11,081,142

0.74%

388,812

BNP PARIBAS ARBITRAGE, SA

$10,684,906

0.71%

374,909

ARROWSTREET CAPITAL, LIMITED PARTNERSHIP

$10,605,163

0.71%

372,111

Qube Research & Technologies Ltd

$10,540,354

0.70%

369,837

VANGUARD FIDUCIARY TRUST CO

$10,446,418

0.69%

366,541

Jupiter Topco LLC

$9,673,413

0.64%

339,418

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	21,900	$624,150	6.5%	Defined
JANUS HENDERSON INVESTORS US LLC	317,518	$9,049,263	93.5%	Defined

ArrowMark Colorado Holdings LLC

$9,648,646

0.64%

338,549

SILVERCREST ASSET MANAGEMENT GROUP LLC

$8,441,700

0.56%

296,200

Connor, Clark & Lunn Investment Management Ltd.

$7,643,899

0.51%

268,207

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 1	268,207	$7,643,899	100.0%	Sole

FMR LLC

$7,219,449

0.48%

253,314

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	252,320	$7,191,120	99.6%	Defined
STRATEGIC ADVISERS LLC	994	$28,329	0.4%	Defined

AQR CAPITAL MANAGEMENT LLC

$6,964,687

0.46%

244,375

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RF&L WEALTH MANAGEMENT, LLC	8	$228	0.0%	Other
Prism Planning Partners LLC	104	$2,964	0.0%	Other
Compound Planning, Inc.	16	$456	0.0%	Other
Arax Advisory Partners	1,066	$30,381	0.4%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	1,049	$29,896	0.4%	Other
Stokes Family Office, LLC	988	$28,158	0.4%	Other
Held directly	241,144	$6,872,604	98.7%	Defined

Aberdeen Group plc

$6,407,626

0.43%

224,829

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
abrdn Inc.	224,829	$6,407,626	100.0%	Defined

ExodusPoint Capital Management, LP

$6,108,747

0.41%

214,342

Legal & General Group Plc

$6,041,058

0.40%

211,967

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	4,979	$141,901	2.3%	Defined
Legal & General Investment Management Ltd joint	206,988	$5,899,157	97.7%	Defined

Bullseye Asset Management LLC

$5,804,310

0.39%

203,660

Parkman Healthcare Partners LLC

$5,768,884

0.38%

202,417

HSBC HOLDINGS PLC

$5,593,124

0.37%

196,250

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
HSBC Global Asset Management (UK) Ltd	1,163	$33,145	0.6%	Defined
HSBC BANK PLC	195,087	$5,559,979	99.4%	Defined

ACADIAN ASSET MANAGEMENT LLC

$5,312,143

0.35%

186,391

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Acadian Inc	186,391	$5,312,143	100.0%	Defined

Bank of New York Mellon Corp

$4,873,897

0.32%

171,014

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Investments Corporation	76,845	$2,190,082	44.9%	Defined
The Bank of New York Mellon	93,993	$2,678,800	55.0%	Defined
BNY Mellon, NA joint	175	$4,987	0.1%	Defined
Held directly	1	$28	0.0%	Defined

FRED ALGER MANAGEMENT, LLC

$4,849,161

0.32%

170,146

Uniplan Investment Counsel, Inc.

$4,478,233

0.30%

157,131

Nuveen, LLC

$4,346,506

0.29%

152,509

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	4,446	$126,711	2.9%	Defined
TEACHERS ADVISORS, LLC joint	148,063	$4,219,795	97.1%	Defined

RAYMOND JAMES FINANCIAL INC

$4,288,622

0.29%

150,478

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	3,024	$86,184	2.0%	Defined
Raymond James Financial Services Advisors, Inc.	48,497	$1,382,164	32.2%	Defined
ClariVest Asset Management LLC joint	98,957	$2,820,274	65.8%	Defined

WELLINGTON MANAGEMENT GROUP LLP

$4,274,629

0.28%

149,987

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wellington Management Company LLP	48,271	$1,375,723	32.2%	Defined
Wellington Trust Company, NA joint	95,034	$2,708,469	63.4%	Defined
Wellington Management Australia Pty Ltd	6,682	$190,437	4.5%	Defined

Squarepoint Ops LLC

$4,036,255

0.27%

141,623

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Squarepoint Capital LLP	141,623	$4,036,255	100.0%	Defined

AMERICAN CENTURY COMPANIES INC

$3,676,272

0.24%

128,992

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AMERICAN CENTURY INVESTMENT MANAGEMENT INC	128,992	$3,676,272	100.0%	Defined

Man Group plc

$3,618,417

0.24%

126,962

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Numeric Investors LLC	126,962	$3,618,417	100.0%	Defined

GLOBEFLEX CAPITAL L P

$3,398,682

0.23%

119,252

STATE BOARD OF ADMINISTRATION OF FLORIDA RETIREMENT SYSTEM

$3,327,831

0.22%

116,766

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$3,135,882

0.21%

110,031

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	2,086	$59,451	1.9%	Defined
UBS Asset Management Switzerland AG	79,923	$2,277,805	72.6%	Sole
UBS Asset Management (UK) Ltd.	20,497	$584,164	18.6%	Other
Held directly	7,525	$214,462	6.8%	Sole

CITADEL ADVISORS LLC

$3,049,044

0.20%

106,984

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	97,312	$2,773,392	91.0%	Defined
Held directly	9,672	$275,652	9.0%	Defined

STATE OF WISCONSIN INVESTMENT BOARD

$2,872,771

0.19%

100,799

IQ EQ FUND MANAGEMENT (IRELAND) Ltd

$2,826,430

0.19%

99,173

LAZARD ASSET MANAGEMENT LLC

$2,700,289

0.18%

94,747

Kera Capital Partners, Inc.

$2,596,578

0.17%

91,108

Swiss National Bank

$2,490,900

0.17%

87,400

PRUDENTIAL FINANCIAL INC

$2,432,760

0.16%

85,360

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PGIM Quantitative Solutions LLC	85,360	$2,432,760	100.0%	Defined

Schonfeld Strategic Advisors LLC

$2,379,693

0.16%

83,498

Palumbo Wealth Management LLC

$2,363,961

0.16%

82,946

Jump Financial, LLC

$2,359,002

0.16%

82,772

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Leap GP LLC joint	82,772	$2,359,002	100.0%	Defined

Regal Partners Ltd

$2,256,772

0.15%

79,185

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Regal Partners Funds Management Pty Ltd	79,185	$2,256,772	100.0%	Defined

FORMULA GROWTH LTD

$2,223,741

0.15%

78,026

MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.

$2,185,437

0.15%

76,682

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Global X Management CO LLC joint	72,990	$2,080,215	95.2%	Defined
Global X Management (AUS) Ltd joint	3,692	$105,222	4.8%	Defined

CITIGROUP INC

$2,171,044

0.14%

76,177

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	76,177	$2,171,044	100.0%	Defined

PRINCIPAL FINANCIAL GROUP INC

$2,134,307

0.14%

74,888

ARS Investment Partners, LLC

$2,115,697

0.14%

74,235

MILLENNIUM MANAGEMENT LLC

$2,103,556

0.14%

73,809

Gotham Asset Management, LLC

$2,097,001

0.14%

73,579

FRANKLIN RESOURCES INC

$1,960,457

0.13%

68,788

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fiduciary Trust Co International of Pennsylvania	6,750	$192,375	9.8%	Defined
FIDUCIARY TRUST INTERNATIONAL LLC	156	$4,446	0.2%	Defined
FRANKLIN ADVISERS INC	59,882	$1,706,637	87.1%	Defined
FRANKLIN ADVISORY SERVICES LLC	1,575	$44,887	2.3%	Defined
Franklin Templeton International Services S.a r.l	425	$12,112	0.6%	Defined

RHUMBLINE ADVISERS

$1,943,871

0.13%

68,206

ALLIANCEBERNSTEIN L.P.

$1,726,530

0.11%

60,580

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	60,580	$1,726,530	100.0%	Defined

DEUTSCHE BANK AG\

$1,547,179

0.10%

54,287

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DWS INVESTMENT MANAGEMENT AMERICAS, INC	884	$25,194	1.6%	Defined
DWS Investments UK Ltd	53,403	$1,521,985	98.4%	Defined

XTX Topco Ltd

$1,531,818

0.10%

53,748

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	53,748	$1,531,818	100.0%	Defined

CANADA PENSION PLAN INVESTMENT BOARD

$1,504,800

0.10%

52,800

AMERIPRISE FINANCIAL INC

$1,492,430

0.10%

52,366

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	52,281	$1,490,008	99.8%	Defined
Ameriprise Financial Services, LLC	85	$2,422	0.2%	Defined

PRICE T ROWE ASSOCIATES INC /MD/

$1,490,293

0.10%

52,291

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	52,291	$1,490,293	100.0%	Defined

WELLS FARGO & COMPANY/MN

$1,477,440

0.10%

51,840

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	46,786	$1,333,401	90.3%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	5,022	$143,127	9.7%	Defined
WELLS FARGO CLEARING SERVICES, LLC	32	$912	0.1%	Defined

Quantinno Capital Management LP

$1,391,370

0.09%

48,820

Trexquant Investment LP

$1,308,378

0.09%

45,908

Quadrature Capital Ltd

$1,299,286

0.09%

45,589

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Quadrature US Inc.	45,589	$1,299,286	100.0%	Other

Susquehanna Portfolio Strategies, LLC

$1,285,977

0.09%

45,122

VIRTUS ADVISERS, LLC

$1,198,311

0.08%

42,046

JPMORGAN CHASE & CO

$1,150,942

0.08%

40,384

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	28,471	$811,423	70.5%	Defined
J.P. Morgan Investment Management Inc.	4,569	$130,216	11.3%	Defined
J.P. Morgan Securities LLC	296	$8,436	0.7%	Defined
J.P. Morgan Securities plc	357	$10,174	0.9%	Defined
J.P. Morgan SE	6,691	$190,693	16.6%	Defined

EXCHANGE TRADED CONCEPTS, LLC

$1,073,908

0.07%

37,681

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	2,340	$66,690	6.4%	Defined
UBS AG	34,127	$972,619	93.6%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CREATIVE PLANNING TRUST Co LLC	79	$2,251	0.2%	Defined
Held directly	35,104	$1,000,464	99.8%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	34,601	$986,128	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	31,500	$897,750	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
METLIFE INC	29,915	$852,577	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI INVESTMENTS MANAGEMENT CORP	1,413	$40,270	5.8%	Sole
SEI TRUST CO	22,939	$653,761	94.2%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Allspring Funds Management, LLC reporting on behalf of Morgan Stanley Smith Barney, LLC	261	$7,438	1.2%	Defined
Allspring Global Investments, LLC reporting on behalf of UBS ASSET MANAGEMENT AMERICAS INC	21,778	$620,673	98.8%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Magnetar Asset Management LLC	10,866	$309,681	59.5%	Sole
Held directly	7,401	$210,928	40.5%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marshall Wace North America L.P. joint	17,898	$510,093	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Capital	15,057	$429,124	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NEEDHAM EMERGING GROWTH PARTNERS LP	10,000	$285,000	66.7%	Defined
Held directly	5,000	$142,500	33.3%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	14,100	$401,850	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dorset Partners LLC	13,630	$388,455	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MANULIFE FINANCIAL CORP joint	13,403	$381,985	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Amundi Asset Management	11,891	$338,893	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
First Sentier Investors RQI Pty Ltd	10,996	$313,386	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 1	10,878	$310,023	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PNC Bank, National Association	10,678	$304,323	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	9,940	$283,290	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	9,600	$273,600	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	6,597	$188,014	82.7%	Defined
BARCLAYS CAPITAL INC.	1,379	$39,301	17.3%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Advisors Asset Management, Inc.	2,394	$68,229	33.5%	Defined
Blue Trust, Inc.	4,750	$135,375	66.5%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ROCKEFELLER & CO. LLC	6,135	$174,847	96.2%	Defined
ROCKEFELLER TRUST COMPANY, N.A.	242	$6,897	3.8%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FEDERATED GLOBAL INVESTMENT MANAGEMENT CORP	3,977	$113,344	82.6%	Defined
FEDERATED MDTA LLC	838	$23,883	17.4%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
EATON VANCE MANAGEMENT	4,655	$132,667	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Optiver VOF	2,621	$74,698	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
KBC Bank NV joint	2,018	$57,513	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC CMA LTD.	1,546	$44,061	91.7%	Defined
RBC Capital Markets, LLC	139	$3,961	8.2%	Defined
Held directly	1	$28	0.1%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
US BANK NATIONAL ASSOCIATION	1,428	$40,698	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Latour Trading LLC (Latour)	1,330	$37,905	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNP PARIBAS ASSET MANAGEMENT Europe	1,172	$33,402	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nykredit Bank A/S	862	$24,567	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
OSAIC WEALTH, INC.	789	$22,486	98.9%	Defined
Osaic Advisory Services, LLC	9	$256	1.1%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fifth Third Bank, National Association	549	$15,646	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BMSS Wesson Wealth Solutions, LLC	104	$2,964	36.8%	Sole
Held directly	179	$5,101	63.2%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
TORONTO DOMINION BANK	42	$1,197	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	21,900	$380,184	6.9%	Defined
JANUS HENDERSON INVESTORS US LLC	294,619	$5,114,585	93.1%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Brevan Howard US Investment Management, LP	17,604	$305,605	100.0%	Defined

Key facts CIK 1217234 CUSIP 14167L103 13F (30d) 212 filings 208 filers Visit website Investor relations