VCYT · Veracyte, Inc. · Holdings

$50.28 +2.08 (+4.32%) Q2 2026 earnings call transcript

S&P 600 member Russell 2000 member Rebalance outlook locked

Market Cap

$4.12B

80.39M

Volume · Oct 27 1.81M Avg daily vol (3M) 1.38M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Use institutional holdings through the 13F API or MCP → See every holder of VCYT: 13F, 13D and 13G, insiders and funds →

The Sep 30, 2026 filing window is still open — showing the latest known positions. Funds that have not filed yet carry their prior-quarter positions; a fund that filed without this stock counts as sold out. Each fund's Change is measured against its own previous reported quarter, shown in the Reported column. Pick a completed quarter in the Quarter selector below to put every fund on one quarter and see the stock-wide activity breakdown.

Quarter:

Holders: 379

Total Value: $5,324,192,771

Total Shares: 91,396,756

Showing 1–100 of 385 holdings by value · page 1 of 4

Institutional Ownership Trend

27 quarters

Superinvestors Holding VCYT

All superinvestors →

Ray Dalio $3,423,312 Joel Greenblatt $1,370,699

Holder Concentration

Across 372 13F filers

Top Holder

15.1%

of institutional value

Top 5

40.5%

of institutional value

Top 10

55.6%

of institutional value

HHI

481

Broadly held

Who Holds VCYT

Last 8 quarters through Sep 30, 2026 (filing window open)

Each kind of 13F filer's share of the common shares 13F filers report. Holder type comes from regulatory records, never the filer's name; style, turnover and horizon are measured from each filer's own 13F book. The quarter selector does not change this card.

Jun 30, 2026, the newest closed quarter: 370 filers, like-for-like change +0.9%; actively managed books -0.2%; index-like books +3.0%.

Holder type	Share	Filers	Change	New	Exited
Registered adviser	47.3%	196	-1.6%	49	29
Exempt adviser	0.1%	3	-54.0%	0	3
Adviser	1.6%	8	-6.9%	0	0
Bank	11.0%	21	-5.4%	0	0
Insurer	0.8%	3	+7.1%	1	0
Broker-dealer	0.0%	1	-13.7%	0	0
Benefit plan	0.2%	5	+2.0%	1	0
Non-US	2.1%	12	+72.6%	1	1
Holding company	30.2%	11	+2.8%	1	0
Unknown	6.6%	110	+14.1%	22	8

Style	Share	Filers	Change	New	Exited
Index-like	36.4%	40	+3.0%	2	0
Systematic-like	2.4%	14	-6.1%	6	11
Concentrated	1.3%	59	-3.7%	18	10
Diversified active	59.8%	257	+0.1%	49	20
Not measured	0.0%	0	—	0	0
Unknown	0.0%	0	—	0	0

Turnover	Share	Filers	Change	New	Exited
Low turnover	57.8%	200	+1.1%	29	11
Medium turnover	28.5%	104	+1.1%	21	11
High turnover	3.0%	33	-15.7%	16	18
Unknown	10.7%	33	+5.0%	9	1
Not measured	0.0%	0	—	0	0

Holding horizon	Share	Filers	Change	New	Exited
Short horizon	11.4%	43	+8.3%	17	8
Medium horizon	17.7%	58	+1.8%	16	11
Long horizon	65.7%	246	-1.5%	35	15
Unknown	5.2%	23	+15.3%	7	7
Not measured	0.0%	0	—	0	0

13F book size	Share	Filers	Change	New	Exited
Under $1B	1.5%	76	+5.4%	24	11
$1B to $10B	5.8%	98	-32.7%	22	16
$10B to $100B	23.8%	122	+6.4%	26	12
$100B or more	68.8%	74	+3.4%	3	2
Not measured	0.0%	0	—	0	0

Client base	Share	Filers	Change	New	Exited
Individuals and high net worth	3.2%	94	+6.4%	26	11
Pooled investment vehicles	5.2%	27	-15.9%	10	11
Registered funds and BDCs	26.0%	30	-1.5%	3	3
Pension and profit sharing plans	0.0%	2	+53.4%	0	0
Insurance companies	0.5%	4	+1.2%	0	0
Governments and sovereign funds	0.8%	8	-20.3%	4	2
Corporations	0.0%	1	new	1	0
Other investment advisers	0.1%	6	+17.3%	1	0
Other	0.1%	2	+12.2%	1	0
Diversified	11.4%	22	+5.3%	3	2
Unknown	52.7%	174	+3.3%	26	12

Private funds	Share	Filers	Change	New	Exited
Hedge funds	19.8%	46	-8.6%	15	18
Private equity funds	0.4%	9	+4.9%	1	0
Venture capital funds	0.0%	1	+96.2%	0	0
Real estate funds	0.0%	4	+29.5%	0	1
Securitized asset funds	0.2%	1	-0.8%	0	0
Other private funds	8.8%	23	+6.0%	4	0
Mixed private funds	0.4%	3	+4.6%	1	0
Private funds, type not stated	0.5%	4	-0.4%	0	0
No private funds	17.2%	108	+2.5%	28	13
Unknown	52.6%	171	+3.6%	26	9

Location	Share	Filers	Change	New	Exited
United States	92.5%	301	-0.2%	62	35
Outside the US	7.5%	69	+17.1%	13	6

The Sep 30, 2026 filing window is still open, so its bar carries the Jun 30, 2026 positions of filers that have not filed yet.

How the mix is measured

Shares are 13F common shares restated to today's split basis; options and principal amounts are left out. The change compares only filers with a 13F in both quarters and keeps each filer in its bucket of the later quarter on both sides; a filer that continues under a new CIK is compared through its successor. Holder type comes from each filer's current regulatory profile; client base and private funds come from the Form ADV roster current at each quarter end (rosters from December 2019, so an earlier quarter reads Unknown). Style, turnover, horizon and book size are measured from each filer's 13F book in the newest measured quarter, and Not measured means none was. Actively managed books are those measured systematic-like, concentrated or diversified active.

Excluded 3 holders with no observed 2026-03-31 13F and 1 holder with no observed 2026-06-30 13F. Compared across CIK changes: JANUS HENDERSON GROUP PLC continues as Jupiter Topco LLC.

Conviction and Crowding

370 13F filers, quarter ended Jun 30, 2026

Measured across every 13F filer's common-share book once the quarter's filing window closed. Crowding counts filers with high turnover or that advise private funds on Form ADV.

Conviction holders

0.541% of filers hold it in their top 10

Mar 31, 2026: 0

Holders above 5%

median weight 0.004%

Mar 31, 2026: 0

Crowding

—

holders exceed shares outstanding: 13F filers report more shares than the cover-page count, which predates an issuance, is misstated, or misses shares lent and reported twice

Private-fund advisers

—

of shares outstanding

Institutional ownership

—

holders exceed shares outstanding: 13F filers report more shares than the cover-page count, which predates an issuance, is misstated, or misses shares lent and reported twice

Top 10 holders

55.7%

of 13F shares, Herfindahl 0.048

Active holders

-0.2%

like-for-like share change of active filers

5% stakes on record

0 13D, 7 13G (7 institutional, 0 passive, 0 exempt)

How long holders have held it value-weighted 16.3 quarters

Under 1 year 19.1% 1 to 3 years 23.1% 3 years or more 57.8% Since coverage start 0.000%

Managers in the top fifth by current three-year FundScore alpha were net buyers of $0 (0 buying, 0 selling).

Estimated Holder Cost Basis

Quarter ended Jun 30, 2026, on that quarter's share basis

Estimated, never filed: each 13F buy is costed at its quarter's average traded price and each sale keeps the average cost, so this is what holders paid on average if they traded evenly through each quarter.

Estimated avg cost

$34.80

estimated per share

Mar 31, 2026: $32.44

Quarter-end price

$58.69

implied by 13F values

Estimated gain or loss

+68.6%

price against the estimated cost

Estimated at a loss

0.000%

of costed 13F value

Estimated cost covers 25.6% of the 13F shares; positions already held when a filer's 13F history starts, or bought in a quarter without a trusted average price, have no estimate.

Shared Holders

Companies VCYT's active holders also own, quarter ended Jun 30, 2026

#	Company	Shared holders	Similarity
1	WGS GeneDx Holdings Corp.	8	0.237
2	VCEL Vericel Corp	8	0.237
3	AXGN Axogen, Inc.	8	0.213
4	CDNA CareDx, Inc.	7	0.207
5	PSNL Personalis, Inc.	6	0.203
6	KNSA Kiniksa Pharmaceuticals International, plc	7	0.198
7	CERS Cerus Corp	4	0.187
8	ECPG Encore Capital Group Inc	5	0.181
9	GH Guardant Health, Inc.	12	0.176
10	LFST LifeStance Health Group, Inc.	4	0.168

Shareholder Targets

13F filers that own its peers but not VCYT, quarter ended Jun 30, 2026

Found through its holder-overlap peers: WGS, VCEL, AXGN, CDNA, PSNL, KNSA, CERS, ECPG, GH, LFST, AGYS, ONTO, RELY, LIVN, ALKS, CAMT, ANIP, NTRA, TWST, HNGE. 1,300 filers in all.

#	Filer	In peers	Of book	Largest peers
1	TPG GP A, LLC Unknown, Concentrated	$1,196,784,815 1 held	27.2%	LFST
2	TWO SIGMA INVESTMENTS, LP Registered adviser, Systematic-like	$1,188,569,179 18 held	0.930%	GH, ONTO, HNGE
3	NORGES BANK Non-US, Index-like	$1,173,359,294 11 held	0.120%	NTRA, ONTO, GH
4	FARALLON CAPITAL MANAGEMENT LLC Registered adviser, Diversified active	$870,307,948 3 held	5.14%	NTRA, TWST, GH
5	Duquesne Family Office LLC Unknown, Diversified active	$864,922,763 1 held	20.0%	NTRA
6	CITADEL ADVISORS LLC Registered adviser, Systematic-like	$841,887,758 18 held	0.490%	GH, ONTO, TWST
7	Deep Track Capital, LP Adviser, Diversified active	$820,353,599 4 held	12.5%	GH, ALKS, PSNL
8	BAILLIE GIFFORD & CO Adviser, Diversified active	$732,013,301 3 held	0.660%	GH, RELY, TWST
9	BAKER BROS. ADVISORS LP Registered adviser, Concentrated	$724,592,867 3 held	3.64%	ALKS, KNSA, CERS
10	Castle Hook Partners LP Registered adviser, Diversified active	$705,857,981 2 held	7.58%	NTRA, TWST

Fund Ownership of VCYT by Fund Type

Registered funds' Form N-PORT reports for the quarter ended Jun 30, 2026

Index funds and index ETFs hold 33.91% of shares outstanding.

Fund type	Of shares outstanding	Change	Shares	Funds
Index funds	4.83%	+0.57 pp	3,879,203	80
Index ETFs	29.08%	+0.76 pp	23,380,728	76
Active ETFs	5.28%	-0.84 pp	4,245,708	32
Active funds	23.94%	+4.22 pp	19,243,917	149
Unclassified	0.02%	-0.01 pp	19,455	7
All registered funds	63.15%	+4.70 pp	50,769,011	344

How fund types are measured

Each fund series counts once, with its latest Form N-PORT report dated inside the quarter; only long share positions count, restated to today's split basis. The fund type is the series' latest Form N-CEN answer on or before the quarter end (the index-fund box and the exchange-traded flag), never its name. Unclassified funds have no usable N-CEN; unit investment trusts are among them, so index ownership is a floor. Funds sit inside their advisers' 13F reports, so these shares overlap the 13F figures and are never added to them.

Holder type: Registered adviser 203 Exempt adviser 3 Adviser 8 Bank 21 Insurer 3 Broker-dealer 1 Benefit plan 5 Non-US 12 Holding company 11 Unknown 112

Showing 100 of 385

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

BlackRock, Inc.

$792,058,437

14.88%

13,486,437

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	119,517	$7,019,233	0.9%	Sole
BlackRock Investment Management (Australia) Ltd	278	$16,326	0.0%	Sole
BlackRock Asset Management Canada Ltd	21,684	$1,273,501	0.2%	Sole
BlackRock Investment Management, LLC	276,643	$16,247,243	2.1%	Sole
BLACKROCK ADVISORS LLC	383,973	$22,550,734	2.8%	Sole
BlackRock Fund Advisors	9,344,128	$548,780,637	69.3%	Sole
BlackRock Institutional Trust Company, N.A.	2,641,270	$155,121,787	19.6%	Sole
BlackRock Japan Co. Ltd	1,649	$96,845	0.0%	Sole
BlackRock Fund Managers Ltd.	14,926	$876,603	0.1%	Sole
BlackRock Investment Management (UK) Ltd.	42,169	$2,476,585	0.3%	Sole
BlackRock (Netherlands) B.V.	87,432	$5,134,881	0.6%	Sole
BlackRock Asset Management Ireland Ltd	423,089	$24,848,016	3.1%	Sole
BlackRock (Luxembourg) S.A.	9,139	$536,733	0.1%	Sole
BlackRock Life Ltd	367	$21,553	0.0%	Sole
BlackRock Asset Management Schweiz AG	13,922	$817,639	0.1%	Sole
Aperio Group, LLC	106,251	$6,240,121	0.8%	Sole

FMR LLC

$452,382,271

8.50%

7,702,746

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	6,633,700	$389,597,201	86.1%	Defined
STRATEGIC ADVISERS LLC	1,656	$97,256	0.0%	Defined
Fidelity Institutional Asset Management Trust Co	117,094	$6,876,930	1.5%	Defined
FIAM LLC	950,296	$55,810,884	12.3%	Defined

Artisan Partners Limited Partnership

$325,270,466

6.11%

5,538,404

VANGUARD PORTFOLIO MANAGEMENT LLC

$276,981,897

5.20%

4,716,191

STATE STREET CORP

$275,921,057

5.18%

4,698,128

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	4,698,128	$275,921,057	100.0%	Defined

VANGUARD CAPITAL MANAGEMENT LLC

$211,071,039

3.96%

3,593,922

DIMENSIONAL FUND ADVISORS LP

$166,001,753

3.12%

2,826,524

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	13,690	$804,013	0.5%	Defined
DFA Australia Ltd.	32,003	$1,879,536	1.1%	Defined
Dimensional Fund Advisors Pte. Ltd.	495	$29,071	0.0%	Defined
Dimensional Ireland Ltd	83,537	$4,906,128	3.0%	Defined
Held directly	2,696,799	$158,383,005	95.4%	Sole

FIRST TRUST ADVISORS LP

$147,289,084

2.77%

2,507,902

ARK Investment Management LLC

$134,841,202

2.53%

2,295,951

WELLINGTON MANAGEMENT GROUP LLP

$133,561,414

2.51%

2,274,160

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wellington Management Company LLP	1,951,842	$114,631,680	85.8%	Defined
Wellington Managment Canada LLC	841	$49,391	0.0%	Defined
Wellington Management International Ltd	180,307	$10,589,430	7.9%	Defined
Wellington Trust Company, NA joint	139,244	$8,177,800	6.1%	Defined
Wellington Management Europe GmbH	1,926	$113,113	0.1%	Defined

GEODE CAPITAL MANAGEMENT, LLC

$123,567,919

2.32%

2,104,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	296,538	$17,415,676	14.1%	Defined
Held directly	1,807,462	$106,152,243	85.9%	Defined

GOLDMAN SACHS GROUP INC

$96,488,161

1.81%

1,642,911

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	672,705	$39,507,964	40.9%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	940,207	$55,218,357	57.2%	Defined
GOLDMAN SACHS INTERNATIONAL	4,507	$264,696	0.3%	Defined
Goldman Sachs Trust Company, N.A. joint	25,492	$1,497,144	1.6%	Defined

CONGRESS ASSET MANAGEMENT CO

$89,971,275

1.69%

1,954,198

FULLER & THALER ASSET MANAGEMENT, INC.

$88,667,558

1.67%

1,509,749

WILLIAM BLAIR INVESTMENT MANAGEMENT, LLC

$84,077,633

1.58%

1,431,596

GW&K Investment Management, LLC

$83,193,570

1.56%

1,416,543

MORGAN STANLEY

$69,993,822

1.31%

1,191,790

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	578	$33,945	0.0%	Defined
MORGAN STANLEY & CO. LLC	30,018	$1,762,957	2.5%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	836,088	$49,103,448	70.2%	Defined
MORGAN STANLEY INVESTMENT MANAGEMENT INC	4,671	$274,327	0.4%	Defined
MORGAN STANLEY SMITH BARNEY LLC	44,946	$2,639,678	3.8%	Defined
Morgan Stanley Wealth Management Australia Pty Ltd	1,677	$98,490	0.1%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	273,812	$16,080,977	23.0%	Defined

BANK OF AMERICA CORP /DE/

$64,736,844

1.22%

1,102,279

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	232,723	$13,667,821	21.1%	Defined
BOFA SECURITIES, INC.	271,970	$15,972,798	24.7%	Defined
MANAGED ACCOUNT ADVISORS LLC joint	597,586	$35,096,225	54.2%	Defined

Driehaus Capital Management LLC

$61,421,360

1.15%

1,045,826

NORTHERN TRUST CORP

$58,504,121

1.10%

996,154

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	897,973	$52,737,952	90.1%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	58,964	$3,462,955	5.9%	Defined
MASON STREET ADVISORS, LLC joint	26,644	$1,564,802	2.7%	Other
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	12,573	$738,412	1.3%	Other

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$55,205,201

1.04%

939,983

OBERWEIS ASSET MANAGEMENT INC/

$51,549,317

0.97%

877,734

Legal & General Group Plc

$50,398,326

0.95%

858,136

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	122,340	$7,185,028	14.3%	Defined
Legal & General Investment Management Ltd joint	735,796	$43,213,298	85.7%	Defined

Sumitomo Mitsui Trust Group, Inc.

$45,160,374

0.85%

768,949

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Amova Asset Management Co., Ltd.	768,949	$45,160,374	100.0%	Defined

Amova Asset Management Americas, Inc.

$45,160,374

0.85%

768,949

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Sumitomo Mitsui Trust Group, Inc. joint	768,949	$45,160,374	100.0%	Sole

Point72 Asset Management, L.P.

$43,331,228

0.81%

737,804

Fisher Asset Management, LLC

$41,145,768

0.77%

700,592

BAMCO INC /NY/

$38,214,788

0.72%

650,686

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARON CAPITAL MANAGEMENT INC /NY/ joint	650,686	$38,214,788	100.0%	Defined

SEGALL BRYANT & HAMILL, LLC

$37,189,421

0.70%

633,227

VANGUARD FIDUCIARY TRUST CO

$33,054,594

0.62%

562,823

ARROWSTREET CAPITAL, LIMITED PARTNERSHIP

$31,219,223

0.59%

531,572

Alyeska Investment Group, L.P.

$30,424,665

0.57%

518,043

JPMORGAN CHASE & CO

$30,272,494

0.57%

515,452

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	50,819	$2,984,599	9.9%	Defined
J.P. Morgan Investment Management Inc.	261,437	$15,354,195	50.7%	Defined
J.P. Morgan Securities LLC	1,295	$76,055	0.3%	Defined
J.P. Morgan Securities plc	6,046	$355,081	1.2%	Defined
JPMorgan Asset Management (UK) Ltd	195,855	$11,502,564	38.0%	Defined

Artemis Investment Management LLP

$29,604,686

0.56%

643,021

ROYAL BANK OF CANADA

$29,569,142

0.56%

503,476

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC CMA LTD.	2,457	$144,299	0.5%	Defined
RBC Capital Markets, LLC	5,232	$307,275	1.0%	Defined
RBC Global Asset Management (U.S.) Inc.	405,866	$23,836,510	80.6%	Defined
RBC Dominion Securities Inc.	449	$26,369	0.1%	Defined
RBC Trust Co (Delaware) Ltd	120	$7,047	0.0%	Defined
CITY NATIONAL BANK	410	$24,079	0.1%	Other
RBC Rochdale, LLC	500	$29,365	0.1%	Defined
Alaska Permanent Fund Corp joint	86,810	$5,098,351	17.2%	Defined
Held directly	1,632	$95,847	0.3%	Sole

PRINCIPAL FINANCIAL GROUP INC

$28,855,164

0.54%

491,319

RAYMOND JAMES FINANCIAL INC

$27,737,764

0.52%

472,293

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	40,695	$2,390,017	8.6%	Defined
Raymond James Financial Services Advisors, Inc.	2,904	$170,551	0.6%	Defined
ClariVest Asset Management LLC joint	61,552	$3,614,948	13.0%	Defined
EAGLE ASSET MANAGEMENT INC joint	367,142	$21,562,248	77.7%	Defined

UBS Group AG

$26,429,320

0.50%

450,014

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	29,716	$1,745,220	6.6%	Defined
UBS AG	412,534	$24,228,121	91.7%	Defined
UBS Switzerland AG	7,764	$455,979	1.7%	Defined

ArrowMark Colorado Holdings LLC

$25,957,837

0.49%

441,986

BNP PARIBAS ARBITRAGE, SA

$25,878,376

0.49%

440,633

Nuveen, LLC

$25,789,516

0.48%

439,120

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	26,826	$1,575,490	6.1%	Defined
TIAA CREF INVESTMENT MANAGEMENT LLC joint	92,059	$5,406,625	21.0%	Defined
TEACHERS ADVISORS, LLC joint	320,235	$18,807,401	72.9%	Defined

Bank of New York Mellon Corp

$25,700,477

0.48%

437,604

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Investments Corporation joint	217,499	$12,773,715	49.7%	Defined
BNY Mellon Investment Adviser, Inc.	64,760	$3,803,354	14.8%	Defined
The Bank of New York Mellon	149,826	$8,799,280	34.2%	Defined
BNY Mellon, NA joint	2,278	$133,786	0.5%	Defined
BNY Mellon Advisors, Inc. joint	1,893	$111,175	0.4%	Other
Newton Investment Management North America, LLC	1,346	$79,050	0.3%	Defined
Held directly	2	$117	0.0%	Defined

Invesco Ltd.

$25,312,627

0.48%

431,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	32,097	$1,885,055	7.4%	Defined
Invesco Trust Co joint	32,282	$1,895,921	7.5%	Defined
Invesco Capital Management LLC	366,621	$21,531,651	85.1%	Defined

D. E. Shaw & Co., Inc.

$24,257,662

0.46%

413,037

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	296,083	$17,388,954	71.7%	Defined
D. E. Shaw Investment Management, L.L.C. joint	116,954	$6,868,708	28.3%	Defined

SEI INVESTMENTS CO

$24,108,488

0.45%

410,497

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI INVESTMENTS MANAGEMENT CORP	1,957	$114,934	0.5%	Sole
SEI TRUST CO	408,540	$23,993,554	99.5%	Sole

NEW YORK STATE COMMON RETIREMENT FUND

$23,741,543

0.45%

404,249

JUPITER ASSET MANAGEMENT LTD

$20,884,622

0.39%

355,604

Public Sector Pension Investment Board

$20,521,730

0.39%

349,425

KAMES CAPITAL plc

$19,151,735

0.36%

326,098

FRED ALGER MANAGEMENT, LLC

$18,870,594

0.35%

321,311

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Weatherbie Capital, LLC	314,582	$18,475,400	97.9%	Sole
Held directly	6,729	$395,194	2.1%	Sole

USS Investment Management Ltd

$18,786,376

0.35%

319,877

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
USS Investment Management Ltd	319,877	$18,786,376	100.0%	Sole

SILVERARC CAPITAL MANAGEMENT, LLC

$18,676,198

0.35%

318,001

Freestone Grove Partners LP

$17,613,773

0.33%

299,911

Casdin Capital, LLC

$16,227,568

0.30%

276,308

Jupiter Topco LLC

$15,845,705

0.30%

269,806

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	34,326	$2,015,965	12.7%	Defined
JANUS HENDERSON INVESTORS US LLC	235,480	$13,829,740	87.3%	Defined

Granite Investment Partners, LLC

$14,854,167

0.28%

252,923

NORDEA INVESTMENT MANAGEMENT AB

$13,450,755

0.25%

229,027

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nordea Funds Ltd	54,831	$3,220,224	23.9%	Defined
Nordea Investment Funds S.A.	174,196	$10,230,531	76.1%	Defined

PRUDENTIAL FINANCIAL INC

$12,896,520

0.24%

219,590

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PGIM Quantitative Solutions LLC	194,090	$11,398,905	88.4%	Defined
Prudential Trust Co	25,500	$1,497,615	11.6%	Defined

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$11,972,518

0.22%

203,857

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	4,168	$244,786	2.0%	Defined
UBS Asset Management Switzerland AG	136,267	$8,002,960	66.8%	Defined
UBS Asset Management (UK) Ltd.	46,477	$2,729,593	22.8%	Defined
Held directly	16,945	$995,179	8.3%	Defined

RHUMBLINE ADVISERS

$11,097,150

0.21%

188,952

BLAIR WILLIAM & CO/IL

$10,554,485

0.20%

179,712

Champlain Investment Partners, LLC

$10,449,828

0.20%

177,930

Palisades Investment Partners, LLC

$10,279,746

0.19%

175,034

Universal- Beteiligungs- und Servicegesellschaft mbH

$9,616,039

0.18%

163,733

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Universal-Investment-Gesellschaft mbH joint	163,733	$9,616,039	100.0%	Defined

Corient Private Wealth LLC

$9,515,493

0.18%

162,021

Swiss National Bank

$9,367,435

0.18%

159,500

AQR CAPITAL MANAGEMENT LLC

$8,788,825

0.17%

149,648

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Petros Family Wealth, LLC	34	$1,996	0.0%	Other
RF&L WEALTH MANAGEMENT, LLC	11	$646	0.0%	Other
Compound Planning, Inc.	25	$1,468	0.0%	Other
Arax Advisory Partners	259	$15,211	0.2%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	407	$23,903	0.3%	Other
Stokes Family Office, LLC	53	$3,112	0.0%	Other
Held directly	148,859	$8,742,489	99.5%	Defined

EXCHANGE TRADED CONCEPTS, LLC

$8,472,683

0.16%

144,265

Focus Partners Wealth

$8,011,359

0.15%

136,410

Candriam S.C.A.

$7,928,256

0.15%

134,995

Allianz Asset Management GmbH

$7,878,629

0.15%

134,150

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Allianz Global Investors GmbH	132,150	$7,761,169	98.5%	Defined
Allianz Global Invesstors Taiwan Ltd.	2,000	$117,460	1.5%	Defined

NEUBERGER BERMAN GROUP LLC

$7,788,654

0.15%

132,618

MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.

$7,755,354

0.15%

132,051

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Global X Management CO LLC joint	127,232	$7,472,335	96.4%	Defined
Global X Management (AUS) Ltd joint	4,819	$283,019	3.6%	Defined

AMERIPRISE FINANCIAL INC

$7,604,359

0.14%

129,480

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	129,413	$7,600,425	99.9%	Defined
Ameriprise Financial Services, LLC	67	$3,934	0.1%	Defined

ARDSLEY ADVISORY PARTNERS LP

$7,341,250

0.14%

125,000

Caption Management, LLC

$7,311,885

0.14%

124,500

FRANKLIN RESOURCES INC

$7,284,455

0.14%

124,033

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FIDUCIARY TRUST CO INTERNATIONAL	42	$2,466	0.0%	Defined
FIDUCIARY TRUST INTERNATIONAL LLC	958	$56,263	0.8%	Defined
FRANKLIN ADVISERS INC	23,630	$1,387,789	19.1%	Defined
Franklin Templeton International Services S.a r.l	215	$12,626	0.2%	Defined
PUTNAM INVESTMENT MANAGEMENT LLC	99,188	$5,825,311	80.0%	Defined

EFG International AG

$7,224,670

0.14%

123,015

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
EFG Asset Management (North America) Corp.	122,960	$7,221,440	100.0%	Defined
EFG Private Wealth Management (Americas) Corp.	55	$3,230	0.0%	Defined

Summit Creek Advisors LLC

$6,986,285

0.13%

118,956

PRICE T ROWE ASSOCIATES INC /MD/

$6,437,101

0.12%

109,605

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	109,605	$6,437,101	100.0%	Defined

ALLIANCEBERNSTEIN L.P.

$6,209,464

0.12%

105,729

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	105,729	$6,209,464	100.0%	Defined

DEERFIELD MANAGEMENT COMPANY, L.P.

$5,931,730

0.11%

101,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Deerfield UGP, LLC	101,000	$5,931,730	100.0%	Defined

DEUTSCHE BANK AG\

$5,913,170

0.11%

100,684

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Deutsche Bank Trust Co Americas	11,217	$658,774	11.1%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC	1,185	$69,595	1.2%	Defined
DWS Investments UK Ltd	85,474	$5,019,888	84.9%	Defined
DWS International GmbH	2,808	$164,913	2.8%	Defined

IQ EQ FUND MANAGEMENT (IRELAND) Ltd

$5,706,089

0.11%

97,158

BROWN CAPITAL MANAGEMENT LLC

$5,639,430

0.11%

96,023

RICE HALL JAMES & ASSOCIATES, LLC

$5,613,765

0.11%

95,586

ExodusPoint Capital Management, LP

$5,523,204

0.10%

94,044

WELLS FARGO & COMPANY/MN

$5,451,023

0.10%

92,815

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	75,249	$4,419,373	81.1%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	2,682	$157,513	2.9%	Defined
WELLS FARGO CLEARING SERVICES, LLC	14,825	$870,672	16.0%	Defined
WELLS FARGO INVESTMENT INSTITUTE, INC.	59	$3,465	0.1%	Defined

CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM

$5,363,282

0.10%

91,321

Russell Investments Group, Ltd.

$5,328,278

0.10%

90,725

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC	60,394	$3,546,939	66.6%	Defined
Russell Investments Capital, LLC	25,220	$1,481,170	27.8%	Defined
Russell Investments Trust Company	5,111	$300,169	5.6%	Defined

ENVESTNET ASSET MANAGEMENT INC

$4,945,829

0.09%

84,213

PINNACLE ASSOCIATES LTD

$4,753,998

0.09%

103,258

CITIGROUP INC

$4,660,518

0.09%

79,355

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	79,355	$4,660,518	100.0%	Defined

GLOBEFLEX CAPITAL L P

$4,379,848

0.08%

74,576

VANGUARD ASSET MANAGEMENT, Ltd

$4,292,928

0.08%

73,096

CASTLEARK MANAGEMENT LLC

$4,229,499

0.08%

72,016

MILLENNIUM MANAGEMENT LLC

$4,227,502

0.08%

71,982

CANADA LIFE ASSURANCE Co

$4,205,479

0.08%

71,607

LOS ANGELES CAPITAL MANAGEMENT LLC

$4,086,727

0.08%

69,585

Informed Momentum Co LLC

$4,029,406

0.08%

68,609

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Key facts CIK 1384101 CUSIP 92337F107 13F (30d) 57 filings 57 filers Visit website Investor relations