CYTK · Cytokinetics Inc

$77.18 -0.56 (-0.72%) At close · Aug 20

Market Cap

$10.65B

139.02M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Quarter:

Holders: 484

Total Value: $14,279,388,072

Total Shares: 167,618,129

Institutional Ownership Trend

26 quarters

Superinvestors Holding CYTK

All superinvestors →

Joel Greenblatt $457,214

Holder Concentration

Across 484 13F filers · Δ vs Mar 31, 2026

Top Holder

12.0%

+0.8 pts

Top 5

35.2%

−2.3 pts

Top 10

49.8%

−1.0 pts

HHI

396

Broadly held−2

Quarterly Activity

vs Mar 31, 2026

New

Increased

237

Decreased

107

Sold Out

Unchanged

Holder count: +33

Institutional value: +38.3%

Shares held: +7.0%

Top Buyers

vs Mar 31, 2026

Institution	Added	Now Holds
Mega BlackRock, Inc.	+3,546,351	20,094,362
Large Avoro Capital Advisors LLC	+1,666,666	4,999,999
Mega BANK OF AMERICA CORP /DE/	+1,510,273	4,635,709
Mega SUSQUEHANNA INTERNATIONAL GROUP, LLP	+1,506,907	3,729,658
Mega T. Rowe Price Investment Management, Inc.	+1,194,658	18,718,498

Top Sellers

vs Mar 31, 2026

Institution	Sold	Now Holds
Mega FMR LLC	−5,691,358	5,031,782
Large Caption Management, LLC	−1,344,200	512,400
Mega WELLINGTON MANAGEMENT GROUP LLP	−1,077,719	6,931,770
Mid Integral Health Asset Management, LLC	−1,000,000	1,600,000
Mega UBS Group AG	−930,890	1,171,287

Top New Positions

vs Mar 31, 2026

Institution	Acquired	Value (USD)
Large Holocene Advisors, LP	1,144,579	$97,506,685
Mega NORGES BANK	1,015,522	$86,512,319
Large Hood River Capital Management LLC	469,273	$39,977,366
Mid Polaris Capital Management, LLC	266,900	$22,737,211
Mega Jupiter Topco LLC	244,417	$20,821,884

Top Sold-Out Positions

vs Mar 31, 2026

Institution	Previously Held	Last Value (USD)
Mid Paradigm Biocapital Advisors LP	1,480,857	$97,603,284
Small Ikarian Capital, LLC	1,242,472	$81,891,329
Mid Logos Global Management LP	875,000	$57,671,250
Mid MPM BIOIMPACT LLC	500,000	$32,955,000
Mid GROUP ONE TRADING LLC	363,997	$23,991,042

Filter: New 92 Increased 237 Reduced 107 Sold out 59 Unchanged 48

Showing 100 of 507

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

BlackRock, Inc.

$1,711,838,691

11.99%

20,094,362

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	341,882	$29,124,927	1.7%	Sole
BlackRock Investment Management (Australia) Ltd	305	$25,982	0.0%	Sole
BlackRock Asset Management Canada Ltd	24,461	$2,083,832	0.1%	Sole
BLACKROCK (SINGAPORE) LTD.	8,226	$700,772	0.0%	Sole
BlackRock Investment Management, LLC	667,932	$56,901,127	3.3%	Sole
BLACKROCK ADVISORS LLC	1,413,603	$120,424,839	7.0%	Sole
BlackRock Fund Advisors	11,492,015	$979,004,757	57.2%	Sole
BlackRock Institutional Trust Company, N.A.	4,750,149	$404,665,193	23.6%	Sole
BlackRock Japan Co. Ltd	3,847	$327,725	0.0%	Sole
BlackRock Fund Managers Ltd.	21,059	$1,794,016	0.1%	Sole
BlackRock Investment Management (UK) Ltd.	121,911	$10,385,598	0.6%	Sole
BlackRock (Netherlands) B.V.	3,737	$318,355	0.0%	Sole
BlackRock Asset Management Ireland Ltd	337,916	$28,787,064	1.7%	Sole
BlackRock (Luxembourg) S.A.	556,812	$47,434,814	2.8%	Sole
BlackRock Life Ltd	527	$44,895	0.0%	Sole
BlackRock Asset Management Schweiz AG	18,617	$1,585,982	0.1%	Sole
Aperio Group, LLC	331,047	$28,201,893	1.6%	Sole
SpiderRock Advisors, LLC	316	$26,920	0.0%	Sole

T. Rowe Price Investment Management, Inc.

$1,594,628,844

11.17%

18,718,498

WELLINGTON MANAGEMENT GROUP LLP

$590,517,483

4.14%

6,931,770

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wellington Management Company LLP	5,980,996	$509,521,049	86.3%	Defined
Wellington Managment Canada LLC	64,323	$5,479,676	0.9%	Defined
Wellington Management Hong Kong Ltd	2,225	$189,547	0.0%	Defined
Wellington Management Singapore Pte. Ltd.	77,916	$6,637,664	1.1%	Defined
Wellington Management International Ltd	198,443	$16,905,359	2.9%	Defined
Wellington Trust Company, NA joint	561,357	$47,822,002	8.1%	Defined
Wellington Management Australia Pty Ltd	14,091	$1,200,412	0.2%	Defined
Wellington Management Europe GmbH	32,419	$2,761,774	0.5%	Defined

VANGUARD PORTFOLIO MANAGEMENT LLC

$571,778,156

4.00%

6,711,799

STATE STREET CORP

$553,243,709

3.87%

6,494,233

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	6,494,233	$553,243,709	100.0%	Defined

VANGUARD CAPITAL MANAGEMENT LLC

$521,151,102

3.65%

6,117,515

FMR LLC

$428,657,506

3.00%

5,031,782

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	4,402,341	$375,035,429	87.5%	Defined
FIDELITY MANAGEMENT TRUST CO	384,700	$32,772,593	7.6%	Defined
STRATEGIC ADVISERS LLC	17,128	$1,459,134	0.3%	Defined
Fidelity Institutional Asset Management Trust Co	116,272	$9,905,211	2.3%	Defined
FIAM LLC	110,016	$9,372,263	2.2%	Defined
Fidelity Diversifying Solutions LLC	1,325	$112,876	0.0%	Defined

Avoro Capital Advisors LLC

$425,949,914

2.98%

4,999,999

BANK OF AMERICA CORP /DE/

$394,916,048

2.77%

4,635,709

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	2,954,648	$251,706,463	63.7%	Defined
MERRILL LYNCH, PIERCE, FENNER & SMITH INC.	73,104	$6,227,729	1.6%	Defined
BOFA SECURITIES, INC.	1,478,864	$125,984,424	31.9%	Defined
MANAGED ACCOUNT ADVISORS LLC joint	129,093	$10,997,432	2.8%	Defined

GEODE CAPITAL MANAGEMENT, LLC

$316,267,959

2.21%

3,712,501

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	670,483	$57,118,446	18.1%	Defined
Held directly	3,042,018	$259,149,513	81.9%	Defined

RTW INVESTMENTS, LP

$243,814,035

1.71%

2,862,003

Polar Capital Holdings Plc

$191,988,358

1.34%

2,253,649

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Polar Capital LLP	2,253,649	$191,988,358	100.0%	Defined

PRICE T ROWE ASSOCIATES INC /MD/

$166,882,609

1.17%

1,958,946

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	187,477	$15,971,165	9.6%	Defined
Held directly	1,771,469	$150,911,444	90.4%	Sole

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$161,060,214

1.13%

1,890,600

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	1,890,600	$161,060,214	100.0%	Other

GOLDMAN SACHS GROUP INC

$150,069,678

1.05%

1,761,588

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	634,503	$54,053,310	36.0%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	1,089,940	$92,851,988	61.9%	Defined
GOLDMAN SACHS INTERNATIONAL	11,612	$989,226	0.7%	Defined
GOLDMAN SACHS BANK AG	4,768	$406,185	0.3%	Defined
GOLDMAN SACHS WEALTH SERVICES, L.P.	10,850	$924,311	0.6%	Defined
Goldman Sachs Trust Company, N.A.	9,915	$844,658	0.6%	Defined

Capital Research Global Investors

$143,683,583

1.01%

1,686,625

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CAPITAL GROUP COMPANIES INC joint	1,686,625	$143,683,583	100.0%	Defined

NORTHERN TRUST CORP

$140,337,319

0.98%

1,647,345

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	1,488,257	$126,784,613	90.3%	Other
NORTHERN TRUST GLOBAL INVESTMENTS LTD	75,534	$6,434,741	4.6%	Defined
MASON STREET ADVISORS, LLC joint	63,022	$5,368,844	3.8%	Other
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	20,532	$1,749,121	1.2%	Other

WILLIAM BLAIR INVESTMENT MANAGEMENT, LLC

$136,597,223

0.96%

1,603,442

Integral Health Asset Management, LLC

$136,304,000

0.95%

1,600,000

JPMORGAN CHASE & CO

$135,357,961

0.95%

1,588,895

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	201,789	$17,190,404	12.7%	Defined
J.P. Morgan Investment Management Inc.	1,272,205	$108,379,143	80.1%	Defined
J.P. Morgan Securities LLC	804	$68,492	0.1%	Defined
JPMorgan Asset Management (UK) Ltd	113,494	$9,668,553	7.1%	Defined
55I, LLC	6	$511	0.0%	Other
J.P. MORGAN MANSART MANAGEMENT Ltd	597	$50,858	0.0%	Defined

Point72 Asset Management, L.P.

$135,257,185

0.95%

1,587,712

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$133,388,883

0.93%

1,565,781

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$131,998,327

0.92%

1,549,458

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	1,549,458	$131,998,327	100.0%	Other

ARMISTICE CAPITAL, LLC

$130,340,700

0.91%

1,530,000

MORGAN STANLEY

$128,663,982

0.90%

1,510,318

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	115	$9,796	0.0%	Defined
MORGAN STANLEY & CO. LLC	162,419	$13,836,474	10.8%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	695,660	$59,263,275	46.1%	Defined
MORGAN STANLEY AIP GP LP	11,449	$975,339	0.8%	Defined
MORGAN STANLEY INVESTMENT MANAGEMENT INC	3,894	$331,729	0.3%	Defined
Morgan Stanley Investment Management LTD	9,264	$789,199	0.6%	Defined
MORGAN STANLEY SMITH BARNEY LLC	265,663	$22,631,830	17.6%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	320,956	$27,342,240	21.3%	Defined
EATON VANCE MANAGEMENT	190	$16,186	0.0%	Defined
Calvert Research & Management	40,708	$3,467,914	2.7%	Defined

PRINCIPAL FINANCIAL GROUP INC

$111,740,655

0.78%

1,311,664

Capital International Investors

$111,245,191

0.78%

1,305,848

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CAPITAL GROUP COMPANIES INC joint	1,305,848	$111,245,191	100.0%	Defined

Invesco Ltd.

$105,237,162

0.74%

1,235,323

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	1,029,996	$87,745,357	83.4%	Defined
Invesco Trust Co joint	52,635	$4,483,975	4.3%	Defined
Invesco Asset Management Limited joint	60,092	$5,119,236	4.9%	Defined
Invesco Capital Management LLC	92,600	$7,888,594	7.5%	Defined

ARMISTICE CAPITAL, LLC

$102,342,324

0.72%

1,201,342

UBS Group AG

$99,781,937

0.70%

1,171,287

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	123,583	$10,528,035	10.6%	Defined
UBS AG	980,245	$83,507,071	83.7%	Defined
UBS Switzerland AG	67,041	$5,711,222	5.7%	Defined
UBS Securities LLC	418	$35,609	0.0%	Defined

Pictet Asset Management Holding SA

$98,859,842

0.69%

1,160,463

Frazier Life Sciences Management, L.P.

$98,810,092

0.69%

1,159,879

Bank of New York Mellon Corp

$98,027,870

0.69%

1,150,697

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNY Mellon Trust of Delaware joint	1,560	$132,895	0.1%	Defined
Mellon Investments Corporation joint	368,252	$31,371,386	32.0%	Defined
BNY Mellon Investment Adviser, Inc.	50,235	$4,279,519	4.4%	Defined
BNY Mellon Securities Corporation	3,485	$296,887	0.3%	Defined
The Bank of New York Mellon	402,664	$34,302,946	35.0%	Defined
BNY Mellon, NA joint	89,454	$7,620,585	7.8%	Defined
BNY Mellon Advisors, Inc. joint	6,591	$561,487	0.6%	Other
Newton Investment Management North America, LLC joint	228,455	$19,462,080	19.9%	Defined
Held directly	1	$85	0.0%	Defined

Holocene Advisors, LP

$97,506,685

0.68%

1,144,579

Woodline Partners LP

$93,769,314

0.66%

1,100,708

D. E. Shaw & Co., Inc.

$89,287,894

0.63%

1,048,103

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	568,302	$48,413,647	54.2%	Defined
D. E. Shaw Investment Management, L.L.C. joint	479,801	$40,874,247	45.8%	Defined

NORGES BANK

$86,512,319

0.61%

1,015,522

Royalty Pharma Sub-Manager, LLC

$83,519,594

0.58%

980,392

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Royalty Pharma Manager, LLC joint	980,392	$83,519,594	100.0%	Defined

NOMURA ASSET MANAGEMENT INTERNATIONAL INC.

$81,937,019

0.57%

961,815

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NOMURA HOLDINGS INC joint	961,815	$81,937,019	100.0%	Defined

GW&K Investment Management, LLC

$78,043,155

0.55%

916,107

VANGUARD FIDUCIARY TRUST CO

$77,218,175

0.54%

906,423

CITADEL ADVISORS LLC

$77,062,874

0.54%

904,600

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	904,600	$77,062,874	100.0%	Defined

FRANKLIN RESOURCES INC

$75,911,698

0.53%

891,087

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FIDUCIARY TRUST CO INTERNATIONAL	35	$2,981	0.0%	Defined
Fiduciary Trust International of California	12	$1,022	0.0%	Defined
Fiduciary Trust International of the South	92	$7,837	0.0%	Defined
FIDUCIARY TRUST INTERNATIONAL LLC	1,692	$144,141	0.2%	Defined
FRANKLIN ADVISERS INC	776,383	$66,140,067	87.1%	Defined
PUTNAM INVESTMENT MANAGEMENT LLC	105,616	$8,997,427	11.9%	Defined
PUTNAM ADVISORY CO LLC	7,257	$618,223	0.8%	Defined

BNP PARIBAS ASSET MANAGEMENT Holding S.A.

$70,521,132

0.49%

827,810

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNP PARIBAS ASSET MANAGEMENT Europe	1,968	$167,653	0.2%	Defined
BNP Paribas Asset Management USA, Inc.	825,842	$70,353,479	99.8%	Defined

Melqart Asset Management (UK) Ltd

$65,385,454

0.46%

767,525

Siren, L.L.C.

$64,655,120

0.45%

758,952

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invus Group, LLC	758,952	$64,655,120	100.0%	Defined

Fisher Asset Management, LLC

$63,355,547

0.44%

743,697

Foresite Capital Management IV, LLC

$60,884,867

0.43%

714,695

Balyasny Asset Management L.P.

$60,627,678

0.42%

711,676

Deep Track Capital, LP

$59,633,000

0.42%

700,000

JANE STREET GROUP, LLC

$59,633,000

0.42%

700,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	700,000	$59,633,000	100.0%	Defined

BNP PARIBAS ARBITRAGE, SA

$58,702,469

0.41%

689,077

Nationale-Nederlanden Powszechne Towarzystwo Emerytalne S.A.

$56,251,808

0.39%

660,310

CITADEL ADVISORS LLC

$54,602,103

0.38%

640,945

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	9	$766	0.0%	Defined
Held directly	640,936	$54,601,337	100.0%	Defined

Qube Research & Technologies Ltd

$48,774,256

0.34%

572,535

BROWN ADVISORY INC

$48,614,865

0.34%

570,664

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BROWN INVESTMENT ADVISORY & TRUST CO	2,400	$204,456	0.4%	Defined
Brown Advisory LLC	536,619	$45,714,572	94.0%	Defined
NEW YORK LIFE INVESTMENT MANAGEMENT LLC	31,645	$2,695,837	5.5%	Defined

EVENTIDE ASSET MANAGEMENT, LLC

$48,081,065

0.34%

564,398

Affinity Asset Advisors, LLC

$46,854,500

0.33%

550,000

MASSACHUSETTS FINANCIAL SERVICES CO /MA/

$46,798,956

0.33%

549,348

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MFS Heritage Trust CO	5,832	$496,828	1.1%	Defined
Held directly	543,516	$46,302,128	98.9%	Sole

FEDERATED HERMES, INC.

$45,834,093

0.32%

538,022

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FEDERATED EQUITY MANAGEMENT CO OF PENNSYLVANIA	11,140	$949,016	2.1%	Defined
FEDERATED GLOBAL INVESTMENT MANAGEMENT CORP	518,813	$44,197,679	96.4%	Defined
FEDERATED MDTA LLC	8,069	$687,398	1.5%	Defined

ALLIANCEBERNSTEIN L.P.

$45,661,243

0.32%

535,993

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	535,993	$45,661,243	100.0%	Defined

First Turn Management, LLC

$44,930,228

0.31%

527,412

AMERICAN CENTURY COMPANIES INC

$44,162,325

0.31%

518,398

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AMERICAN CENTURY INVESTMENT MANAGEMENT INC	518,398	$44,162,325	100.0%	Defined

Caption Management, LLC

$43,651,356

0.31%

512,400

Checkpoint Capital L.P.

$40,465,250

0.28%

475,000

Hood River Capital Management LLC

$39,977,366

0.28%

469,273

EMERALD ADVISERS, LLC

$39,013,271

0.27%

457,956

ADAR1 Capital Management, LLC

$39,009,097

0.27%

457,907

AMERIPRISE FINANCIAL INC

$38,748,756

0.27%

454,851

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	446,179	$38,009,989	98.1%	Defined
Ameriprise Financial Services, LLC	1,008	$85,871	0.2%	Defined
Ameriprise Bank, FSB	64	$5,452	0.0%	Defined
Columbia Threadneedle Management Ltd	7,600	$647,444	1.7%	Defined

MARSHALL WACE, LLP

$38,460,473

0.27%

451,467

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marshall Wace North America L.P. joint	451,467	$38,460,473	100.0%	Other

Nuveen, LLC

$38,095,689

0.27%

447,185

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	33,661	$2,867,580	7.5%	Defined
TIAA CREF INVESTMENT MANAGEMENT LLC joint	31,400	$2,674,966	7.0%	Defined
TEACHERS ADVISORS, LLC joint	382,124	$32,553,143	85.5%	Defined

Candriam S.C.A.

$37,744,792

0.26%

443,066

RHUMBLINE ADVISERS

$35,933,056

0.25%

421,799

CITADEL ADVISORS LLC

$35,882,028

0.25%

421,200

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	421,200	$35,882,028	100.0%	Defined

Handelsbanken Fonder AB

$34,501,950

0.24%

405,000

GILDER GAGNON HOWE & CO LLC

$34,096,871

0.24%

400,245

TD Asset Management Inc

$32,809,224

0.23%

385,130

VOYA INVESTMENT MANAGEMENT LLC

$31,409,382

0.22%

368,698

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	368,698	$31,409,382	100.0%	Defined

GSK plc

$29,001,572

0.20%

340,434

CITIGROUP INC

$27,510,576

0.19%

322,932

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	322,932	$27,510,576	100.0%	Defined

TWO SIGMA INVESTMENTS, LP

$27,040,243

0.19%

317,411

E. Ohman J:or Asset Management AB

$25,949,811

0.18%

304,611

Russell Investments Group, Ltd.

$25,690,403

0.18%

301,566

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC	164,911	$14,048,767	54.7%	Defined
Russell Investments Canada Limited	520	$44,298	0.2%	Defined
Russell Investments Capital, LLC	68,471	$5,833,044	22.7%	Defined
Russell Investments France SAS	20,463	$1,743,242	6.8%	Defined
Russell Investments Limited	501	$42,680	0.2%	Defined
Russell Investments Trust Company joint	46,700	$3,978,372	15.5%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$24,671,024

0.17%

289,600

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	289,600	$24,671,024	100.0%	Other

Trails Edge Capital Partners, LP

$24,542,301

0.17%

288,089

BANK OF MONTREAL /CAN/

$24,337,759

0.17%

285,688

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BMO Asset Management Inc.	9,765	$831,880	3.4%	Defined
BMO Bank N.A.	3,681	$313,584	1.3%	Defined
BMO Family Office, LLC	898	$76,500	0.3%	Defined
BMO NESBITT BURNS INC.	744	$63,381	0.3%	Defined
Stoker Ostler Wealth Advisors, Inc.	600	$51,114	0.2%	Defined
Held directly	270,000	$23,001,300	94.5%	Defined

EMERALD MUTUAL FUND ADVISERS TRUST

$23,833,521

0.17%

279,769

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$23,259,507

0.16%

273,031

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	3,817	$325,170	1.4%	Defined
UBS Asset Management Switzerland AG	182,982	$15,588,236	67.0%	Defined
UBS Asset Management (UK) Ltd.	63,803	$5,435,376	23.4%	Defined
Held directly	22,429	$1,910,725	8.2%	Sole

Polaris Capital Management, LLC

$22,737,211

0.16%

266,900

SEGALL BRYANT & HAMILL, LLC

$21,899,196

0.15%

257,063

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NEW YORK LIFE INVESTMENT MANAGEMENT LLC	257,063	$21,899,196	100.0%	Sole

Alyeska Investment Group, L.P.

$21,297,500

0.15%

250,000

ExodusPoint Capital Management, LP

$21,234,374

0.15%

249,259

Capital World Investors

$21,196,549

0.15%

248,815

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CAPITAL GROUP COMPANIES INC joint	248,815	$21,196,549	100.0%	Defined

Jupiter Topco LLC

$20,821,884

0.15%

244,417

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	49,864	$4,247,914	20.4%	Defined
JANUS HENDERSON INVESTORS US LLC	194,553	$16,573,970	79.6%	Defined

Walleye Capital LLC

$20,422,428

0.14%

239,728

Sarissa Capital Management LP

$19,476,393

0.14%

228,623

Swiss National Bank

$18,946,256

0.13%

222,400

SPHERA FUNDS MANAGEMENT LTD.

$18,740,692

0.13%

219,987

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Sphera Global Healthcare Management LP joint	219,987	$18,740,692	100.0%	Defined

DEUTSCHE BANK AG\

$18,447,039

0.13%

216,540

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Deutsche Bank Trust Co Americas	9	$766	0.0%	Defined
DWS Investment GmbH	67,812	$5,776,904	31.3%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC	248	$21,127	0.1%	Defined
DWS Investments UK Ltd	146,184	$12,453,414	67.5%	Defined
DBX Advisors LLC	188	$16,015	0.1%	Defined
DWS International GmbH	2,099	$178,813	1.0%	Defined

PRUDENTIAL FINANCIAL INC

$18,276,236

0.13%

214,535

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PGIM Quantitative Solutions LLC	187,035	$15,933,511	87.2%	Defined
Prudential Trust Co	27,500	$2,342,725	12.8%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
IMPAX ASSET MANAGEMENT LLC	165,000	$14,056,350	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	78,652	$6,700,363	47.8%	Defined
Legal & General Investment Management Ltd joint	86,036	$7,329,406	52.2%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Compound Planning, Inc.	2	$170	0.0%	Other
Arax Advisory Partners	174	$14,823	0.1%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	27	$2,300	0.0%	Other
Held directly	161,700	$13,775,223	99.9%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 1 joint	157,100	$13,383,349	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MANULIFE FINANCIAL CORP joint	156,477	$13,330,275	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	118,298	$10,077,806	77.5%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	7,622	$649,318	5.0%	Defined
WELLS FARGO CLEARING SERVICES, LLC	26,615	$2,267,331	17.4%	Defined
Wells Fargo Securities, LLC	6	$511	0.0%	Defined
WELLS FARGO INVESTMENT INSTITUTE, INC.	51	$4,344	0.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
HSBC Bank USA, N.A.	182	$15,504	0.1%	Defined
HSBC Global Asset Management (UK) Ltd	3,564	$303,617	2.4%	Defined
HSBC GLOBAL ASSET MANAGEMENT (FRANCE)	147,139	$12,534,771	97.4%	Defined
HSBC SECURITIES USA INC	240	$20,445	0.2%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	149,500	$12,735,905	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CREATIVE PLANNING TRUST Co LLC	3,807	$324,318	2.7%	Defined
Held directly	138,132	$11,767,465	97.3%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Schroder Investment Management Limited	139,190	$11,857,596	99.4%	Sole
Schroder Investment Management North America Inc.	909	$77,437	0.6%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
METLIFE INC	140,046	$11,930,518	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
abrdn Inc.	119,435	$10,174,667	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Universal-Investment-Gesellschaft mbH	116,828	$9,952,577	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Sphera Global Healthcare Management LP joint	100,000	$8,519,000	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI INVESTMENTS MANAGEMENT CORP	1,700	$144,823	1.7%	Sole
SEI TRUST CO	96,462	$8,217,597	98.3%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	24,635	$2,098,655	27.9%	Defined
Raymond James Financial Services Advisors, Inc.	9,389	$799,848	10.6%	Defined
CHARTWELL INVESTMENT PARTNERS, LLC	54,235	$4,620,279	61.4%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	80,743	$6,878,496	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	40,367	$3,438,864	57.1%	Defined
BARCLAYS CAPITAL INC.	28,970	$2,467,954	41.0%	Defined
BARCLAYS CAPITAL SECURITIES LTD	60	$5,111	0.1%	Defined
Barclays Investment Solutions Ltd	1,245	$106,061	1.8%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wabon Partners, L.P.	66,900	$5,699,211	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
INTERNAL MGMT S&P400 INDEX	37,226	$3,171,282	56.1%	Sole
WELLINGTON SMALL CAP EQUITY	25,294	$2,154,795	38.1%	Sole
MED S&P 400 STOCK INDEX	3,812	$324,744	5.7%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Global X Management CO LLC joint	60,916	$5,189,434	92.7%	Defined
Global X Management (AUS) Ltd joint	4,781	$407,293	7.3%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
EFG Asset Management (North America) Corp.	57,905	$4,932,926	100.0%	Defined
EFG Private Wealth Management (Americas) Corp.	24	$2,044	0.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STIFEL NICOLAUS & CO INC \MO\	41,145	$3,505,142	72.4%	Defined
Stifel Independent Advisors, LLC	69	$5,878	0.1%	Defined
1919 Investment Counsel, LLC	15,510	$1,321,296	27.3%	Defined
Held directly	110	$9,370	0.2%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	56,000	$4,770,640	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Natixis Investment Managers, L.P.	53,358	$4,545,568	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC	11,834	$1,008,138	23.0%	Other
Held directly	39,609	$3,374,290	77.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
OSAIC WEALTH, INC.	2,090	$178,047	4.2%	Defined
CW Advisors, LLC	47,696	$4,063,222	95.2%	Sole
Osaic Advisory Services, LLC	301	$25,642	0.6%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ADVISOR PARTNERS II, LLC	43,873	$3,737,540	92.1%	Defined
Held directly	3,760	$320,314	7.9%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PEAK6 Investments LLC joint	46,200	$3,935,778	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Capital	39,014	$3,323,602	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
EATON VANCE MANAGEMENT	36,992	$3,151,348	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC CMA LTD.	5,176	$440,943	14.2%	Defined
RBC Capital Markets, LLC	11,113	$946,716	30.5%	Defined
RBC Dominion Securities Inc.	12,887	$1,097,843	35.3%	Defined
RBC Trust Co (Delaware) Ltd	6,750	$575,032	18.5%	Defined
RBC Rochdale, LLC	199	$16,952	0.5%	Defined
BREWIN DOLPHIN WEALTH MANAGEMENT LTD	170	$14,482	0.5%	Defined
Held directly	180	$15,334	0.5%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wabon Partners, L.P.	34,653	$2,952,089	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NOMURA SECURITIES INTERNATIONAL INC	34,000	$2,896,460	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	33,100	$2,819,789	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GUGGENHEIM FUNDS DISTRIBUTORS, LLC	4,505	$383,780	14.2%	Defined
SECURITY INVESTORS LLC	27,153	$2,313,164	85.8%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHWESTERN MUTUAL LIFE INSURANCE CO	30,541	$2,601,787	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fifth Third Bank, National Association	30,303	$2,581,512	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Edward D. Jones & Co., L.P.	26,593	$2,265,457	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Landscape High Leverage Fund, LP	4,698	$400,222	19.0%	Defined
LANDSCAPE OFFSHORE HL FUND LTD.	12,828	$1,092,817	51.9%	Defined
Held directly	7,208	$614,049	29.1%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Quadrature US Inc.	20,162	$1,717,600	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Bellevue Asset Management AG	19,007	$1,619,206	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
J. Safra Sarasin Fund Management (Luxembourg) S.A.	18,211	$1,551,395	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AHL Partners LLP	14,012	$1,193,682	80.6%	Defined
Numeric Investors LLC	3,381	$288,027	19.4%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ALLSTATE INVESTMENTS LLC	16,889	$1,438,773	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Blue Trust, Inc.	15,042	$1,281,427	94.6%	Defined
Held directly	856	$72,922	5.4%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DNB FUND	12,676	$1,079,868	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manufacturers & Traders Trust Company joint	875	$74,541	7.0%	Defined
Wilmington Trust Investment Advisors, Inc. joint	177	$15,077	1.4%	Defined
Wilmington Trust National Association joint	8,177	$696,598	65.5%	Defined
Wilmington Trust Company	3,259	$277,634	26.1%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BOHRER NATHANIEL joint	12,400	$1,056,356	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Amundi Asset Management	12,089	$1,029,861	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Brevan Howard US Investment Management, LP	8,006	$682,031	67.3%	Defined
Brevan Howard (Hong Kong) Ltd	3,885	$330,963	32.7%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CIBC Private Wealth Advisors, Inc.	1,999	$170,294	17.4%	Defined
CIBC National Trust Co	2,159	$183,925	18.8%	Defined
CIBC WORLD MARKETS CORP	7,303	$622,142	63.7%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Latour Trading LLC (Latour)	10,305	$877,882	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
VOYA INVESTMENTS, LLC	10,000	$851,900	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PNC Bank, National Association	9,318	$793,799	94.0%	Other
PNC Delaware Trust Company	241	$20,530	2.4%	Defined
PNC Wealth Management LLC	351	$29,901	3.5%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GW&K Investment Management	9,831	$837,502	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Desjardins Global Asset Management Inc.	8,659	$737,660	97.8%	Defined
Desjardins Securities inc.	195	$16,612	2.2%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Arrowpoint Investment Partners (Hong Kong) Ltd	8,836	$752,738	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Pacific Investment Management Company LLC	157	$13,374	1.8%	Defined
Allianz Global Invesstors Taiwan Ltd.	8,500	$724,115	98.2%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Allspring Funds Management, LLC reporting on behalf of Morgan Stanley Smith Barney, LLC	770	$65,596	10.2%	Defined
Allspring Global Investments, LLC reporting on behalf of UBS ASSET MANAGEMENT AMERICAS INC	6,761	$575,969	89.8%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
TRUIST BANK /NC/	3,960	$337,351	57.4%	Defined
Truist Advisory Services, Inc.	2,944	$250,799	42.6%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Aperio Group, LLC	6,812	$580,314	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
First Sentier Investors RQI Pty Ltd	6,593	$561,657	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	1,233	$105,039	20.3%	Defined
Jane Street Global Trading, LLC	4,851	$413,256	79.7%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CAPITAL GROUP COMPANIES INC joint	5,890	$501,769	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fiduciary Trust Co of New England	5,144	$438,217	88.8%	Defined
Held directly	646	$55,032	11.2%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Alpha Planning & Financial Services	56	$4,770	1.1%	Other
Arcadia Wealth Management, LLC	17	$1,448	0.3%	Other
B.O.S.S. Retirement Advisors, LLC	98	$8,348	1.8%	Other
BHC Wealth Management	70	$5,963	1.3%	Other
Blackston Financial Advisory Group, LLC	143	$12,182	2.7%	Other
Capital A Wealth Management, LLC	453	$38,591	8.5%	Other
Capstar Financial Services	30	$2,555	0.6%	Other
Family Legacy Planning	7	$596	0.1%	Other
FSA Advisors, Inc.	24	$2,044	0.5%	Other
Fletcher Financial Group	9	$766	0.2%	Other
Pineridge Advisors LLC	238	$20,275	4.5%	Other
Harlow Wealth Managment, Inc.	184	$15,674	3.5%	Other
Hobart Private Capital, LLC	435	$37,057	8.2%	Other
HOWARD BAILEY SECURITIES, LLC	356	$30,327	6.7%	Other
Mach-1 Financial Group, LLC	158	$13,460	3.0%	Other
Montgomery Financial Services	31	$2,640	0.6%	Other
POM Investment Strategies, LLC	444	$37,824	8.4%	Other
Retirement Income Strategies	50	$4,259	0.9%	Other
RSG Investments LLC	13	$1,107	0.2%	Other
Strategy Financial Group	178	$15,163	3.3%	Other
The Holland Group	31	$2,640	0.6%	Other
The Lifewealth Group	116	$9,882	2.2%	Other
Lloyd Advisory Services, LLC.	35	$2,981	0.7%	Other
Held directly	2,140	$182,305	40.3%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
COMMERCE BANCSHARES INC /MO/	5,235	$445,968	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nykredit Bank A/S	5,213	$444,095	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
US BANK NATIONAL ASSOCIATION	3,740	$318,610	93.7%	Other
U.S. BANCORP ADVISORS, LLC	250	$21,297	6.3%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Blackstone Alternative Asset Management L.P.	3,629	$309,154	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
KBC Bank NV joint	3,470	$295,609	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 1	3,452	$294,075	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ROCKEFELLER & CO. LLC	492	$41,913	15.5%	Defined
ROCKEFELLER TRUST COMPANY, N.A.	2,057	$175,235	64.6%	Defined
Rockefeller Trust Co (Delaware)	283	$24,108	8.9%	Defined
ROCKEFELLER FINANCIAL LLC	351	$29,901	11.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Orion Custom Indexing	1,801	$153,427	58.1%	Other
AQR Capital Management	101	$8,604	3.3%	Other
Parametric	50	$4,259	1.6%	Other
Natixis UMA	76	$6,474	2.5%	Other
Aperio	154	$13,119	5.0%	Other
Vanguard Direct Indexing	45	$3,833	1.5%	Other
Dimensional Fund Advisors	1	$85	0.0%	Other
Held directly	870	$74,115	28.1%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 1	2,991	$254,803	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
TOWNSQUARE CAPITAL LLC	2,957	$251,906	98.9%	Other
Held directly	33	$2,811	1.1%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
O'Shaughnessy	3	$255	0.1%	Other
Assetmark	66	$5,622	2.3%	Other
Orion	2,658	$226,435	91.4%	Other
Held directly	181	$15,419	6.2%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Magnetar Asset Management LLC	1,317	$112,195	46.6%	Sole
Held directly	1,512	$128,807	53.4%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Squarepoint Capital LLP	2,655	$226,179	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AQR Flex joint	2,593	$220,897	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Intesa Sanpaolo S.p.A.	2,337	$199,089	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BESSEMER TRUST CO	935	$79,652	47.4%	Other
BESSEMER TRUST Co NATIONAL ASSOCIATION	788	$67,128	39.9%	Defined
Bessemer Trust CO of Delaware, N.A.	250	$21,297	12.7%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Acadian Inc	952	$81,100	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BMSS Wesson Wealth Solutions, LLC	510	$43,445	56.5%	Sole
Held directly	392	$33,393	43.5%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Optiver VOF	660	$56,225	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MITSUBISHI UFJ FINANCIAL GROUP INC	589	$50,176	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
National Bank Financial Inc.	159	$13,545	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JOHNSON BANK	35	$2,981	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SOUTHSTATE PRIVATE CAPITAL MANAGEMENT LLC	15	$1,277	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PARAMETRICA ASSET MANAGEMENT LTD.	9	$766	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DIMENSIONAL FUND ADVISORS LP	1	$85	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	53,264	$3,510,630	18.9%	Defined
JANUS HENDERSON INVESTORS US LLC	228,857	$15,083,964	81.1%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JEFFERIES LLC	50,000	$3,295,500	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dorset Partners LLC	23,644	$1,558,376	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Daiwa America Strategic Advisors Corp	22,200	$1,463,202	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marex Financial	11,300	$744,783	100.0%	Defined
Held directly	1	$65	0.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
HANTZ FINANCIAL SERVICES, INC. joint	10,581	$697,393	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
55I, LLC	3	$197	0.5%	Sole
Fidelity Institutional Asset Management Trust Co	4	$263	0.7%	Sole
Held directly	550	$36,250	98.7%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Rakuten Group, Inc. joint	170	$11,204	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wolverine Holdings, LLC	120	$7,909	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Frost Investment Services, LLC	35	$2,306	100.0%	Defined

Key facts CIK 1061983 CUSIP 23282W605 13F (30d) 443 filings 439 filers Visit website Investor relations