ARWR · Arrowhead Pharmaceuticals, Inc. · Holdings

$61.48 -0.06 (-0.10%) At close · Oct 7

S&P 400 member Russell 1000 member Rebalance outlook locked

Market Cap

$9.36B

141.25M

Volume · Oct 7 1.82M Avg daily vol (3M) 1.9M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Use institutional holdings through the 13F API or MCP →

Some SEC filings for this period have not finished importing. Holder totals and changes may be incomplete; these filings remain queued for recovery.

The Sep 30, 2026 filing window is still open — showing the latest known positions. Funds that have not filed yet carry their prior-quarter positions; a fund that filed without this stock counts as sold out. Each fund's Change is measured against its own previous reported quarter, shown in the Reported column. Pick a completed quarter in the Quarter selector below to put every fund on one quarter and see the stock-wide activity breakdown.

Quarter:

Holders: 509

Total Value: $9,929,228,351

Total Shares: 121,854,036

Showing 1–100 of 524 holdings by value · page 1 of 6

Institutional Ownership Trend

27 quarters

Superinvestors Holding ARWR

All superinvestors →

Joel Greenblatt $545,790

Holder Concentration

Across 509 13F filers

Top Holder

12.2%

of institutional value

Top 5

38.3%

of institutional value

Top 10

55.0%

of institutional value

HHI

418

Broadly held

Showing 100 of 524

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

BlackRock, Inc.

$1,208,838,367

12.17%

14,830,553

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	197,668	$16,111,918	1.3%	Sole
BlackRock Investment Management (Australia) Ltd	331	$26,979	0.0%	Sole
BlackRock Asset Management Canada Ltd	17,655	$1,439,059	0.1%	Sole
BlackRock Investment Management, LLC	303,506	$24,738,774	2.0%	Sole
BLACKROCK ADVISORS LLC	143,392	$11,687,881	1.0%	Sole
BlackRock Fund Advisors	9,022,834	$735,451,199	60.8%	Sole
BlackRock Institutional Trust Company, N.A.	4,236,648	$345,329,178	28.6%	Sole
BlackRock Fund Managers Ltd.	22,363	$1,822,808	0.2%	Sole
BlackRock Investment Management (UK) Ltd.	54,434	$4,436,915	0.4%	Sole
BlackRock (Netherlands) B.V.	44,448	$3,622,956	0.3%	Sole
BlackRock Asset Management Ireland Ltd	498,788	$40,656,209	3.4%	Sole
BlackRock Advisors (UK) Ltd	2,140	$174,431	0.0%	Sole
BlackRock (Luxembourg) S.A.	2,036	$165,954	0.0%	Sole
BlackRock Life Ltd	511	$41,651	0.0%	Sole
BlackRock Asset Management Schweiz AG	20,091	$1,637,617	0.1%	Sole
Aperio Group, LLC	263,620	$21,487,666	1.8%	Sole
SpiderRock Advisors, LLC	88	$7,172	0.0%	Sole

Avoro Capital Advisors LLC

$925,591,288

9.32%

11,355,555

FMR LLC

$655,324,421

6.60%

8,039,804

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	4,810,131	$392,073,777	59.8%	Defined
FIDELITY MANAGEMENT TRUST CO	3,222,325	$262,651,710	40.1%	Defined
STRATEGIC ADVISERS LLC	5,293	$431,432	0.1%	Defined
FIAM LLC	260	$21,192	0.0%	Defined
Fidelity Diversifying Solutions LLC	1,795	$146,310	0.0%	Defined

VANGUARD PORTFOLIO MANAGEMENT LLC

$517,698,946

5.21%

6,351,355

VANGUARD CAPITAL MANAGEMENT LLC

$498,103,779

5.02%

6,110,953

STATE STREET CORP

$464,170,106

4.67%

5,694,640

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	5,694,640	$464,170,106	100.0%	Defined

NORGES BANK

$317,606,649

3.20%

3,896,536

Invesco Ltd.

$316,582,882

3.19%

3,883,976

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	1,612,017	$131,395,504	41.5%	Defined
Invesco Trust Co joint	55,200	$4,499,352	1.4%	Defined
Invesco Capital Management LLC	2,216,759	$180,688,026	57.1%	Defined

GEODE CAPITAL MANAGEMENT, LLC

$287,780,183

2.90%

3,530,612

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	543,049	$44,263,923	15.4%	Defined
Held directly	2,987,563	$243,516,260	84.6%	Defined

FRED ALGER MANAGEMENT, LLC

$269,139,417

2.71%

3,301,919

BAKER BROS. ADVISORS LP

$162,431,742

1.64%

1,992,783

Siren, L.L.C.

$139,828,448

1.41%

1,715,476

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invus Group, LLC	1,715,476	$139,828,448	100.0%	Defined

Vestal Point Capital, LP

$128,378,250

1.29%

1,575,000

DEERFIELD MANAGEMENT COMPANY, L.P.

$113,221,873

1.14%

1,389,055

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Deerfield UGP, LLC	1,389,055	$113,221,873	100.0%	Defined

FIRST TRUST ADVISORS LP

$110,276,346

1.11%

1,352,918

Driehaus Capital Management LLC

$110,083,819

1.11%

1,350,556

Jupiter Topco LLC

$99,572,289

1.00%

1,221,596

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	53,745	$4,380,754	4.4%	Defined
JANUS HENDERSON INVESTORS US LLC	1,167,851	$95,191,535	95.6%	Defined

NORTHERN TRUST CORP

$97,230,668

0.98%

1,192,868

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	1,042,873	$85,004,577	87.4%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	84,612	$6,896,723	7.1%	Defined
MASON STREET ADVISORS, LLC joint	65,383	$5,329,368	5.5%	Other

DIMENSIONAL FUND ADVISORS LP

$96,312,866

0.97%

1,181,608

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	2,609	$212,659	0.2%	Defined
DFA Australia Ltd.	35,091	$2,860,267	3.0%	Defined
Dimensional Fund Advisors Pte. Ltd.	793	$64,637	0.1%	Defined
Dimensional Ireland Ltd	68,660	$5,596,476	5.8%	Defined
Held directly	1,074,455	$87,578,827	90.9%	Sole

GOLDMAN SACHS GROUP INC

$93,737,069

0.94%

1,150,007

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	677,498	$55,222,861	58.9%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	465,257	$37,923,098	40.5%	Defined
Goldman Sachs Trust Company, N.A.	7,252	$591,110	0.6%	Defined

BANK OF AMERICA CORP /DE/

$92,905,911

0.94%

1,139,810

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	258,459	$21,066,993	22.7%	Defined
MERRILL LYNCH, PIERCE, FENNER & SMITH INC.	125,900	$10,262,109	11.0%	Defined
BOFA SECURITIES, INC.	605,207	$49,330,422	53.1%	Defined
MERRILL LYNCH INTERNATIONAL	3,351	$273,140	0.3%	Defined
MANAGED ACCOUNT ADVISORS LLC joint	146,893	$11,973,247	12.9%	Defined

Hood River Capital Management LLC

$91,352,087

0.92%

1,120,747

Holocene Advisors, LP

$90,164,487

0.91%

1,106,177

MORGAN STANLEY

$88,424,894

0.89%

1,084,835

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	17,727	$1,444,927	1.6%	Defined
MORGAN STANLEY & CO. LLC	63,783	$5,198,952	5.9%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	389,785	$31,771,375	35.9%	Defined
MORGAN STANLEY AIP GP LP	773	$63,006	0.1%	Defined
MORGAN STANLEY INVESTMENT MANAGEMENT INC	7,580	$617,845	0.7%	Defined
Morgan Stanley Investment Management LTD	10,084	$821,946	0.9%	Defined
MORGAN STANLEY SMITH BARNEY LLC	55,568	$4,529,347	5.1%	Defined
Parametric Sas	500	$40,755	0.0%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	506,375	$41,274,625	46.7%	Defined
EATON VANCE MANAGEMENT	7,000	$570,570	0.6%	Defined
Calvert Research & Management	25,660	$2,091,546	2.4%	Defined

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$78,614,678

0.79%

964,479

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	57,251	$4,666,529	5.9%	Defined
UBS Asset Management Switzerland AG	566,372	$46,164,979	58.7%	Defined
UBS Asset Management (UK) Ltd.	19,337	$1,576,158	2.0%	Defined
Held directly	321,519	$26,207,012	33.3%	Defined

Fisher Asset Management, LLC

$78,061,393

0.79%

957,691

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$75,309,289

0.76%

923,927

VANGUARD FIDUCIARY TRUST CO

$73,093,114

0.74%

896,738

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$70,718,076

0.71%

867,600

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	867,600	$70,718,076	100.0%	Other

UBS Group AG

$70,222,494

0.71%

861,520

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	205,408	$16,742,806	23.8%	Defined
UBS AG	363,324	$29,614,539	42.2%	Defined
UBS Switzerland AG	292,788	$23,865,149	34.0%	Defined

Bank of New York Mellon Corp

$70,060,935

0.71%

859,538

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNY Mellon Trust of Delaware joint	2,850	$232,302	0.3%	Defined
Mellon Investments Corporation joint	382,604	$31,186,051	44.5%	Other
BNY Mellon Investment Adviser, Inc.	52,210	$4,255,637	6.1%	Defined
The Bank of New York Mellon	307,692	$25,079,974	35.8%	Defined
BNY Mellon, NA joint	91,581	$7,464,765	10.7%	Defined
BNY Mellon Advisors, Inc.	2,753	$224,397	0.3%	Defined
Newton Investment Management North America, LLC joint	19,846	$1,617,646	2.3%	Defined
Held directly	2	$163	0.0%	Defined

PRICE T ROWE ASSOCIATES INC /MD/

$69,557,372

0.70%

853,360

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	137,696	$11,223,600	16.1%	Defined
Held directly	715,664	$58,333,772	83.9%	Sole

SUMMIT PARTNERS PUBLIC ASSET MANAGEMENT, LLC

$67,673,432

0.68%

830,247

Western Financial Corp/CA

$66,857,680

0.67%

820,239

JPMORGAN CHASE & CO

$51,954,389

0.52%

637,399

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	130,145	$10,608,118	20.4%	Defined
J.P. Morgan Investment Management Inc.	325,400	$26,523,354	51.1%	Defined
J.P. Morgan Securities LLC	27,904	$2,274,455	4.4%	Defined
J.P. Morgan Securities plc	2,041	$166,361	0.3%	Defined
JPMorgan Asset Management (UK) Ltd	150,523	$12,269,129	23.6%	Defined
J.P. MORGAN MANSART MANAGEMENT Ltd	1,386	$112,972	0.2%	Defined

Connor, Clark & Lunn Investment Management Ltd.

$50,990,129

0.51%

625,569

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 1	625,569	$50,990,129	100.0%	Sole

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$48,626,665

0.49%

596,573

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	596,573	$48,626,665	100.0%	Other

TWO SIGMA INVESTMENTS, LP

$47,325,439

0.48%

580,609

Pictet Asset Management Holding SA

$46,739,382

0.47%

573,419

BNP PARIBAS ARBITRAGE, SA

$45,534,990

0.46%

558,643

LORD, ABBETT & CO. LLC

$44,053,383

0.44%

540,466

1832 Asset Management L.P.

$43,893,135

0.44%

538,500

PRINCIPAL FINANCIAL GROUP INC

$41,949,773

0.42%

514,658

HRT FINANCIAL LP

$40,645,369

0.41%

498,655

VOYA INVESTMENT MANAGEMENT LLC

$39,409,675

0.40%

483,495

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	483,495	$39,409,675	100.0%	Defined

Balyasny Asset Management L.P.

$38,371,158

0.39%

470,754

RENAISSANCE TECHNOLOGIES LLC

$38,221,750

0.38%

468,921

FRANKLIN RESOURCES INC

$37,668,456

0.38%

462,133

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Brandywine Global Investment Management, LLC	158	$12,878	0.0%	Defined
FIDUCIARY TRUST CO INTERNATIONAL	41	$3,341	0.0%	Defined
Fiduciary Trust International of California	3	$244	0.0%	Defined
FIDUCIARY TRUST INTERNATIONAL LLC	592	$48,253	0.1%	Defined
FRANKLIN ADVISERS INC	131,995	$10,758,912	28.6%	Defined
PUTNAM INVESTMENT MANAGEMENT LLC	210,264	$17,138,618	45.5%	Defined
PUTNAM ADVISORY CO LLC	119,080	$9,706,210	25.8%	Defined

D. E. Shaw & Co., Inc.

$35,176,454

0.35%

431,560

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	322,317	$26,272,058	74.7%	Defined
D. E. Shaw Investment Management, L.L.C. joint	109,243	$8,904,396	25.3%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$34,600,995

0.35%

424,500

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	424,500	$34,600,995	100.0%	Other

Nuveen, LLC

$33,898,621

0.34%

415,883

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	113,997	$9,291,894	27.4%	Defined
TIAA CREF INVESTMENT MANAGEMENT LLC joint	113,256	$9,231,496	27.2%	Defined
TEACHERS ADVISORS, LLC joint	188,630	$15,375,231	45.4%	Defined

RHUMBLINE ADVISERS

$30,557,854

0.31%

374,897

EVENTIDE ASSET MANAGEMENT, LLC

$30,158,700

0.30%

370,000

AMERICAN CENTURY COMPANIES INC

$29,686,838

0.30%

364,211

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AMERICAN CENTURY INVESTMENT MANAGEMENT INC	364,211	$29,686,838	100.0%	Defined

MILLENNIUM MANAGEMENT LLC

$26,844,258

0.27%

329,337

Caption Management, LLC

$25,724,556

0.26%

315,600

LPL Financial LLC

$25,330,129

0.26%

310,761

Artisan Partners Limited Partnership

$25,325,646

0.26%

310,706

MARSHALL WACE, LLP

$24,863,075

0.25%

305,031

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marshall Wace North America L.P. joint	305,031	$24,863,075	100.0%	Other

SEI INVESTMENTS CO

$24,503,454

0.25%

300,619

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI INVESTMENTS MANAGEMENT CORP	190,277	$15,509,478	63.3%	Sole
SEI TRUST CO	110,342	$8,993,976	36.7%	Sole

NORDEA INVESTMENT MANAGEMENT AB

$23,163,103

0.23%

284,175

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nordea Funds Ltd	66,831	$5,447,394	23.5%	Defined
Nordea Investment Funds S.A.	217,344	$17,715,709	76.5%	Defined

Russell Investments Group, Ltd.

$23,156,662

0.23%

284,096

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC	194,299	$15,837,310	68.4%	Defined
Russell Investments Canada Limited	246	$20,051	0.1%	Defined
Russell Investments Capital, LLC	46,800	$3,814,668	16.5%	Defined
Russell Investments France SAS	14,075	$1,147,253	5.0%	Defined
Russell Investments Limited	414	$33,745	0.1%	Defined
Russell Investments Trust Company	28,262	$2,303,635	9.9%	Defined

CITADEL ADVISORS LLC

$21,755,019

0.22%

266,900

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	266,900	$21,755,019	100.0%	Defined

CITIGROUP INC

$21,573,903

0.22%

264,678

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citibank, N.A. joint	42	$3,423	0.0%	Defined
Citigroup Financial Products Inc. joint	264,636	$21,570,480	100.0%	Defined

California Public Employees Retirement System

$20,892,154

0.21%

256,314

Squarepoint Ops LLC

$20,858,979

0.21%

255,907

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Squarepoint Capital LLP	255,907	$20,858,979	100.0%	Defined

ENVESTNET ASSET MANAGEMENT INC

$20,722,124

0.21%

254,228

Aberdeen Group plc

$20,537,911

0.21%

251,968

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
abrdn Inc.	251,968	$20,537,911	100.0%	Defined

Swiss National Bank

$19,472,739

0.20%

238,900

Legal & General Group Plc

$19,335,475

0.19%

237,216

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	88,930	$7,248,684	37.5%	Defined
Legal & General Investment Management Ltd joint	148,286	$12,086,791	62.5%	Defined

Privium Fund Management B.V.

$19,091,435

0.19%

234,222

ARROWSTREET CAPITAL, LIMITED PARTNERSHIP

$17,780,020

0.18%

218,133

Life Cycle Investment Partners Ltd

$17,041,132

0.17%

209,068

HSBC HOLDINGS PLC

$16,611,738

0.17%

203,800

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
HSBC GLOBAL ASSET MANAGEMENT (FRANCE)	110,000	$8,966,100	54.0%	Defined
HSBC BANK PLC	93,800	$7,645,638	46.0%	Defined

Assenagon Asset Management S.A.

$16,459,069

0.17%

201,927

Woodline Partners LP

$16,316,997

0.16%

200,184

Integral Health Asset Management, LLC

$16,302,000

0.16%

200,000

AQR CAPITAL MANAGEMENT LLC

$16,207,609

0.16%

198,842

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Arax Advisory Partners	1,044	$85,096	0.5%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	1,092	$89,008	0.5%	Other
Stokes Family Office, LLC	1,160	$94,551	0.6%	Other
Held directly	195,546	$15,938,954	98.3%	Defined

JANE STREET GROUP, LLC

$15,992,669

0.16%

196,205

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	88,261	$7,194,154	45.0%	Defined
Jane Street Global Trading, LLC	107,944	$8,798,515	55.0%	Defined

MUTUAL OF AMERICA CAPITAL MANAGEMENT LLC

$15,603,622

0.16%

191,432

ALLIANCEBERNSTEIN L.P.

$14,122,830

0.14%

173,265

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	173,265	$14,122,830	100.0%	Defined

WCM INVESTMENT MANAGEMENT, LLC

$13,957,120

0.14%

171,232

PINNACLE ASSOCIATES LTD

$13,867,377

0.14%

170,131

RAYMOND JAMES FINANCIAL INC

$13,849,607

0.14%

169,913

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	9,763	$795,782	5.7%	Defined
Raymond James Financial Services Advisors, Inc.	16,101	$1,312,392	9.5%	Defined
ClariVest Asset Management LLC joint	64,013	$5,217,699	37.7%	Defined
CHARTWELL INVESTMENT PARTNERS, LLC	80,036	$6,523,734	47.1%	Defined

SEB Asset Management AB

$12,895,289

0.13%

158,205

Clearbridge Investments, LLC

$12,806,036

0.13%

157,110

AMERIPRISE FINANCIAL INC

$12,708,060

0.13%

155,908

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	148,851	$12,132,845	95.5%	Defined
Ameriprise Financial Services, LLC	6,972	$568,287	4.5%	Defined
Ameriprise Bank, FSB	85	$6,928	0.1%	Defined

Affinity Asset Advisors, LLC

$12,226,500

0.12%

150,000

DEUTSCHE BANK AG\

$12,164,550

0.12%

149,240

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Deutsche Bank Trust Co Americas	5	$407	0.0%	Defined
DWS Investment GmbH	100,852	$8,220,446	67.6%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC	22,654	$1,846,527	15.2%	Defined
DWS Investments UK Ltd	23,665	$1,928,934	15.9%	Defined
DWS International GmbH	2,064	$168,236	1.4%	Defined

WEALTH EFFECTS LLC

$12,006,423

0.12%

147,300

Retirement Systems of Alabama

$11,874,947

0.12%

145,687

TORONTO DOMINION BANK

$11,813,080

0.12%

144,928

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
TD SECURITIES INC	1	$81	0.0%	Defined
Held directly	144,927	$11,812,999	100.0%	Sole

HighTower Advisors, LLC

$11,443,514

0.12%

140,394

CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM

$11,397,217

0.11%

139,826

JANE STREET GROUP, LLC

$11,134,266

0.11%

136,600

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	136,600	$11,134,266	100.0%	Defined

Weiss Asset Management LP

$10,923,481

0.11%

134,014

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WEISS ANDREW M joint	134,014	$10,923,481	100.0%	Defined

BANK OF NOVA SCOTIA

$10,906,445

0.11%

133,805

PFM Health Sciences, LP

$10,732,177

0.11%

131,667

Tetragon Partners GP Ltd

$10,635,750

0.11%

130,484

WELLS FARGO & COMPANY/MN

$10,497,589

0.11%

128,789

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	97,208	$7,923,424	75.5%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	3,280	$267,352	2.5%	Defined
WELLS FARGO CLEARING SERVICES, LLC	28,254	$2,302,983	21.9%	Defined
WELLS FARGO INVESTMENT INSTITUTE, INC.	47	$3,830	0.0%	Defined

Key facts CIK 879407 CUSIP 04280A100 13F (30d) 11 filings 11 filers Visit website Investor relations