OCUL · Ocular Therapeutix, Inc

$10.04 -0.06 (-0.59%) At close · Aug 14

Market Cap

$2.20B

219.62M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Quarter:

Holders: 259

Total Value: $2,176,338,470

Total Shares: 221,623,078

Institutional Ownership Trend

26 quarters

Holder Concentration

Across 259 13F filers · Δ vs Mar 31, 2026

Top Holder

15.9%

+0.5 pts

Top 5

46.8%

+3.0 pts

Top 10

64.9%

+4.4 pts

HHI

601

Broadly held+60

Quarterly Activity

vs Mar 31, 2026

New

Increased

104

Decreased

Sold Out

Unchanged

Holder count: -15

Institutional value: +21.7%

Shares held: +4.9%

Top Buyers

vs Mar 31, 2026

Institution	Added	Now Holds
Mega BlackRock, Inc.	+18,471,392	34,604,323
Mega DIMENSIONAL FUND ADVISORS LP	+2,133,611	2,783,367
Mega PRICE T ROWE ASSOCIATES INC /MD/	+1,726,384	7,623,533
Mid Logos Global Management LP	+1,450,000	5,300,000
Mega Nuveen, LLC	+1,286,194	1,697,274

Top Sellers

vs Mar 31, 2026

Institution	Sold	Now Holds
Mega ALLIANCEBERNSTEIN L.P.	−3,902,667	227,020
Large Point72 Asset Management, L.P.	−2,242,792	3,292,193
Large Caption Management, LLC	−1,585,459	39,800
Mega JANE STREET GROUP, LLC	−1,150,089	84,700
Mid Octagon Capital Advisors LP	−1,091,600	2,075,000

Top New Positions

vs Mar 31, 2026

Institution	Acquired	Value (USD)
Large ADAGE CAPITAL PARTNERS GP, L.L.C.	1,550,000	$15,221,000
Large Trexquant Investment LP	418,781	$4,112,429
Mid ORBIMED ADVISORS LLC	220,000	$2,160,400
Mid CSM Advisors, LLC	190,194	$1,867,705
Small Exome Asset Management LLC	181,588	$1,783,194

Top Sold-Out Positions

vs Mar 31, 2026

Institution	Previously Held	Last Value (USD)
Mid Commodore Capital LP	2,575,000	$21,810,250
Small DELTEC ASSET MANAGEMENT LLC	2,340,620	$19,825,051
Large Artisan Partners Limited Partnership	902,931	$7,647,825
Large READYSTATE ASSET MANAGEMENT LP	883,621	$7,484,269
Large Aberdeen Group plc	558,380	$4,729,478

Filter: New 34 Increased 104 Reduced 78 Sold out 49 Unchanged 43

Showing 100 of 271

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

FMR LLC

$322,528,500

14.82%

32,844,043

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	18,966,716	$186,253,151	57.7%	Defined
FIDELITY MANAGEMENT TRUST CO	13,134,074	$128,976,606	40.0%	Defined
STRATEGIC ADVISERS LLC	423	$4,153	0.0%	Defined
Fidelity Institutional Asset Management Trust Co	5,880	$57,741	0.0%	Defined
FIAM LLC	736,950	$7,236,849	2.2%	Defined

Deep Track Capital, LP

$193,319,289

8.88%

19,686,282

BlackRock, Inc.

$190,277,763

8.74%

19,376,555

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	633,086	$6,216,904	3.3%	Sole
BlackRock Investment Management (Australia) Ltd	508	$4,988	0.0%	Sole
BlackRock Asset Management Canada Ltd	1,091	$10,713	0.0%	Sole
BlackRock Investment Management, LLC	744,795	$7,313,886	3.8%	Sole
BLACKROCK ADVISORS LLC	1,983,588	$19,478,834	10.2%	Sole
BlackRock Fund Advisors	8,145,702	$79,990,793	42.0%	Sole
BlackRock Institutional Trust Company, N.A.	6,652,500	$65,327,550	34.3%	Sole
BlackRock Japan Co. Ltd	17,100	$167,922	0.1%	Sole
BlackRock Fund Managers Ltd.	32,438	$318,541	0.2%	Sole
BlackRock Investment Management (UK) Ltd.	270,323	$2,654,571	1.4%	Sole
BlackRock (Netherlands) B.V.	22,317	$219,152	0.1%	Sole
BlackRock Asset Management Ireland Ltd	352,624	$3,462,767	1.8%	Sole
BlackRock (Luxembourg) S.A.	353,707	$3,473,402	1.8%	Sole
BlackRock Asset Management Schweiz AG	24,182	$237,467	0.1%	Sole
Aperio Group, LLC	142,594	$1,400,273	0.7%	Sole

BlackRock, Inc.

$149,536,681

6.87%

15,227,768

Avoro Capital Advisors LLC

$123,142,800

5.66%

12,540,000

Summer Road LLC

$119,885,466

5.51%

12,208,296

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Data LLC	12,208,296	$119,885,466	100.0%	Defined

Venrock Adviser, LLC

$101,606,813

4.67%

10,346,926

VANGUARD CAPITAL MANAGEMENT LLC

$86,736,082

3.99%

8,832,595

PRICE T ROWE ASSOCIATES INC /MD/

$74,863,093

3.44%

7,623,533

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	281,293	$2,762,297	3.7%	Defined
Held directly	7,342,240	$72,100,796	96.3%	Sole

Logos Global Management LP

$52,046,000

2.39%

5,300,000

STATE STREET CORP

$51,390,829

2.36%

5,233,282

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	5,233,282	$51,390,829	100.0%	Defined

GEODE CAPITAL MANAGEMENT, LLC

$49,307,123

2.27%

5,021,092

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	688,311	$6,759,214	13.7%	Defined
Held directly	4,332,781	$42,547,909	86.3%	Defined

Point72 Asset Management, L.P.

$32,329,335

1.49%

3,292,193

DIMENSIONAL FUND ADVISORS LP

$27,332,662

1.26%

2,783,367

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	17,219	$169,090	0.6%	Defined
DFA Australia Ltd.	48,053	$471,880	1.7%	Defined
Dimensional Ireland Ltd	107,740	$1,058,006	3.9%	Defined
Held directly	2,610,355	$25,633,686	93.8%	Sole

Balyasny Asset Management L.P.

$22,298,342

1.02%

2,270,707

Octagon Capital Advisors LP

$20,376,500

0.94%

2,075,000

VANGUARD PORTFOLIO MANAGEMENT LLC

$19,376,136

0.89%

1,973,130

NORTHERN TRUST CORP

$17,670,361

0.81%

1,799,426

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	1,677,593	$16,473,962	93.2%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	89,973	$883,534	5.0%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	31,860	$312,865	1.8%	Other

GOLDMAN SACHS GROUP INC

$17,300,423

0.79%

1,761,754

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	1,417,347	$13,918,347	80.5%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	331,898	$3,259,238	18.8%	Defined
Goldman Sachs Trust Company, N.A.	12,509	$122,838	0.7%	Defined

Nuveen, LLC

$16,667,229

0.77%

1,697,274

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	29,303	$287,755	1.7%	Defined
TIAA CREF INVESTMENT MANAGEMENT LLC joint	357,840	$3,513,988	21.1%	Defined
TEACHERS ADVISORS, LLC joint	1,310,131	$12,865,486	77.2%	Defined

Woodline Partners LP

$16,375,920

0.75%

1,667,609

TCG Crossover Management, LLC

$16,180,924

0.74%

1,647,752

UBS Group AG

$15,230,829

0.70%

1,551,001

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	1,148,521	$11,278,476	74.1%	Defined
UBS AG	383,670	$3,767,639	24.7%	Defined
UBS Switzerland AG	18,810	$184,714	1.2%	Defined

ADAGE CAPITAL PARTNERS GP, L.L.C.

$15,221,000

0.70%

1,550,000

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$15,058,361

0.69%

1,533,438

Defilade Capital Management, L.P.

$14,634,608

0.67%

1,490,286

Squarepoint Ops LLC

$13,904,265

0.64%

1,415,913

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Squarepoint Capital LLP	1,415,913	$13,904,265	100.0%	Defined

Invesco Ltd.

$13,201,917

0.61%

1,344,391

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	30,614	$300,628	2.3%	Defined
Invesco Trust Co joint	81,754	$802,824	6.1%	Defined
Invesco Capital Management LLC	1,232,023	$12,098,465	91.6%	Defined

Boxer Capital Management, LLC

$12,766,000

0.59%

1,300,000

VANGUARD FIDUCIARY TRUST CO

$12,659,944

0.58%

1,289,200

TWO SIGMA INVESTMENTS, LP

$11,965,797

0.55%

1,218,513

Knoll Capital Management, LLC

$11,296,928

0.52%

1,150,400

HRT FINANCIAL LP

$10,577,023

0.49%

1,077,090

Balyasny Asset Management L.P.

$9,576,464

0.44%

975,200

RAYMOND JAMES FINANCIAL INC

$9,459,762

0.43%

963,316

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	377,826	$3,710,251	39.2%	Defined
Raymond James Financial Services Advisors, Inc.	585,490	$5,749,511	60.8%	Defined

AQR CAPITAL MANAGEMENT LLC

$9,332,316

0.43%

950,338

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Petros Family Wealth, LLC	65	$638	0.0%	Other
RF&L WEALTH MANAGEMENT, LLC	28	$274	0.0%	Other
Compound Planning, Inc.	11	$108	0.0%	Other
Arax Advisory Partners	2,604	$25,571	0.3%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	1,591	$15,623	0.2%	Other
Stokes Family Office, LLC	291	$2,857	0.0%	Other
Held directly	945,748	$9,287,245	99.5%	Defined

MORGAN STANLEY

$8,954,177

0.41%

911,831

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	7,926	$77,833	0.9%	Defined
MORGAN STANLEY & CO. LLC	65,511	$643,318	7.2%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	599,978	$5,891,783	65.8%	Defined
MORGAN STANLEY INVESTMENT MANAGEMENT INC	927	$9,103	0.1%	Defined
MORGAN STANLEY SMITH BARNEY LLC	94,206	$925,102	10.3%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	143,283	$1,407,038	15.7%	Defined

Candriam S.C.A.

$8,914,978

0.41%

907,839

Empire Life Investments Inc.

$8,560,015

0.39%

871,692

Cetera Investment Advisers

$7,715,888

0.35%

785,732

PEREGRINE CAPITAL MANAGEMENT LLC

$7,399,320

0.34%

753,495

Aptus Capital Advisors, LLC

$6,271,945

0.29%

638,691

Second Line Capital, LLC

$6,271,444

0.29%

638,640

Opaleye Management Inc.

$5,892,000

0.27%

600,000

ALGERT GLOBAL LLC

$5,816,474

0.27%

592,309

Bank of New York Mellon Corp

$5,432,775

0.25%

553,236

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Investments Corporation	190,367	$1,869,403	34.4%	Defined
The Bank of New York Mellon	362,868	$3,563,363	65.6%	Defined
Held directly	1	$9	0.0%	Defined

Man Group plc

$5,337,975

0.25%

543,582

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Numeric Investors LLC	543,582	$5,337,975	100.0%	Defined

Sio Capital Management, LLC

$5,297,821

0.24%

539,493

ACUTA CAPITAL PARTNERS, LLC

$5,228,963

0.24%

532,481

TUDOR INVESTMENT CORP ET AL

$5,132,550

0.24%

522,663

BANK OF AMERICA CORP /DE/

$5,098,845

0.23%

519,231

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	207,386	$2,036,530	39.9%	Defined
MERRILL LYNCH, PIERCE, FENNER & SMITH INC.	41,331	$405,870	8.0%	Defined
BOFA SECURITIES, INC.	131,823	$1,294,501	25.4%	Defined
MANAGED ACCOUNT ADVISORS LLC joint	138,691	$1,361,944	26.7%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$4,754,058

0.22%

484,120

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	484,120	$4,754,058	100.0%	Other

JPMORGAN CHASE & CO

$4,736,116

0.22%

482,293

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	111,845	$1,098,317	23.2%	Defined
J.P. Morgan Investment Management Inc.	361,448	$3,549,419	74.9%	Defined
J.P. Morgan Securities LLC	9,000	$88,380	1.9%	Defined

ESSEX INVESTMENT MANAGEMENT CO LLC

$4,734,752

0.22%

482,154

BEACON INVESTMENT ADVISORS LLC

$4,272,986

0.20%

435,131

BNP PARIBAS ASSET MANAGEMENT Holding S.A.

$4,153,967

0.19%

423,011

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNP PARIBAS ASSET MANAGEMENT Europe	2,292	$22,507	0.5%	Defined
AXA INVESTMENT MANAGERS UK LIMITED	420,719	$4,131,460	99.5%	Defined

Trexquant Investment LP

$4,112,429

0.19%

418,781

HighVista Strategies LLC

$4,011,803

0.18%

408,534

E. Ohman J:or Asset Management AB

$3,780,700

0.17%

385,000

Rock Springs Capital Management LP

$3,768,091

0.17%

383,716

RENAISSANCE TECHNOLOGIES LLC

$3,719,364

0.17%

378,754

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$3,627,508

0.17%

369,400

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	369,400	$3,627,508	100.0%	Other

Propel Bio Management, LLC

$3,515,226

0.16%

357,966

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Simplify Asset Management Inc.	357,966	$3,515,226	100.0%	Defined

Jump Financial, LLC

$3,446,201

0.16%

350,937

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jump Trading Holdings, LLC joint	5,137	$50,445	1.5%	Defined
Leap GP LLC joint	345,800	$3,395,756	98.5%	Defined

CITADEL ADVISORS LLC

$3,440,928

0.16%

350,400

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	350,400	$3,440,928	100.0%	Defined

JACOBS LEVY EQUITY MANAGEMENT, INC

$3,296,240

0.15%

335,666

CITADEL ADVISORS LLC

$3,048,952

0.14%

310,484

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	284,314	$2,791,963	91.6%	Defined
Held directly	26,170	$256,989	8.4%	Defined

BNP PARIBAS ARBITRAGE, SA

$3,033,427

0.14%

308,903

Swiss National Bank

$3,002,956

0.14%

305,800

KVP Capital Advisors, LP

$2,946,000

0.14%

300,000

Verition Fund Management LLC

$2,938,134

0.14%

299,199

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$2,859,584

0.13%

291,200

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	291,200	$2,859,584	100.0%	Other

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$2,819,887

0.13%

287,158

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	7,615	$74,779	2.7%	Defined
UBS Asset Management Switzerland AG	213,233	$2,093,948	74.3%	Sole
UBS Asset Management (UK) Ltd.	32,828	$322,369	11.4%	Defined
Held directly	33,482	$328,791	11.7%	Defined

RHUMBLINE ADVISERS

$2,657,704

0.12%

270,642

BARCLAYS PLC

$2,461,775

0.11%

250,690

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	247,393	$2,429,399	98.7%	Defined
BARCLAYS CAPITAL INC.	3,297	$32,376	1.3%	Defined

NOMURA ASSET MANAGEMENT INTERNATIONAL INC.

$2,435,831

0.11%

248,048

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NOMURA HOLDINGS INC joint	248,048	$2,435,831	100.0%	Defined

WELLS FARGO & COMPANY/MN

$2,269,136

0.10%

231,073

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	185,168	$1,818,349	80.1%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	500	$4,910	0.2%	Defined
WELLS FARGO CLEARING SERVICES, LLC	45,300	$444,846	19.6%	Defined
WELLS FARGO INVESTMENT INSTITUTE, INC.	105	$1,031	0.0%	Defined

Advisory Services Network, LLC

$2,263,058

0.10%

230,454

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Institutional Asset Management Trust Co	21	$206	0.0%	Other
Held directly	230,433	$2,262,852	100.0%	Sole

TD Asset Management Inc

$2,256,636

0.10%

229,800

MANUFACTURERS LIFE INSURANCE COMPANY, THE

$2,251,293

0.10%

229,256

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MANULIFE FINANCIAL CORP joint	229,256	$2,251,293	100.0%	Defined

ALLIANCEBERNSTEIN L.P.

$2,229,336

0.10%

227,020

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	227,020	$2,229,336	100.0%	Defined

Rhenman & Partners Asset Management AB

$2,209,500

0.10%

225,000

DEUTSCHE BANK AG\

$2,201,103

0.10%

224,145

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DWS INVESTMENT MANAGEMENT AMERICAS, INC	3,105	$30,491	1.4%	Defined
DWS Investments UK Ltd	211,704	$2,078,933	94.4%	Defined
DWS International GmbH	9,336	$91,679	4.2%	Defined

ORBIMED ADVISORS LLC

$2,160,400

0.10%

220,000

SEI INVESTMENTS CO

$2,048,942

0.09%

208,650

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI INVESTMENTS MANAGEMENT CORP	123	$1,207	0.1%	Sole
SEI TRUST CO	208,527	$2,047,735	99.9%	Sole

Walleye Capital LLC

$1,981,499

0.09%

201,782

Russell Investments Group, Ltd.

$1,888,599

0.09%

192,322

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC joint	88,880	$872,800	46.2%	Defined
Russell Investments Capital, LLC joint	57,084	$560,564	29.7%	Defined
Russell Investments France SAS	9,870	$96,923	5.1%	Defined
Russell Investments Trust Company joint	36,488	$358,312	19.0%	Other

CSM Advisors, LLC

$1,867,705

0.09%

190,194

SCHRODER INVESTMENT MANAGEMENT GROUP

$1,849,773

0.08%

188,368

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Schroder Investment Management Limited	187,557	$1,841,809	99.6%	Sole
Schroder Investment Management North America Inc.	811	$7,964	0.4%	Sole

BOOTHBAY FUND MANAGEMENT, LLC

$1,830,015

0.08%

186,356

Exome Asset Management LLC

$1,783,194

0.08%

181,588

TUDOR INVESTMENT CORP ET AL

$1,683,148

0.08%

171,400

Universal- Beteiligungs- und Servicegesellschaft mbH

$1,660,208

0.08%

169,064

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Universal-Investment-Gesellschaft mbH	169,064	$1,660,208	100.0%	Defined

Steward Partners Investment Advisory, LLC

$1,621,694

0.07%

165,142

Wealthspire Advisors, LLC

$1,600,267

0.07%

162,960

CITADEL ADVISORS LLC

$1,519,154

0.07%

154,700

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	154,700	$1,519,154	100.0%	Defined

WESBANCO BANK INC

$1,463,180

0.07%

149,000

CITIGROUP INC

$1,381,850

0.06%

140,718

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	140,718	$1,381,850	100.0%	Defined

SIMPLEX TRADING, LLC

$1,338,033

0.06%

136,256

MARSHALL WACE, LLP

$1,237,643

0.06%

126,033

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marshall Wace North America L.P. joint	126,033	$1,237,643	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. Shaw Investment Management, L.L.C. joint	124,457	$1,222,167	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Truist Advisory Services, Inc.	121,000	$1,188,220	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
METLIFE INC	120,033	$1,178,724	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	19,994	$196,341	17.0%	Defined
Legal & General Investment Management Ltd joint	97,921	$961,583	83.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK VONTOBEL AG	111,368	$1,093,633	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	101,471	$996,445	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Global X Management CO LLC joint	94,818	$931,112	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	91,375	$897,302	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Brown Advisory LLC	90,000	$883,800	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Prudential Financial Inc	75,829	$744,640	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wellington Management Company LLP	36,149	$354,983	50.6%	Defined
Wellington Managment Canada LLC	2,018	$19,816	2.8%	Defined
Wellington Trust Company, NA joint	28,715	$281,981	40.2%	Defined
Wellington Management Europe GmbH	4,619	$45,358	6.5%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	68,500	$672,670	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC CMA LTD.	6,263	$61,502	10.0%	Defined
RBC Capital Markets, LLC	56,163	$551,520	89.9%	Defined
Held directly	35	$343	0.1%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CREATIVE PLANNING TRUST Co LLC	92	$903	0.2%	Defined
Held directly	61,193	$600,915	99.8%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Capital	60,732	$596,388	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DNB FUND	52,722	$517,730	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
OSAIC WEALTH, INC.	29,730	$291,948	57.5%	Defined
OSAIC INSTITUTIONS, INC.	200	$1,964	0.4%	Defined
Osaic Advisory Services, LLC	21,810	$214,174	42.2%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	49,000	$481,180	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	28,979	$284,573	80.5%	Defined
Ameriprise Financial Services, LLC	7,001	$68,749	19.5%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	35,700	$350,574	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 1	34,966	$343,366	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PGIM Quantitative Solutions LLC	33,700	$330,934	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PEAK6 Investments LLC joint	26,656	$261,761	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Brevan Howard US Investment Management, LP	24,659	$242,151	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CIBC Private Wealth Advisors, Inc.	8,040	$78,952	41.4%	Defined
CIBC WORLD MARKETS CORP	11,368	$111,633	58.6%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
EATON VANCE MANAGEMENT	18,688	$183,516	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Yasumitsu Shigeta	17,000	$166,940	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ALLSTATE INVESTMENTS LLC	16,047	$157,581	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Allianz Global Invesstors Taiwan Ltd.	15,200	$149,264	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF NOVA SCOTIA	13,337	$130,969	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
KBC Bank NV joint	6,462	$63,456	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Latour Trading LLC (Latour)	6,022	$59,136	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Edward D. Jones & Co., L.P.	3,744	$36,766	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ROCKEFELLER & CO. LLC	558	$5,479	17.5%	Defined
ROCKEFELLER TRUST COMPANY, N.A.	2,223	$21,829	69.9%	Defined
ROCKEFELLER FINANCIAL LLC	400	$3,928	12.6%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nykredit Bank A/S	1,931	$18,962	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PNC Bank, National Association	957	$9,397	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BMSS Wesson Wealth Solutions, LLC	41	$402	8.3%	Sole
Held directly	451	$4,426	91.7%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
US BANK NATIONAL ASSOCIATION	447	$4,389	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
TORONTO DOMINION BANK	81	$795	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
abrdn Inc.	558,380	$4,729,478	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Thakore Nainoor	125,000	$1,058,750	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	68,500	$580,195	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NOMURA SECURITIES INTERNATIONAL INC	50,000	$423,500	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Virtu Americas LLC	44,560	$377,423	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JEFFERIES LLC	22,100	$187,187	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wabon Partners, L.P.	20,000	$169,400	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Landscape High Leverage Fund, LP	3,030	$25,664	19.6%	Defined
LANDSCAPE OFFSHORE HL FUND LTD.	8,316	$70,436	53.7%	Defined
Held directly	4,141	$35,074	26.7%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Hudson Bay Capital UK LLP	12,540	$106,213	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manufacturers & Traders Trust Company joint	759	$6,427	7.1%	Defined
Wilmington Trust Investment Advisors, Inc. joint	27	$228	0.3%	Defined
Wilmington Trust National Association joint	9,327	$78,999	87.4%	Defined
Wilmington Trust Company	558	$4,726	5.2%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Amundi Asset Management	10,589	$89,688	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
National Bank Financial Inc.	125	$1,058	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Optiver VOF	10	$84	100.0%	Defined

Key facts CIK 1393434 CUSIP 67576A100 13F (30d) 252 filings 247 filers Visit website Investor relations