ARCT · Arcturus Therapeutics Holdings Inc. · Shareholders & Ownership

$9.84 -0.89 (-8.29%) At close · Oct 23

Russell 2000 member Rebalance outlook locked

Market Cap

$285.23M

28.42M

Volume · Oct 23 475K Avg daily vol (3M) 939.9K

Every disclosed holder of ARCT in one list: 13F institutions, Schedule 13D and 13G filers, insiders (Forms 3, 4 and 5) and registered funds, each from its own latest filing. Holders nest by CIK and SEC reporting links, never by name, and each is summed once.

How the holder list picks a filing →

Who Owns ARCT

13F quarter Sep 30, 2026, combined view: filers that have not filed yet carry their Jun 30, 2026 positions

Holder group	Holders	Shares	% Out	Summed
Institutional (13F)	136	24,929,486	87.71%	Yes
Insiders (Forms 3, 4, 5)	2	223,965	0.79%	Yes
13D/G-only blocks	0	0	0.00%	Yes
Remaining	—	3,269,618	11.50%	Rest
Registered funds (N-PORT, not summed)	142	11,370,423	40.00%	No

Shares outstanding: 28,423,069, the issuer's latest cover-page count restated onto today's split basis.

13D/G-only counts each filer's latest 13D or 13G that no 13F report of the listing contains, that names no 13F filer and that no counted insider files. Funds are never summed: their shares sit inside their advisers' 13F reports.

Holders of ARCT

One row per holder Every filing By ultimate parent

All 13F Funds Insiders 13D/G Non-US

#	Holder	Shares	% Out	Change	As of	Notes
1	ARK Investment Management LLC 13F CIK 1697748 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	2,901,635	10.21%	+132,117	Jun 30, 2026	carried, not yet filed for the newest quarter
1.1	ARK Genomic Revolution ETF Fund S000042975 · Registered fund · Jul 31, 2026 ARK ETF Trust, not summed	2,741,123	9.64%	-67,920	Jul 31, 2026	ARK ETF Trust, not summed
2	BlackRock, Inc. 13F CIK 2012383 · Holding company · Jun 30, 2026 carried, not yet filed for the newest quarter	2,614,012	9.20%	+731	Jun 30, 2026	carried, not yet filed for the newest quarter
2.1	iShares Russell 2000 ETF Fund S000004344 · Registered fund · Jun 30, 2026 iShares Trust, not summed	712,395	2.51%	+49,406	Jun 30, 2026	iShares Trust, not summed
2.2	iShares Russell 2000 Value ETF Fund S000004342 · Registered fund · Jun 30, 2026 iShares Trust, not summed	254,094	0.89%	+104,109	Jun 30, 2026	iShares Trust, not summed
2.3	iShares Biotechnology ETF Fund S000004350 · Registered fund · Jun 30, 2026 iShares Trust, not summed	183,754	0.65%	-3,213	Jun 30, 2026	iShares Trust, not summed
2.4	BlackRock Advantage Small Cap Core Fund Fund S000039982 · Registered fund · May 29, 2026 BlackRock Funds, not summed	135,078	0.48%	0	May 29, 2026	BlackRock Funds, not summed
2.5	iShares Micro-Cap ETF Fund S000004439 · Registered fund · Jun 30, 2026 iShares Trust, not summed	90,153	0.32%	+23,214	Jun 30, 2026	iShares Trust, not summed
3	Sumitomo Mitsui Trust Group, Inc. 13F CIK 1475365 · Bank · Jun 30, 2026 carried, not yet filed for the newest quarter	1,963,123	6.91%	-325,812	Jun 30, 2026	carried, not yet filed for the newest quarter
4	Federated Hermes Kaufmann Fund Fund S000009729 · Registered fund · Jul 31, 2026 Federated Hermes Equity Funds, not summed	1,962,580	6.90%	-109,920	Jul 31, 2026	Federated Hermes Equity Funds, not summed
5	Amova Asset Management Americas, Inc. 13F CIK 1480751 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	1,960,458	6.90%	-328,477	Jun 30, 2026	carried, not yet filed for the newest quarter
6	FEDERATED HERMES, INC. 13G CIK 1056288 · Investment adviser · Sep 30, 2026 Rule 13d-1(b), qualified institution, not summed	1,678,069	5.90%	-1,625,571	Sep 30, 2026	Rule 13d-1(b), qualified institution, not summed
6.1	FEDERATED HERMES, INC. 13F CIK 1056288 · Investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	3,303,640	11.62%	0	Jun 30, 2026	carried, not yet filed for the newest quarter
7	Federated Hermes Kaufmann Small Cap Fund Fund S000009730 · Registered fund · Jul 31, 2026 Federated Hermes Equity Funds, not summed	1,186,700	4.18%	0	Jul 31, 2026	Federated Hermes Equity Funds, not summed
8	VANGUARD CAPITAL MANAGEMENT LLC 13F CIK 2100119 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	1,114,956	3.92%	+7,936	Jun 30, 2026	carried, not yet filed for the newest quarter
9	TWO SIGMA INVESTMENTS, LP 13F CIK 1179392 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	1,091,137	3.84%	+9,353	Jun 30, 2026	carried, not yet filed for the newest quarter
10	VANGUARD PORTFOLIO MANAGEMENT LLC 13F CIK 2100121 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	951,575	3.35%	-26,347	Jun 30, 2026	carried, not yet filed for the newest quarter
11	VANGUARD TOTAL STOCK MARKET INDEX FUND Fund S000002848 · Registered fund · Jun 30, 2026 VANGUARD INDEX FUNDS, not summed	797,154	2.80%	0	Jun 30, 2026	VANGUARD INDEX FUNDS, not summed
12	GEODE CAPITAL MANAGEMENT, LLC 13F CIK 1214717 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	693,660	2.44%	+32,732	Jun 30, 2026	carried, not yet filed for the newest quarter
13	AQR CAPITAL MANAGEMENT LLC 13F CIK 1167557 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	675,014	2.37%	+513,736	Jun 30, 2026	carried, not yet filed for the newest quarter
13.1	AQR Small Cap Multi-Style Fund Fund S000040064 · Registered fund · Jun 30, 2026 AQR Funds, not summed	23,162	0.08%	—	Jun 30, 2026	AQR Funds, not summed
14	JACOBS LEVY EQUITY MANAGEMENT, INC 13F CIK 884414 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	647,511	2.28%	+448,783	Jun 30, 2026	carried, not yet filed for the newest quarter
15	STATE STREET CORP 13F CIK 93751 · Bank · Jun 30, 2026 carried, not yet filed for the newest quarter	569,598	2.00%	+25,735	Jun 30, 2026	carried, not yet filed for the newest quarter
15.1	STATE STREET SMALL/MID CAP EQUITY INDEX PORTFOLIO Fund S000050332 · Registered fund · Jun 30, 2026 STATE STREET INSTITUTIONAL INVESTMENT TRUST, not summed	9,935	0.04%	+1,900	Jun 30, 2026	STATE STREET INSTITUTIONAL INVESTMENT TRUST, not summed
15.2	State Street Total Return V.I.S. Fund Fund S000001637 · Registered fund · Jun 30, 2026 STATE STREET VARIABLE INSURANCE SERIES FUNDS INC, not summed	630	0.00%	0	Jun 30, 2026	STATE STREET VARIABLE INSURANCE SERIES FUNDS INC, not summed
15.3	State Street Balanced Index Fund Fund S000082802 · Registered fund · Jun 30, 2026 STATE STREET INSTITUTIONAL INVESTMENT TRUST, not summed	166	0.00%	0	Jun 30, 2026	STATE STREET INSTITUTIONAL INVESTMENT TRUST, not summed
16	UBS Group AG 13F CIK 1610520 · Bank · Jun 30, 2026 carried, not yet filed for the newest quarter	490,632	1.73%	+32,355	Jun 30, 2026	carried, not yet filed for the newest quarter
17	D. E. Shaw & Co., Inc. 13F CIK 1009207 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	392,358	1.38%	+140,324	Jun 30, 2026	carried, not yet filed for the newest quarter
18	Empire Life Investments Inc. 13F CIK 1640951 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	366,217	1.29%	-17,276	Jun 30, 2026	carried, not yet filed for the newest quarter
19	Hennion & Walsh Asset Management, Inc. 13F CIK 1861642 · Investment adviser · Sep 30, 2026	349,102	1.23%	+2,895	Sep 30, 2026
20	MILLENNIUM MANAGEMENT LLC 13F CIK 1273087 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	336,996	1.19%	-141,561	Jun 30, 2026	carried, not yet filed for the newest quarter
21	GOLDMAN SACHS GROUP INC 13F CIK 886982 · Holding company · Jun 30, 2026 carried, not yet filed for the newest quarter	309,863	1.09%	+111,182	Jun 30, 2026	carried, not yet filed for the newest quarter
21.1	Goldman Sachs ActiveBeta(R) U.S. Small Cap Equity ETF Fund S000048296 · Registered fund · May 31, 2026 Goldman Sachs ETF Trust, not summed	12,950	0.05%	-11,793	May 31, 2026	Goldman Sachs ETF Trust, not summed
21.2	Goldman Sachs Absolute Return Tracker Fund Fund S000020314 · Registered fund · Jun 30, 2026 GOLDMAN SACHS TRUST, not summed	3,896	0.01%	+1,911	Jun 30, 2026	GOLDMAN SACHS TRUST, not summed
22	VANGUARD EXTENDED MARKET INDEX FUND Fund S000002841 · Registered fund · Jun 30, 2026 VANGUARD INDEX FUNDS, not summed	293,104	1.03%	+7,431	Jun 30, 2026	VANGUARD INDEX FUNDS, not summed
23	VANGUARD EXPLORER FUND Fund S000002578 · Registered fund · Jul 31, 2026 VANGUARD EXPLORER FUND, not summed	285,663	1.01%	0	Jul 31, 2026	VANGUARD EXPLORER FUND, not summed
24	BNP PARIBAS ARBITRAGE, SA 13F CIK 1166588 · Non-US institution · Jun 30, 2026 carried, not yet filed for the newest quarter	261,789	0.92%	+66,929	Jun 30, 2026	carried, not yet filed for the newest quarter
25	VANGUARD STRATEGIC SMALL-CAP EQUITY FUND Fund S000012432 · Registered fund · Jun 30, 2026 VANGUARD HORIZON FUNDS, not summed	253,764	0.89%	0	Jun 30, 2026	VANGUARD HORIZON FUNDS, not summed
26	CHARLES SCHWAB INVESTMENT MANAGEMENT INC 13F CIK 884546 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	231,628	0.81%	+15,128	Jun 30, 2026	carried, not yet filed for the newest quarter
26.1	Schwab U.S. Small-Cap ETF Fund S000026636 · Registered fund · May 31, 2026 SCHWAB STRATEGIC TRUST, not summed	131,117	0.46%	-3,732	May 31, 2026	SCHWAB STRATEGIC TRUST, not summed
26.2	Schwab Small-Cap Index Fund Fund S000005912 · Registered fund · Jul 31, 2026 SCHWAB CAPITAL TRUST, not summed	61,652	0.22%	+5,692	Jul 31, 2026	SCHWAB CAPITAL TRUST, not summed
26.3	Schwab U.S. Broad Market ETF Fund S000026631 · Registered fund · May 31, 2026 SCHWAB STRATEGIC TRUST, not summed	25,985	0.09%	+10,985	May 31, 2026	SCHWAB STRATEGIC TRUST, not summed
26.4	Schwab Total Stock Market Index Fund Fund S000005913 · Registered fund · Jul 31, 2026 SCHWAB CAPITAL TRUST, not summed	13,000	0.05%	0	Jul 31, 2026	SCHWAB CAPITAL TRUST, not summed
27	Point72 Asset Management, L.P. 13F CIK 1603466 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	231,582	0.81%	+167,737	Jun 30, 2026	carried, not yet filed for the newest quarter
28	SASSINE ANDY Insider CIK 1515322 · Aug 15, 2025 Director, Chief Financial Officer, issuer-wide	221,764	0.78%	+1,238	Aug 15, 2025	Director, Chief Financial Officer, issuer-wide
29	DIMENSIONAL FUND ADVISORS LP 13F CIK 354204 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	221,322	0.78%	-67,033	Jun 30, 2026	carried, not yet filed for the newest quarter
29.1	U.S. MICRO CAP PORTFOLIO Fund S000000948 · Registered fund · Jul 31, 2026 DFA INVESTMENT DIMENSIONS GROUP INC, not summed	69,508	0.24%	+67,520	Jul 31, 2026	DFA INVESTMENT DIMENSIONS GROUP INC, not summed
29.2	U.S. SMALL CAP PORTFOLIO Fund S000000971 · Registered fund · Jul 31, 2026 DFA INVESTMENT DIMENSIONS GROUP INC, not summed	33,975	0.12%	-5,557	Jul 31, 2026	DFA INVESTMENT DIMENSIONS GROUP INC, not summed
29.3	Dimensional U.S. Core Equity 2 ETF Fund S000070903 · Registered fund · Jul 31, 2026 Dimensional ETF Trust, not summed	24,073	0.08%	0	Jul 31, 2026	Dimensional ETF Trust, not summed
29.4	Dimensional U.S. Small Cap ETF Fund S000070906 · Registered fund · Jul 31, 2026 Dimensional ETF Trust, not summed	14,428	0.05%	-80,135	Jul 31, 2026	Dimensional ETF Trust, not summed
29.5	Dimensional US Core Equity Market ETF Fund S000069432 · Registered fund · Jul 31, 2026 Dimensional ETF Trust, not summed	3,714	0.01%	0	Jul 31, 2026	Dimensional ETF Trust, not summed
30	HEARTLAND ADVISORS INC 13F CIK 937394 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	219,300	0.77%	-36,700	Jun 30, 2026	carried, not yet filed for the newest quarter
31	NORTHERN TRUST CORP 13F CIK 73124 · Bank · Jun 30, 2026 carried, not yet filed for the newest quarter	204,578	0.72%	-15,035	Jun 30, 2026	carried, not yet filed for the newest quarter
31.1	Small Cap Index Fund Fund S000001245 · Registered fund · Jun 30, 2026 Northern Funds, not summed	11,693	0.04%	+1,257	Jun 30, 2026	Northern Funds, not summed
32	JPMORGAN CHASE & CO 13F CIK 19617 · Bank · Jun 30, 2026 carried, not yet filed for the newest quarter	195,704	0.69%	-2,111	Jun 30, 2026	carried, not yet filed for the newest quarter
33	VANGUARD FIDUCIARY TRUST CO 13F CIK 933478 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	174,048	0.61%	+4,236	Jun 30, 2026	carried, not yet filed for the newest quarter
34	Jump Financial, LLC 13F CIK 1831577 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	138,209	0.49%	+69,916	Jun 30, 2026	carried, not yet filed for the newest quarter
35	VANGUARD RUSSELL 2000 INDEX FUND Fund S000030003 · Registered fund · May 31, 2026 VANGUARD SCOTTSDALE FUNDS, not summed	136,565	0.48%	+11,352	May 31, 2026	VANGUARD SCOTTSDALE FUNDS, not summed
36	ACADIAN ASSET MANAGEMENT LLC 13F CIK 916542 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	131,451	0.46%	-149,049	Jun 30, 2026	carried, not yet filed for the newest quarter
37	IQ EQ FUND MANAGEMENT (IRELAND) Ltd 13F CIK 1620943 · Exempt reporting adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	126,390	0.44%	+12,135	Jun 30, 2026	carried, not yet filed for the newest quarter
38	VANGUARD STRATEGIC EQUITY FUND Fund S000002593 · Registered fund · Jun 30, 2026 VANGUARD HORIZON FUNDS, not summed	118,011	0.42%	0	Jun 30, 2026	VANGUARD HORIZON FUNDS, not summed
39	Invesco Ltd. 13F CIK 914208 · Investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	104,188	0.37%	-9,382	Jun 30, 2026	carried, not yet filed for the newest quarter
39.1	Invesco RAFI US 1500 Small-Mid ETF Fund S000013120 · Registered fund · Jul 31, 2026 Invesco Exchange-Traded Fund Trust, not summed	92,217	0.32%	-5,944	Jul 31, 2026	Invesco Exchange-Traded Fund Trust, not summed
39.2	Invesco Nasdaq Biotechnology ETF Fund S000072469 · Registered fund · May 31, 2026 Invesco Exchange-Traded Fund Trust II, not summed	1,489	0.01%	-121	May 31, 2026	Invesco Exchange-Traded Fund Trust II, not summed
40	MORGAN STANLEY 13F CIK 895421 · Holding company · Jun 30, 2026 carried, not yet filed for the newest quarter	97,406	0.34%	-28,729	Jun 30, 2026	carried, not yet filed for the newest quarter
41	Quantinno Capital Management LP 13F CIK 1759654 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	85,875	0.30%	+14,522	Jun 30, 2026	carried, not yet filed for the newest quarter
42	Bank of New York Mellon Corp 13F CIK 1390777 · Bank · Jun 30, 2026 carried, not yet filed for the newest quarter	80,314	0.28%	+5,317	Jun 30, 2026	carried, not yet filed for the newest quarter
43	VANGUARD HEALTH CARE INDEX FUND Fund S000004450 · Registered fund · May 31, 2026 VANGUARD WORLD FUND, not summed	78,018	0.27%	+4,143	May 31, 2026	VANGUARD WORLD FUND, not summed
44	MARSHALL WACE, LLP 13F CIK 1318757 · Exempt reporting adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	77,269	0.27%	+77,269	Jun 30, 2026	carried, not yet filed for the newest quarter
45	Russell Investments Group, Ltd. 13F CIK 1692234 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	75,903	0.27%	+44,199	Jun 30, 2026	carried, not yet filed for the newest quarter
45.1	Russell Investments U.S. Small Cap Equity ETF Fund S000089990 · Registered fund · Jun 30, 2026 Russell Investments Exchange Traded Funds, not summed	16,752	0.06%	—	Jun 30, 2026	Russell Investments Exchange Traded Funds, not summed
45.2	U.S. Small Cap Equity Fund Fund S000001586 · Registered fund · Jul 31, 2026 RUSSELL INVESTMENT CO, not summed	8,600	0.03%	-1,862	Jul 31, 2026	RUSSELL INVESTMENT CO, not summed
45.3	U.S. Small Cap Equity Fund Fund S000006864 · Registered fund · Jun 30, 2026 RUSSELL INVESTMENT FUNDS, not summed	2,245	0.01%	+494	Jun 30, 2026	RUSSELL INVESTMENT FUNDS, not summed
46	AMERICAN CENTURY COMPANIES INC 13F CIK 748054 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	65,361	0.23%	-651	Jun 30, 2026	carried, not yet filed for the newest quarter
47	AMERIPRISE FINANCIAL INC 13F CIK 820027 · Investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	65,282	0.23%	+65,282	Jun 30, 2026	carried, not yet filed for the newest quarter
48	MASSACHUSETTS FINANCIAL SERVICES CO /MA/ 13F CIK 912938 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	57,480	0.20%	-2,290	Jun 30, 2026	carried, not yet filed for the newest quarter
48.1	MFS Blended Research Small Cap Equity Fund Fund S000050659 · Registered fund · May 31, 2026 MFS SERIES TRUST X, not summed	38,570	0.14%	-1,493	May 31, 2026	MFS SERIES TRUST X, not summed
48.2	MFS Blended Research Small Cap Equity Portfolio Fund S000008071 · Registered fund · Jun 30, 2026 MFS VARIABLE INSURANCE TRUST III, not summed	9,719	0.03%	-720	Jun 30, 2026	MFS VARIABLE INSURANCE TRUST III, not summed
49	HSBC HOLDINGS PLC 13F CIK 873630 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	56,895	0.20%	+4,070	Jun 30, 2026	carried, not yet filed for the newest quarter
50	Avantis U.S. Small Cap Equity ETF Fund S000074835 · Registered fund · May 31, 2026 American Century ETF Trust, not summed	56,780	0.20%	-65	May 31, 2026	American Century ETF Trust, not summed

7 more nested rows are not shown here; Every filing lists them all.

Shares are restated onto today's split basis. Change compares each row with the same holder's previous filing in the same source; a dash means there is none to compare. Nested 13D/G, insider and fund rows are not summed, since a 13F report already holds them.

Showing 1–50 of 221 holders

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Key facts CIK 1768224 CUSIP 03969T109 13F (30d) 19 filings 19 filers Visit website Investor relations