DYN · Dyne Therapeutics, Inc.

$26.20 +0.22 (+0.83%)

Market Cap

$4.85B

186.75M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Quarter:

Holders: 246

Total Value: $4,234,270,168

Total Shares: 190,707,075

Institutional Ownership Trend

24 quarters

Holder Concentration

Across 246 13F filers · Δ vs Mar 31, 2026

Top Holder

16.2%

−0.1 pts

Top 5

38.8%

−0.6 pts

Top 10

53.9%

−1.0 pts

HHI

491

Broadly held−13

Quarterly Activity

vs Mar 31, 2026

New

Increased

Decreased

Sold Out

Unchanged

Holder count: +16

Institutional value: +24.1%

Shares held: +1.4%

Top Buyers

vs Mar 31, 2026

Institution	Added	Now Holds
Mega FMR LLC	+2,298,020	11,207,000
Mega PRICE T ROWE ASSOCIATES INC /MD/	+1,608,433	3,936,806
Large MARSHALL WACE, LLP	+1,405,770	1,488,917
Mid Logos Global Management LP	+1,150,000	2,550,000
Mega DIMENSIONAL FUND ADVISORS LP	+994,890	1,946,479

Top Sellers

vs Mar 31, 2026

Institution	Sold	Now Holds
Small Atlas Venture Life Science Advisors, LLC	−2,330,366	6,800,099
Mid PERCEPTIVE ADVISORS LLC	−1,979,604	2,689,235
Large ADAGE CAPITAL PARTNERS GP, L.L.C.	−850,000	150,000
Small FCPM III SERVICES B.V.	−796,130	3,848,256
Mid BRAIDWELL LP	−760,400	921,598

Top New Positions

vs Mar 31, 2026

Institution	Acquired	Value (USD)
Mega Jupiter Topco LLC	11,997,738	$266,469,760
Mid NOVO HOLDINGS A/S	960,394	$21,330,350
Mid Granahan Investment Management, LLC	338,565	$7,519,528
Large Alyeska Investment Group, L.P.	245,278	$5,447,624
Large ALPS ADVISORS INC	167,082	$3,710,891

Top Sold-Out Positions

vs Mar 31, 2026

Institution	Previously Held	Last Value (USD)
Mega JANUS HENDERSON GROUP PLC	13,647,376	$247,426,926
Small STEMPOINT CAPITAL LP	506,075	$9,175,139
Mega Qube Research & Technologies Ltd	253,153	$4,589,663
Mid PDT Partners, LLC	182,986	$3,317,536
Mid Burkehill Global Management, LP	155,000	$2,810,150

Filter: New 49 Increased 89 Reduced 75 Sold out 33 Unchanged 33

Showing 100 of 252

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

T. Rowe Price Investment Management, Inc.

$686,561,894

16.21%

30,912,287

BlackRock, Inc.

$275,500,161

6.51%

12,404,330

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	145,123	$3,223,181	1.2%	Sole
BlackRock Investment Management (Australia) Ltd	194	$4,308	0.0%	Sole
BlackRock Asset Management Canada Ltd	27,909	$619,858	0.2%	Sole
BlackRock Investment Management, LLC	390,794	$8,679,534	3.2%	Sole
BLACKROCK ADVISORS LLC	835,407	$18,554,389	6.7%	Sole
BlackRock Fund Advisors	6,277,366	$139,420,298	50.6%	Sole
BlackRock Institutional Trust Company, N.A.	4,300,173	$95,506,842	34.7%	Sole
BlackRock Fund Managers Ltd.	20,784	$461,612	0.2%	Sole
BlackRock Investment Management (UK) Ltd.	46,078	$1,023,392	0.4%	Sole
BlackRock Asset Management Ireland Ltd	235,507	$5,230,610	1.9%	Sole
BlackRock Asset Management Schweiz AG	17,685	$392,783	0.1%	Sole
Aperio Group, LLC	107,189	$2,380,667	0.9%	Sole
SpiderRock Advisors, LLC	121	$2,687	0.0%	Sole

Jupiter Topco LLC

$266,469,760

6.29%

11,997,738

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	93,770	$2,082,631	0.8%	Defined
JANUS HENDERSON INVESTORS US LLC	11,903,968	$264,387,129	99.2%	Defined

FMR LLC

$248,907,468

5.88%

11,207,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	7,213,053	$160,201,907	64.4%	Defined
FIDELITY MANAGEMENT TRUST CO	3,992,775	$88,679,532	35.6%	Defined
STRATEGIC ADVISERS LLC	31	$688	0.0%	Defined
FIAM LLC	1,141	$25,341	0.0%	Defined

STATE STREET CORP

$165,648,021

3.91%

7,458,263

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	7,458,263	$165,648,021	100.0%	Defined

Orbis Allan Gray Ltd

$155,615,297

3.68%

7,006,542

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ORBIS INVESTMENT MANAGEMENT LTD	6,983,862	$155,111,575	99.7%	Defined
Held directly	22,680	$503,722	0.3%	Defined

Atlas Venture Life Science Advisors, LLC

$151,030,198

3.57%

6,800,099

VANGUARD CAPITAL MANAGEMENT LLC

$140,166,621

3.31%

6,310,969

TCG Crossover Management, LLC

$102,502,348

2.42%

4,615,144

VANGUARD PORTFOLIO MANAGEMENT LLC

$89,915,585

2.12%

4,048,428

PRICE T ROWE ASSOCIATES INC /MD/

$87,436,460

2.06%

3,936,806

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	195,776	$4,348,184	5.0%	Defined
Held directly	3,741,030	$83,088,276	95.0%	Sole

FCPM III SERVICES B.V.

$85,469,765

2.02%

3,848,256

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Forbion IV Management B.V.	2,421,886	$53,790,088	62.9%	Defined
Held directly	1,426,370	$31,679,677	37.1%	Defined

GEODE CAPITAL MANAGEMENT, LLC

$83,141,513

1.96%

3,743,427

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	533,493	$11,848,879	14.3%	Defined
Held directly	3,209,934	$71,292,634	85.7%	Defined

Vestal Point Capital, LP

$74,958,750

1.77%

3,375,000

ORBIMED ADVISORS LLC

$71,864,541

1.70%

3,235,684

Frazier Life Sciences Management, L.P.

$70,029,951

1.65%

3,153,082

JPMORGAN CHASE & CO

$69,998,276

1.65%

3,151,656

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
J.P. Morgan Trust Co of Delaware	2,564	$56,945	0.1%	Defined
JPMorgan Chase Bank, N.A.	93,080	$2,067,306	3.0%	Defined
J.P. Morgan Investment Management Inc.	1,519,693	$33,752,381	48.2%	Defined
J.P. Morgan Securities LLC	25,861	$574,372	0.8%	Defined
JPMorgan Asset Management (UK) Ltd	1,510,458	$33,547,272	47.9%	Defined

ARMISTICE CAPITAL, LLC

$68,567,289

1.62%

3,087,226

PERCEPTIVE ADVISORS LLC

$59,727,909

1.41%

2,689,235

Logos Global Management LP

$56,635,500

1.34%

2,550,000

BANK OF AMERICA CORP /DE/

$49,603,768

1.17%

2,233,398

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	1,428,443	$31,725,719	64.0%	Defined
BOFA SECURITIES, INC.	635,544	$14,115,432	28.5%	Defined
MANAGED ACCOUNT ADVISORS LLC joint	169,411	$3,762,617	7.6%	Defined

Venrock Adviser, LLC

$48,282,607

1.14%

2,173,913

GOLDMAN SACHS GROUP INC

$47,911,300

1.13%

2,157,195

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	1,874,159	$41,625,071	86.9%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	283,036	$6,286,229	13.1%	Defined

Siren, L.L.C.

$47,287,866

1.12%

2,129,125

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invus Group, LLC	2,129,125	$47,287,866	100.0%	Defined

FRANKLIN RESOURCES INC

$45,104,489

1.07%

2,030,819

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FIDUCIARY TRUST INTERNATIONAL LLC	68	$1,510	0.0%	Defined
FRANKLIN ADVISERS INC	2,030,751	$45,102,979	100.0%	Defined

DIMENSIONAL FUND ADVISORS LP

$43,231,297

1.02%

1,946,479

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	10,560	$234,537	0.5%	Defined
DFA Australia Ltd.	35,683	$792,519	1.8%	Defined
Dimensional Ireland Ltd	60,274	$1,338,685	3.1%	Defined
Held directly	1,839,962	$40,865,556	94.5%	Sole

MPM BIOIMPACT LLC

$42,689,685

1.01%

1,922,093

SAMLYN CAPITAL, LLC

$38,972,063

0.92%

1,754,708

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SAMLYN ONSHORE FUND, LP joint	282,960	$6,284,541	16.1%	Defined
Samlyn Offshore Master Fund, Ltd.	669,773	$14,875,658	38.2%	Defined
Samlyn Net Neutral Master Fund, Ltd.	623,420	$13,846,158	35.5%	Defined
Samlyn Long Alpha Master Fund, Ltd.	178,555	$3,965,706	10.2%	Defined

Sofinnova Investments, Inc.

$36,832,708

0.87%

1,658,384

Palo Alto Investors LP

$36,591,441

0.86%

1,647,521

Vivo Capital, LLC

$34,591,319

0.82%

1,557,466

Aberdeen Group plc

$34,190,829

0.81%

1,539,434

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
abrdn Inc.	1,539,434	$34,190,829	100.0%	Defined

Deep Track Capital, LP

$33,315,000

0.79%

1,500,000

MARSHALL WACE, LLP

$33,068,845

0.78%

1,488,917

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marshall Wace North America L.P. joint	1,488,917	$33,068,845	100.0%	Other

Polar Capital Holdings Plc

$28,580,072

0.67%

1,286,811

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Polar Capital LLP	1,286,811	$28,580,072	100.0%	Defined

NORTHERN TRUST CORP

$28,386,176

0.67%

1,278,081

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	1,191,832	$26,470,587	93.3%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	65,094	$1,445,737	5.1%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	21,155	$469,852	1.7%	Other

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$26,700,928

0.63%

1,202,203

CITADEL ADVISORS LLC

$25,407,307

0.60%

1,143,958

Saturn V Capital Management LP

$23,486,919

0.55%

1,057,493

VANGUARD FIDUCIARY TRUST CO

$21,964,557

0.52%

988,949

NOVO HOLDINGS A/S

$21,330,350

0.50%

960,394

Rock Springs Capital Management LP

$20,925,240

0.49%

942,154

BRAIDWELL LP

$20,468,691

0.48%

921,598

FRONTIER CAPITAL MANAGEMENT CO LLC

$19,414,604

0.46%

874,138

BAKER BROS. ADVISORS LP

$18,066,702

0.43%

813,449

Fiera Capital Corp

$17,545,655

0.41%

789,989

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fiera Capital Inc.	773,391	$17,177,014	97.9%	Other
Held directly	16,598	$368,641	2.1%	Sole

MORGAN STANLEY

$16,573,100

0.39%

746,200

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	3,291	$73,093	0.4%	Defined
MORGAN STANLEY & CO. LLC	47,541	$1,055,885	6.4%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	500,547	$11,117,148	67.1%	Defined
MORGAN STANLEY SMITH BARNEY LLC	69,072	$1,534,089	9.3%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	125,749	$2,792,885	16.9%	Defined

Artisan Partners Limited Partnership

$14,653,847

0.35%

659,786

RAYMOND JAMES FINANCIAL INC

$12,800,733

0.30%

576,350

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	291,263	$6,468,951	50.5%	Defined
Raymond James Financial Services Advisors, Inc.	285,087	$6,331,782	49.5%	Defined

TD Asset Management Inc

$12,173,567

0.29%

548,112

ALGERT GLOBAL LLC

$11,244,323

0.27%

506,273

WASATCH ADVISORS LP

$9,853,866

0.23%

443,668

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 1	443,668	$9,853,866	100.0%	Sole

UBS Group AG

$9,845,334

0.23%

443,284

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	12,073	$268,141	2.7%	Defined
UBS AG	422,919	$9,393,030	95.4%	Defined
UBS Switzerland AG	7,970	$177,013	1.8%	Defined
UBS Europe SE	146	$3,242	0.0%	Defined
UBS Securities LLC	176	$3,908	0.0%	Defined

MILLENNIUM MANAGEMENT LLC

$9,591,388

0.23%

431,850

Clearbridge Investments, LLC

$9,220,059

0.22%

415,131

Bank of New York Mellon Corp

$9,101,676

0.21%

409,801

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Investments Corporation joint	145,472	$3,230,932	35.5%	Defined
The Bank of New York Mellon	255,650	$5,677,986	62.4%	Defined
BNY Mellon, NA joint	2,719	$60,388	0.7%	Defined
BNY Mellon Advisors, Inc. joint	5,959	$132,348	1.5%	Defined
Held directly	1	$22	0.0%	Defined

ArrowMark Colorado Holdings LLC

$8,822,278

0.21%

397,221

Nuveen, LLC

$8,590,538

0.20%

386,787

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	19,750	$438,647	5.1%	Defined
TIAA CREF INVESTMENT MANAGEMENT LLC joint	84,539	$1,877,611	21.9%	Defined
TEACHERS ADVISORS, LLC joint	282,498	$6,274,280	73.0%	Defined

RENAISSANCE TECHNOLOGIES LLC

$8,008,304

0.19%

360,572

Tetragon Partners GP Ltd

$7,921,351

0.19%

356,657

NEW YORK STATE COMMON RETIREMENT FUND

$7,905,716

0.19%

355,953

Granahan Investment Management, LLC

$7,519,528

0.18%

338,565

University of Wisconsin Foundation

$6,964,145

0.16%

313,559

MOORE CAPITAL MANAGEMENT, LP

$6,791,684

0.16%

305,794

Squarepoint Ops LLC

$6,664,643

0.16%

300,074

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Squarepoint Capital LLP	300,074	$6,664,643	100.0%	Defined

Fisher Asset Management, LLC

$6,629,796

0.16%

298,505

AMERIPRISE FINANCIAL INC

$6,625,620

0.16%

298,317

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	298,317	$6,625,620	100.0%	Defined

Ensign Peak Advisors, Inc

$6,434,903

0.15%

289,730

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$5,996,360

0.14%

269,985

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	2,509	$55,724	0.9%	Defined
UBS Asset Management Switzerland AG	174,078	$3,866,271	64.5%	Defined
UBS Asset Management (UK) Ltd.	59,063	$1,311,787	21.9%	Defined
Held directly	34,335	$762,578	12.7%	Defined

Alyeska Investment Group, L.P.

$5,447,624

0.13%

245,278

Catalio Capital Management, LP

$5,439,873

0.13%

244,929

DEUTSCHE BANK AG\

$5,383,524

0.13%

242,392

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Deutsche Bank Trust Co Americas	2,295	$50,971	0.9%	Defined
DWS Investment GmbH	100,000	$2,221,000	41.3%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC	895	$19,877	0.4%	Defined
DWS Investments UK Ltd	139,202	$3,091,676	57.4%	Defined

Swiss National Bank

$5,152,720

0.12%

232,000

WELLS FARGO & COMPANY/MN

$4,900,946

0.12%

220,664

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	120,358	$2,673,151	54.5%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	18,967	$421,257	8.6%	Defined
WELLS FARGO CLEARING SERVICES, LLC	81,306	$1,805,806	36.8%	Defined
WELLS FARGO INVESTMENT INSTITUTE, INC.	33	$732	0.0%	Defined

SPHERA FUNDS MANAGEMENT LTD.

$4,432,982

0.10%

199,594

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Sphera Global Healthcare Management LP joint	199,594	$4,432,982	100.0%	Defined

SummitTX Capital, L.P.

$4,323,021

0.10%

194,643

CITIGROUP INC

$4,173,591

0.10%

187,915

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	187,915	$4,173,591	100.0%	Defined

RHUMBLINE ADVISERS

$4,082,175

0.10%

183,799

VOYA INVESTMENT MANAGEMENT LLC

$3,874,689

0.09%

174,457

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	174,457	$3,874,689	100.0%	Defined

ALLIANCEBERNSTEIN L.P.

$3,821,896

0.09%

172,080

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	172,080	$3,821,896	100.0%	Defined

HighVista Strategies LLC

$3,819,253

0.09%

171,961

ALPS ADVISORS INC

$3,710,891

0.09%

167,082

Invesco Ltd.

$3,655,719

0.09%

164,598

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	18,998	$421,944	11.5%	Defined
Invesco Trust Co joint	54,595	$1,212,554	33.2%	Defined
Invesco Capital Management LLC	91,005	$2,021,221	55.3%	Defined

BNP PARIBAS ASSET MANAGEMENT Holding S.A.

$3,507,313

0.08%

157,916

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNP PARIBAS ASSET MANAGEMENT Europe	1,780	$39,533	1.1%	Defined
AXA INVESTMENT MANAGERS UK LIMITED	156,136	$3,467,780	98.9%	Defined

Y-Intercept (Hong Kong) Ltd

$3,469,268

0.08%

156,203

Darwin Wealth Management, LLC

$3,448,546

0.08%

155,270

AXXCESS WEALTH MANAGEMENT, LLC

$3,448,546

0.08%

155,270

Aegis Wealth Management LLC

$3,332,166

0.08%

150,030

ADAGE CAPITAL PARTNERS GP, L.L.C.

$3,331,500

0.08%

150,000

Woodline Partners LP

$3,317,907

0.08%

149,388

Schonfeld Strategic Advisors LLC

$2,992,841

0.07%

134,752

XTX Topco Ltd

$2,793,707

0.07%

125,786

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	125,786	$2,793,707	100.0%	Defined

Jain Global LLC

$2,583,933

0.06%

116,341

Rafferty Asset Management, LLC

$2,556,415

0.06%

115,102

Russell Investments Group, Ltd.

$2,344,330

0.06%

105,553

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC	59,708	$1,326,114	56.6%	Defined
Russell Investments Capital, LLC	36,414	$808,754	34.5%	Defined
Russell Investments Trust Company	9,431	$209,462	8.9%	Defined

Legal & General Group Plc

$2,175,246

0.05%

97,940

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	13,244	$294,149	13.5%	Defined
Legal & General Investment Management Ltd joint	84,696	$1,881,097	86.5%	Defined

SG Americas Securities, LLC

$2,168,762

0.05%

97,648

FIDEURAM - INTESA SANPAOLO PRIVATE BANKING S.P.A.

$1,933,802

0.05%

87,069

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Intesa Sanpaolo S.p.A.	87,069	$1,933,802	100.0%	Defined

AQR CAPITAL MANAGEMENT LLC

$1,866,837

0.04%

84,054

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RF&L WEALTH MANAGEMENT, LLC	6	$133	0.0%	Other
Prism Planning Partners LLC	12	$266	0.0%	Other
Arax Advisory Partners	170	$3,775	0.2%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	905	$20,100	1.1%	Other
Stokes Family Office, LLC	116	$2,576	0.1%	Other
Held directly	82,845	$1,839,987	98.6%	Defined

VANGUARD ASSET MANAGEMENT, Ltd

$1,804,295

0.04%

81,238

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
METLIFE INC	79,775	$1,771,802	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MANULIFE FINANCIAL CORP joint	78,622	$1,746,194	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Global X Management CO LLC joint	62,824	$1,395,321	82.5%	Defined
Global X Management (AUS) Ltd joint	13,371	$296,969	17.5%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fifth Third Bank, National Association	75,859	$1,684,828	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC CMA LTD.	4,149	$92,149	6.4%	Defined
RBC Capital Markets, LLC	60,593	$1,345,770	93.2%	Defined
RBC Trust Co (Delaware) Ltd	98	$2,176	0.2%	Defined
Held directly	175	$3,886	0.3%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	62,600	$1,390,346	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	62,000	$1,377,020	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	29,500	$655,195	49.0%	Defined
D. E. Shaw Investment Management, L.L.C. joint	30,661	$680,980	51.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wellington Management Company LLP	28,274	$627,965	53.9%	Defined
Wellington Managment Canada LLC	1,379	$30,627	2.6%	Defined
Wellington Trust Company, NA joint	19,618	$435,715	37.4%	Defined
Wellington Management Europe GmbH	3,156	$70,094	6.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	47,127	$1,046,690	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI INVESTMENTS MANAGEMENT CORP	577	$12,815	1.3%	Sole
SEI TRUST CO	43,477	$965,624	98.7%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Capital	40,235	$893,619	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Brevan Howard Investment Products Ltd	37,214	$826,522	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	32,089	$712,696	86.3%	Defined
BARCLAYS CAPITAL INC.	5,091	$113,071	13.7%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CREATIVE PLANNING TRUST Co LLC	650	$14,436	1.9%	Defined
Held directly	34,207	$759,737	98.1%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	31,900	$708,499	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Blackstone Alternative Asset Management L.P.	22,578	$501,457	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PGIM Quantitative Solutions LLC	22,480	$499,280	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	19,100	$424,211	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Allianz Global Invesstors Taiwan Ltd.	15,000	$333,150	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STIFEL NICOLAUS & CO INC \MO\	14,820	$329,152	99.1%	Defined
Held directly	132	$2,931	0.9%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	14,900	$330,929	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ROCKEFELLER & CO. LLC	599	$13,303	4.2%	Defined
ROCKEFELLER TRUST COMPANY, N.A.	2,383	$52,926	16.7%	Defined
ROCKEFELLER FINANCIAL LLC	11,296	$250,884	79.1%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
EATON VANCE MANAGEMENT	12,381	$274,982	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Universal-Investment-Gesellschaft mbH	12,111	$268,985	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	11,600	$257,636	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GUGGENHEIM FUNDS DISTRIBUTORS, LLC	9,738	$216,280	88.1%	Defined
SECURITY INVESTORS LLC	1,313	$29,161	11.9%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Latour Trading LLC (Latour)	5,187	$115,203	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nykredit Bank A/S	5,102	$113,315	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Edward D. Jones & Co., L.P.	4,928	$109,450	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
KBC Bank NV joint	3,405	$75,625	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
OSAIC WEALTH, INC.	1,208	$26,829	97.5%	Defined
Osaic Advisory Services, LLC	31	$688	2.5%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
US BANK NATIONAL ASSOCIATION	972	$21,587	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Optiver VOF	747	$16,590	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PNC Bank, National Association	534	$11,859	99.4%	Defined
PNC Wealth Management LLC	3	$66	0.6%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BMSS Wesson Wealth Solutions, LLC	1	$22	0.9%	Sole
Held directly	110	$2,441	99.1%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BESSEMER TRUST Co NATIONAL ASSOCIATION	97	$2,154	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Deka Vermoegensmanagement GmbH	60,000	$1,067	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	99,863	$1,810,516	0.7%	Defined
JANUS HENDERSON INVESTORS US LLC	13,547,513	$245,616,410	99.3%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AHL Partners LLP	45,618	$827,054	47.7%	Defined
Numeric Investors LLC	50,064	$907,660	52.3%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Arrowpoint Investment Partners (Hong Kong) Ltd	17,213	$312,071	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Inspire Investing, LLC	11,845	$214,749	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Capstone Investment Advisors (UK), LLP	11,217	$203,364	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jump Trading Holdings, LLC joint	10	$181	0.1%	Defined
Leap GP LLC joint	10,600	$192,178	99.9%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 1 joint	1,098	$19,906	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
National Bank Financial Inc.	200	$3,626	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Institutional Asset Management Trust Co	14	$253	48.3%	Sole
Held directly	15	$271	51.7%	Sole

Key facts CIK 1818794 CUSIP 26818M108 13F (30d) 237 filings 231 filers Visit website Investor relations