PCVX · Vaxcyte, Inc.

$63.30 +3.51 (+5.87%) At close · Aug 19

Market Cap

$8.99B

148.81M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Quarter:

Holders: 334

Total Value: $10,039,709,491

Total Shares: 172,711,390

Institutional Ownership Trend

25 quarters

Superinvestors Holding PCVX

All superinvestors →

Ray Dalio $2,919,114

Holder Concentration

Across 334 13F filers · Δ vs Mar 31, 2026

Top Holder

12.1%

+2.0 pts

Top 5

43.2%

+2.3 pts

Top 10

60.3%

+2.8 pts

HHI

489

Broadly held+48

Quarterly Activity

vs Mar 31, 2026

New

Increased

134

Decreased

102

Sold Out

Unchanged

Holder count: -5

Institutional value: +4.5%

Shares held: +4.5%

Top Buyers

vs Mar 31, 2026

Institution	Added	Now Holds
Mega FMR LLC	+4,360,046	20,852,907
Mega PRICE T ROWE ASSOCIATES INC /MD/	+2,012,570	2,988,999
Mega WELLINGTON MANAGEMENT GROUP LLP	+1,386,968	7,001,565
Mega SUSQUEHANNA INTERNATIONAL GROUP, LLP	+1,320,733	2,078,506
Mega BlackRock, Inc.	+1,096,313	12,068,129

Top Sellers

vs Mar 31, 2026

Institution	Sold	Now Holds
Mid Paradigm Biocapital Advisors LP	−2,665,168	2,127,852
Mega JENNISON ASSOCIATES LLC	−1,221,795	476,216
Large WESTFIELD CAPITAL MANAGEMENT CO LP	−1,173,710	2,472,428
Large Assenagon Asset Management S.A.	−1,063,380	101,433
Mega D. E. Shaw & Co., Inc.	−963,446	1,901,872

Top New Positions

vs Mar 31, 2026

Institution	Acquired	Value (USD)
Mega Jupiter Topco LLC	17,965,847	$1,044,354,683
Mega NORGES BANK	1,245,805	$72,418,644
Small First Turn Management, LLC	443,836	$25,800,186
Mid Jump Financial, LLC	135,000	$7,847,550
Mid Nicholas Investment Partners, LP	131,133	$7,622,761

Top Sold-Out Positions

vs Mar 31, 2026

Institution	Previously Held	Last Value (USD)
Mega JANUS HENDERSON GROUP PLC	16,575,875	$963,224,094
Large FARALLON CAPITAL MANAGEMENT LLC	1,261,000	$73,276,710
Mid BAUPOST GROUP LLC/MA	800,000	$46,488,000
Small Trails Edge Capital Partners, LP	662,470	$38,496,131
Small Jones Hill Capital LP	644,831	$37,471,129

Filter: New 50 Increased 134 Reduced 102 Sold out 55 Unchanged 48

Showing 100 of 350

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

FMR LLC

$1,212,179,481

12.07%

20,852,907

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC joint	13,178,987	$766,094,514	63.2%	Defined
FIDELITY MANAGEMENT TRUST CO	1,002,020	$58,247,422	4.8%	Defined
STRATEGIC ADVISERS LLC	1,702,376	$98,959,116	8.2%	Defined
Fidelity Institutional Asset Management Trust Co	873,656	$50,785,623	4.2%	Defined
FIAM LLC	4,095,868	$238,092,806	19.6%	Defined

Jupiter Topco LLC

$1,044,354,683

10.40%

17,965,847

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	179,089	$10,410,443	1.0%	Defined
JANUS HENDERSON INVESTORS US LLC	17,531,066	$1,019,080,866	97.6%	Defined
Held directly	255,692	$14,863,374	1.4%	Defined

T. Rowe Price Investment Management, Inc.

$709,619,068

7.07%

12,207,450

BlackRock, Inc.

$701,520,331

6.99%

12,068,129

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	169,695	$9,864,370	1.4%	Sole
BlackRock Investment Management (Australia) Ltd	280	$16,276	0.0%	Sole
BlackRock Asset Management Canada Ltd	21,599	$1,255,549	0.2%	Sole
BlackRock Investment Management, LLC	377,186	$21,925,822	3.1%	Sole
BLACKROCK ADVISORS LLC	656,339	$38,152,986	5.4%	Sole
BlackRock Fund Advisors	6,083,522	$353,635,133	50.4%	Sole
BlackRock Institutional Trust Company, N.A.	4,149,705	$241,222,351	34.4%	Sole
BlackRock Japan Co. Ltd	7,208	$419,001	0.1%	Sole
BlackRock Fund Managers Ltd.	22,648	$1,316,528	0.2%	Sole
BlackRock Investment Management (UK) Ltd.	42,744	$2,484,708	0.4%	Sole
BlackRock (Netherlands) B.V.	1,435	$83,416	0.0%	Sole
BlackRock Asset Management Ireland Ltd	285,279	$16,583,268	2.4%	Sole
BlackRock (Luxembourg) S.A.	621	$36,098	0.0%	Sole
BlackRock Life Ltd	1,053	$61,210	0.0%	Sole
BlackRock Asset Management Schweiz AG	20,078	$1,167,134	0.2%	Sole
Aperio Group, LLC	228,647	$13,291,250	1.9%	Sole
SpiderRock Advisors, LLC	90	$5,231	0.0%	Sole

RA CAPITAL MANAGEMENT, L.P.

$667,018,439

6.64%

11,474,599

WELLINGTON MANAGEMENT GROUP LLP

$407,000,970

4.05%

7,001,565

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wellington Management Company LLP	5,925,755	$344,464,138	84.6%	Defined
Wellington Managment Canada LLC	64,718	$3,762,057	0.9%	Defined
Wellington Management Singapore Pte. Ltd.	84,905	$4,935,527	1.2%	Defined
Wellington Management International Ltd	229,717	$13,353,449	3.3%	Defined
Wellington Trust Company, NA joint	653,713	$38,000,336	9.3%	Defined
Wellington Management Australia Pty Ltd	12,892	$749,411	0.2%	Defined
Wellington Management Europe GmbH	29,865	$1,736,052	0.4%	Defined

VANGUARD PORTFOLIO MANAGEMENT LLC

$383,283,119

3.82%

6,593,551

VANGUARD CAPITAL MANAGEMENT LLC

$369,048,710

3.68%

6,348,679

STATE STREET CORP

$311,094,204

3.10%

5,351,698

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	5,351,698	$311,094,204	100.0%	Defined

Deep Track Capital, LP

$249,959,000

2.49%

4,300,000

GEODE CAPITAL MANAGEMENT, LLC

$201,092,770

2.00%

3,459,363

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	487,464	$28,336,282	14.1%	Defined
Held directly	2,971,899	$172,756,488	85.9%	Defined

PRICE T ROWE ASSOCIATES INC /MD/

$173,750,511

1.73%

2,988,999

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	187,816	$10,917,744	6.3%	Defined
Held directly	2,801,183	$162,832,767	93.7%	Sole

Kynam Capital Management, LP

$169,875,333

1.69%

2,922,335

WESTFIELD CAPITAL MANAGEMENT CO LP

$143,722,239

1.43%

2,472,428

Caligan Partners LP

$127,225,410

1.27%

2,188,636

Paradigm Biocapital Advisors LP

$123,692,036

1.23%

2,127,852

Bellevue Group AG

$114,628,581

1.14%

1,971,935

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BB Biotech AG joint	1,967,377	$114,363,625	99.8%	Defined
Bellevue Asset Management AG	4,558	$264,956	0.2%	Defined

D. E. Shaw & Co., Inc.

$110,555,819

1.10%

1,901,872

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	1,895,547	$110,188,147	99.7%	Defined
D. E. Shaw Investment Management, L.L.C. joint	6,325	$367,672	0.3%	Defined

MAVERICK CAPITAL LTD

$93,777,757

0.93%

1,613,242

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$85,149,521

0.85%

1,464,812

Pivotal bioVenture Partners Investment Advisor LLC

$82,862,571

0.83%

1,425,470

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Pivotal bioVenture Partners Fund I U.G.P. Ltd	1,425,470	$82,862,571	100.0%	Defined

EVENTIDE ASSET MANAGEMENT, LLC

$80,232,130

0.80%

1,380,219

GOLDMAN SACHS GROUP INC

$78,519,677

0.78%

1,350,760

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	868,642	$50,494,159	64.3%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	474,968	$27,609,889	35.2%	Defined
Goldman Sachs Trust Company, N.A.	7,150	$415,629	0.5%	Defined

NORTHERN TRUST CORP

$74,416,395

0.74%

1,280,172

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	1,175,227	$68,315,944	91.8%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	84,494	$4,911,635	6.6%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	20,451	$1,188,816	1.6%	Other

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$72,929,898

0.73%

1,254,600

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	1,254,600	$72,929,898	100.0%	Other

NORGES BANK

$72,418,644

0.72%

1,245,805

JPMORGAN CHASE & CO

$71,345,621

0.71%

1,227,346

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
J.P. Morgan Trust Co of Delaware	1,465	$85,160	0.1%	Other
JPMorgan Chase Bank, N.A.	84,328	$4,901,986	6.9%	Defined
J.P. Morgan Investment Management Inc.	926,900	$53,880,697	75.5%	Defined
J.P. Morgan Securities LLC	170	$9,882	0.0%	Defined
JPMorgan Asset Management (UK) Ltd	214,478	$12,467,606	17.5%	Defined
55I, LLC	5	$290	0.0%	Other

DIMENSIONAL FUND ADVISORS LP

$69,451,047

0.69%

1,194,754

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	16,676	$969,375	1.4%	Defined
DFA Australia Ltd.	5,004	$290,882	0.4%	Defined
Dimensional Fund Advisors Pte. Ltd.	874	$50,805	0.1%	Defined
Dimensional Ireland Ltd	40,922	$2,378,795	3.4%	Defined
Held directly	1,131,278	$65,761,190	94.7%	Sole

Polar Capital Holdings Plc

$68,938,866

0.69%

1,185,943

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Polar Capital LLP	1,185,943	$68,938,866	100.0%	Defined

BANK OF AMERICA CORP /DE/

$62,710,526

0.62%

1,078,798

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	396,419	$23,043,836	36.7%	Defined
MERRILL LYNCH, PIERCE, FENNER & SMITH INC.	110,971	$6,450,744	10.3%	Defined
BOFA SECURITIES, INC.	126,070	$7,328,449	11.7%	Defined
MANAGED ACCOUNT ADVISORS LLC joint	445,338	$25,887,497	41.3%	Defined

Caption Management, LLC

$60,873,736

0.61%

1,047,200

VANGUARD FIDUCIARY TRUST CO

$54,098,219

0.54%

930,642

ADAGE CAPITAL PARTNERS GP, L.L.C.

$51,154,400

0.51%

880,000

Saturn V Capital Management LP

$49,579,890

0.49%

852,914

Capital Research Global Investors

$48,640,916

0.48%

836,761

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CAPITAL GROUP COMPANIES INC joint	836,761	$48,640,916	100.0%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$47,242,251

0.47%

812,700

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	812,700	$47,242,251	100.0%	Other

NOVO HOLDINGS A/S

$46,504,000

0.46%

800,000

MORGAN STANLEY

$46,371,342

0.46%

797,718

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	59	$3,429	0.0%	Defined
MORGAN STANLEY & CO. LLC	157,321	$9,145,069	19.7%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	42,312	$2,459,596	5.3%	Defined
MORGAN STANLEY INVESTMENT MANAGEMENT INC	46,884	$2,725,366	5.9%	Defined
MORGAN STANLEY SMITH BARNEY LLC	1,275	$74,115	0.2%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	491,599	$28,576,649	61.6%	Defined
EATON VANCE MANAGEMENT	10,000	$581,300	1.3%	Defined
Calvert Research & Management	48,268	$2,805,818	6.1%	Defined

AMERICAN CENTURY COMPANIES INC

$45,468,355

0.45%

782,184

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AMERICAN CENTURY INVESTMENT MANAGEMENT INC	782,184	$45,468,355	100.0%	Defined

UBS Group AG

$45,138,581

0.45%

776,511

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	1,797	$104,459	0.2%	Defined
UBS AG	648,852	$37,717,766	83.6%	Defined
UBS Switzerland AG	123,886	$7,201,492	16.0%	Defined
UBS Securities LLC	1,976	$114,864	0.3%	Defined

Clearbridge Investments, LLC

$43,713,701

0.44%

751,999

Pictet Asset Management Holding SA

$40,384,015

0.40%

694,719

Caption Management, LLC

$37,767,061

0.38%

649,700

Eversept Partners, LP

$36,619,109

0.36%

629,952

BOOTHBAY FUND MANAGEMENT, LLC

$35,726,698

0.36%

614,600

JANE STREET GROUP, LLC

$35,162,837

0.35%

604,900

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	604,900	$35,162,837	100.0%	Defined

FEDERATED HERMES, INC.

$34,945,488

0.35%

601,161

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FEDERATED GLOBAL INVESTMENT MANAGEMENT CORP	505,865	$29,405,932	84.1%	Defined
FEDERATED MDTA LLC	95,296	$5,539,556	15.9%	Defined

BOOTHBAY FUND MANAGEMENT, LLC

$31,050,895

0.31%

534,163

ALGERT GLOBAL LLC

$29,956,888

0.30%

515,343

Bank of New York Mellon Corp

$29,189,102

0.29%

502,135

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNY Mellon Trust of Delaware joint	585	$34,005	0.1%	Defined
Mellon Investments Corporation joint	220,900	$12,840,916	44.0%	Defined
The Bank of New York Mellon	238,659	$13,873,247	47.5%	Defined
BNY Mellon, NA joint	38,624	$2,245,211	7.7%	Defined
BNY Mellon Advisors, Inc. joint	3,366	$195,665	0.7%	Other
Held directly	1	$58	0.0%	Defined

BROWN ADVISORY INC

$28,310,297

0.28%

487,017

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Brown Advisory LLC	463,735	$26,956,915	95.2%	Defined
NEW YORK LIFE INVESTMENT MANAGEMENT LLC	23,282	$1,353,382	4.8%	Defined

JENNISON ASSOCIATES LLC

$27,682,436

0.28%

476,216

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Prudential Financial Inc	476,216	$27,682,436	100.0%	Defined

FRANKLIN RESOURCES INC

$27,670,111

0.28%

476,004

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FIDUCIARY TRUST CO INTERNATIONAL	81	$4,708	0.0%	Defined
FIDUCIARY TRUST INTERNATIONAL LLC	1,983	$115,271	0.4%	Defined
FRANKLIN ADVISERS INC	445,031	$25,869,652	93.5%	Defined
PUTNAM INVESTMENT MANAGEMENT LLC	28,909	$1,680,480	6.1%	Defined

BNP PARIBAS ARBITRAGE, SA

$27,058,178

0.27%

465,477

VICTORY CAPITAL MANAGEMENT INC

$26,316,787

0.26%

452,723

Elmind Capital, LP

$26,294,524

0.26%

452,340

First Turn Management, LLC

$25,800,186

0.26%

443,836

AMERIPRISE FINANCIAL INC

$25,606,553

0.26%

440,505

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	417,842	$24,289,155	94.9%	Defined
Ameriprise Trust Co	12,485	$725,753	2.8%	Defined
Ameriprise Financial Services, LLC	22	$1,278	0.0%	Defined
Ameriprise Bank, FSB	23	$1,336	0.0%	Defined
Columbia Threadneedle Management Ltd	10,133	$589,031	2.3%	Defined

Ikarian Capital, LLC

$22,594,084

0.23%

388,682

Ensign Peak Advisors, Inc

$22,187,407

0.22%

381,686

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$21,598,196

0.22%

371,550

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	1,840	$106,959	0.5%	Defined
UBS Asset Management Switzerland AG	325,526	$18,922,825	87.6%	Sole
UBS Asset Management (UK) Ltd.	22,810	$1,325,943	6.1%	Sole
Held directly	21,374	$1,242,469	5.8%	Other

RAYMOND JAMES FINANCIAL INC

$21,473,569

0.21%

369,406

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	113,928	$6,622,634	30.8%	Defined
Raymond James Financial Services Advisors, Inc.	20,584	$1,196,547	5.6%	Defined
EAGLE ASSET MANAGEMENT INC joint	234,894	$13,654,388	63.6%	Defined

Qube Research & Technologies Ltd

$21,113,106

0.21%

363,205

Aberdeen Group plc

$21,087,006

0.21%

362,756

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
abrdn Inc.	362,756	$21,087,006	100.0%	Defined

Alyeska Investment Group, L.P.

$21,058,871

0.21%

362,272

First Light Asset Management, LLC

$20,063,976

0.20%

345,157

Boxer Capital Management, LLC

$18,892,250

0.19%

325,000

Nuveen, LLC

$18,878,413

0.19%

324,762

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	38,348	$2,229,168	11.8%	Defined
TEACHERS ADVISORS, LLC joint	286,414	$16,649,245	88.2%	Defined

Spruce Street Capital LP

$17,790,512

0.18%

306,047

Woodline Partners LP

$15,970,636

0.16%

274,740

TD Asset Management Inc

$15,964,823

0.16%

274,640

TWO SIGMA INVESTMENTS, LP

$15,920,411

0.16%

273,876

Artisan Partners Limited Partnership

$15,729,396

0.16%

270,590

DEUTSCHE BANK AG\

$15,017,187

0.15%

258,338

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DWS Investment GmbH	85,132	$4,948,723	33.0%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC	37,474	$2,178,363	14.5%	Defined
DWS Investments UK Ltd	135,732	$7,890,101	52.5%	Defined

JANE STREET GROUP, LLC

$14,881,280

0.15%

256,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	256,000	$14,881,280	100.0%	Defined

Swiss National Bank

$14,270,915

0.14%

245,500

WCM INVESTMENT MANAGEMENT, LLC

$14,154,771

0.14%

243,502

University of Wisconsin Foundation

$13,936,609

0.14%

239,749

Nantahala Capital Management, LLC

$12,913,346

0.13%

222,146

California Public Employees Retirement System

$12,350,997

0.12%

212,472

Candriam S.C.A.

$12,094,179

0.12%

208,054

CITIGROUP INC

$11,781,380

0.12%

202,673

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citibank, N.A. joint	39	$2,267	0.0%	Defined
Citigroup Financial Products Inc. joint	202,634	$11,779,113	100.0%	Defined

Rosalind Advisors, Inc.

$11,626,000

0.12%

200,000

C WorldWide Group Holding A/S

$11,335,350

0.11%

195,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 2	195,000	$11,335,350	100.0%	Defined

SEGALL BRYANT & HAMILL, LLC

$10,885,946

0.11%

187,269

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NEW YORK LIFE INVESTMENT MANAGEMENT LLC	187,269	$10,885,946	100.0%	Sole

CITADEL ADVISORS LLC

$10,556,408

0.11%

181,600

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	181,600	$10,556,408	100.0%	Defined

CITADEL ADVISORS LLC

$10,259,945

0.10%

176,500

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	176,500	$10,259,945	100.0%	Defined

Invesco Ltd.

$10,082,007

0.10%

173,439

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	19,772	$1,149,345	11.4%	Defined
Invesco Trust Co joint	52,341	$3,042,582	30.2%	Defined
Invesco Capital Management LLC	101,326	$5,890,080	58.4%	Defined

RHUMBLINE ADVISERS

$9,795,893

0.10%

168,517

Russell Investments Group, Ltd.

$9,552,734

0.10%

164,334

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC	71,918	$4,180,592	43.8%	Defined
Russell Investments Capital, LLC	92,416	$5,372,142	56.2%	Defined

ALLIANCEBERNSTEIN L.P.

$9,295,510

0.09%

159,909

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	159,909	$9,295,510	100.0%	Defined

READYSTATE ASSET MANAGEMENT LP

$9,073,511

0.09%

156,090

NEW YORK STATE COMMON RETIREMENT FUND

$8,728,742

0.09%

150,159

HighVista Strategies LLC

$8,704,328

0.09%

149,739

ADAR1 Capital Management, LLC

$8,488,956

0.08%

146,034

SILVERARC CAPITAL MANAGEMENT, LLC

$8,428,850

0.08%

145,000

MILLENNIUM MANAGEMENT LLC

$7,942,011

0.08%

136,625

Jump Financial, LLC

$7,847,550

0.08%

135,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Leap GP LLC joint	135,000	$7,847,550	100.0%	Defined

WELLS FARGO & COMPANY/MN

$7,702,978

0.08%

132,513

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	121,815	$7,081,105	91.9%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	1,536	$89,287	1.2%	Defined
WELLS FARGO CLEARING SERVICES, LLC	9,122	$530,261	6.9%	Defined
WELLS FARGO INVESTMENT INSTITUTE, INC.	40	$2,325	0.0%	Defined

Nicholas Investment Partners, LP

$7,622,761

0.08%

131,133

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CREATIVE PLANNING TRUST Co LLC	3,527	$205,024	2.9%	Defined
Held directly	117,244	$6,815,393	97.1%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MANULIFE FINANCIAL CORP joint	108,504	$6,307,337	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	12,794	$743,715	13.1%	Defined
Legal & General Investment Management Ltd joint	84,657	$4,921,110	86.9%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AHL Partners LLP	81,632	$4,745,268	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
METLIFE INC	76,628	$4,454,385	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Global X Management CO LLC joint	60,681	$3,527,386	89.6%	Defined
Global X Management (AUS) Ltd joint	7,060	$410,397	10.4%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Universal-Investment-Gesellschaft mbH joint	66,572	$3,869,830	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	65,110	$3,784,843	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Chen Bihua	50,000	$2,906,500	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PGIM Quantitative Solutions LLC	35,970	$2,090,936	86.5%	Defined
Prudential Trust Co	5,600	$325,528	13.5%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	33,231	$1,931,718	85.1%	Defined
BARCLAYS CAPITAL INC.	5,661	$329,073	14.5%	Defined
BARCLAYS CAPITAL SECURITIES LTD	161	$9,358	0.4%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GUGGENHEIM FUNDS DISTRIBUTORS, LLC	5,813	$337,909	14.9%	Defined
SECURITY INVESTORS LLC	33,095	$1,923,812	85.1%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Capital	38,867	$2,259,338	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RF&L WEALTH MANAGEMENT, LLC	9	$523	0.0%	Other
Compound Planning, Inc.	1	$58	0.0%	Other
Arax Advisory Partners	104	$6,045	0.3%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	177	$10,289	0.5%	Other
Stokes Family Office, LLC	19	$1,104	0.1%	Other
Held directly	34,253	$1,991,126	99.1%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI INVESTMENTS MANAGEMENT CORP	2,178	$126,607	6.5%	Sole
SEI TRUST CO	31,338	$1,821,677	93.5%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLINGTON SMALL CAP EQUITY	30,576	$1,777,382	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	25,100	$1,459,063	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ADVISOR PARTNERS II, LLC	20,839	$1,211,371	87.3%	Defined
Willow Street Trust Co of Wyoming, LLC	460	$26,739	1.9%	Other
Held directly	2,577	$149,801	10.8%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NOMURA HOLDINGS INC joint	20,000	$1,162,600	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Deka Vermoegensmanagement GmbH	20,000	$1,162,600	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	18,191	$1,057,442	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Janus Henderson Group Ltd.	18,084	$1,051,222	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DNB FUND	18,041	$1,048,723	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Metzler Asset Management GmbH	17,235	$1,001,870	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Brevan Howard US Investment Management, LP	16,935	$984,431	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CIBC Private Wealth Advisors, Inc.	7,700	$447,601	47.1%	Defined
CIBC National Trust Co	1,356	$78,824	8.3%	Defined
CIBC WORLD MARKETS CORP	7,276	$422,953	44.6%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ALLSTATE INVESTMENTS LLC	14,370	$835,328	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Robert W. Baird & Co. Inc	13,813	$802,949	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Landscape High Leverage Fund, LP	2,277	$132,362	18.6%	Defined
LANDSCAPE OFFSHORE HL FUND LTD.	6,330	$367,962	51.8%	Defined
Held directly	3,604	$209,500	29.5%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marex Financial	12,127	$704,942	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
EATON VANCE MANAGEMENT	11,960	$695,234	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PNC Bank, National Association	11,618	$675,353	98.5%	Defined
PNC Wealth Management LLC	182	$10,579	1.5%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Thrivent Asset Management, LLC	4,558	$264,956	38.8%	Defined
Held directly	7,184	$417,605	61.2%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	11,206	$651,404	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC CMA LTD.	4,075	$236,879	38.0%	Defined
RBC Capital Markets, LLC	6,598	$383,541	61.5%	Defined
RBC Dominion Securities Inc.	26	$1,511	0.2%	Defined
RBC Rochdale, LLC	35	$2,034	0.3%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Desjardins Global Asset Management Inc.	10,062	$584,904	96.9%	Defined
Desjardins Securities inc.	320	$18,601	3.1%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ROCKEFELLER & CO. LLC	192	$11,160	1.9%	Defined
ROCKEFELLER TRUST COMPANY, N.A.	786	$45,690	7.8%	Defined
Rockefeller Trust Co (Delaware)	18	$1,046	0.2%	Defined
ROCKEFELLER FINANCIAL LLC	9,137	$531,133	90.2%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Amundi Asset Management	10,003	$581,474	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Daiwa Asset Management Co. Ltd	9,114	$529,796	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Blue Trust, Inc.	6,643	$386,157	79.5%	Defined
Held directly	1,718	$99,867	20.5%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
OSAIC WEALTH, INC.	2,170	$126,142	27.0%	Defined
CW Advisors, LLC	5,818	$338,200	72.5%	Sole
Osaic Advisory Services, LLC	40	$2,325	0.5%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Pacific Investment Management Company LLC	691	$40,167	8.9%	Defined
Allianz Global Invesstors Taiwan Ltd.	7,100	$412,723	91.1%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Allspring Funds Management, LLC reporting on behalf of Morgan Stanley Smith Barney, LLC	904	$52,549	12.3%	Defined
Allspring Global Investments, LLC reporting on behalf of UBS ASSET MANAGEMENT AMERICAS INC	6,460	$375,519	87.7%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Amova Asset Management Co., Ltd.	7,304	$424,581	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	418	$24,298	6.5%	Defined
Jane Street Global Trading, LLC	5,983	$347,791	93.5%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Edward D. Jones & Co., L.P.	6,068	$352,732	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Strive Asset Management, LLC	5,424	$315,297	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Tower Research Capital Investments LLC	25	$1,453	0.5%	Other
Latour Trading LLC (Latour)	4,963	$288,499	99.5%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nykredit Bank A/S	4,823	$280,360	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Capstone Investment Advisors (UK), LLP	4,666	$271,234	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manufacturers & Traders Trust Company joint	352	$20,460	8.2%	Defined
Wilmington Trust Investment Advisors, Inc. joint	60	$3,487	1.4%	Defined
Wilmington Trust National Association joint	3,191	$185,491	74.6%	Defined
Wilmington Trust Company	672	$39,063	15.7%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
HSBC Bank USA, N.A.	14	$813	0.3%	Defined
HSBC Global Asset Management (UK) Ltd	3,912	$227,404	94.2%	Defined
HSBC BANK PLC	51	$2,964	1.2%	Defined
HSBC SECURITIES USA INC	174	$10,114	4.2%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
KBC Bank NV joint	3,898	$226,590	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
First Sentier Investors RQI Pty Ltd	3,146	$182,876	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNP PARIBAS ASSET MANAGEMENT Europe	1,961	$113,992	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
US BANK NATIONAL ASSOCIATION	1,502	$87,311	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Intesa Sanpaolo S.p.A.	1,383	$80,393	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fifth Third Bank, National Association	1,379	$80,161	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BMSS Wesson Wealth Solutions, LLC	428	$24,879	58.2%	Sole
Held directly	307	$17,844	41.8%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Optiver VOF	731	$42,493	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Frost Investment Services, LLC	466	$27,088	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MITSUBISHI UFJ FINANCIAL GROUP INC	372	$21,624	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 1	361	$20,984	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BESSEMER TRUST Co NATIONAL ASSOCIATION	39	$2	0.0%	Defined
Bessemer Trust CO of Delaware, N.A.	123	$7,149	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Rakuten Group, Inc. joint	101	$5,871	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JOHNSON BANK	33	$1,918	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
O'SHAUGHNESSY ASSET MANAGEMENT, LLC	5	$290	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	170,121	$9,885,731	1.0%	Defined
JANUS HENDERSON INVESTORS US LLC joint	16,405,754	$953,338,363	99.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dynasty Wealth Management, LLC joint	9,370	$544,490	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Blackstone Alternative Asset Management L.P.	4,544	$264,051	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Virtu Americas LLC	4,429	$257,369	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Acadian Inc	1,072	$62,293	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Institutional Asset Management Trust Co	10	$581	20.0%	Sole
Held directly	40	$2,324	80.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
TORONTO DOMINION BANK	3	$174	100.0%	Defined

Key facts CIK 1649094 CUSIP 92243G108 13F (30d) 319 filings 315 filers Visit website Investor relations