ATYR · aTYR PHARMA INC · Shareholders & Ownership

$0.17 0.00 (-0.58%) At close · Oct 21

Market Cap

$17.33M

98.09M

Volume · Oct 21 1.23M Avg daily vol (3M) 3.25M

Every disclosed holder of ATYR in one list: 13F institutions, Schedule 13D and 13G filers, insiders (Forms 3, 4 and 5) and registered funds, each from its own latest filing. Holders nest by CIK and SEC reporting links, never by name, and each is summed once.

How the holder list picks a filing →

Who Owns ATYR

13F quarter Sep 30, 2026, combined view: filers that have not filed yet carry their Jun 30, 2026 positions

Holder group	Holders	Shares	% Out	Summed
Institutional (13F)	84	38,759,831	39.52%	Yes
Insiders (Forms 3, 4, 5)	5	1,655,636	1.69%	Yes
13D/G-only blocks	3	2,817,271	2.87%	Yes
Remaining	—	54,854,687	55.92%	Rest
Registered funds (N-PORT, not summed)	40	20,244,620	20.64%	No

Shares outstanding: 98,087,425, the issuer's latest cover-page count restated onto today's split basis.

13D/G-only counts each filer's latest 13D or 13G that no 13F report of the listing contains, that names no 13F filer and that no counted insider files. Funds are never summed: their shares sit inside their advisers' 13F reports.

Holders of ATYR

One row per holder Every filing By ultimate parent

All 13F Funds Insiders 13D/G Non-US

#	Holder	Shares	% Out	Change	As of	Notes
1	FEDERATED HERMES, INC. 13G CIK 1056288 · Investment adviser · Sep 30, 2026 Rule 13d-1(b), qualified institution, not summed	4,965,414	5.06%	-3,544,264	Sep 30, 2026	Rule 13d-1(b), qualified institution, not summed
1.1	FEDERATED HERMES, INC. 13F CIK 1056288 · Investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	8,509,678	8.68%	-2,000,000	Jun 30, 2026	carried, not yet filed for the newest quarter
2	Federated Hermes Kaufmann Fund Fund S000009729 · Registered fund · Jul 31, 2026 Federated Hermes Equity Funds, not summed	4,367,290	4.45%	-1,217,710	Jul 31, 2026	Federated Hermes Equity Funds, not summed
3	Federated Hermes Kaufmann Small Cap Fund Fund S000009730 · Registered fund · Jul 31, 2026 Federated Hermes Equity Funds, not summed	4,061,288	4.14%	-745,589	Jul 31, 2026	Federated Hermes Equity Funds, not summed
4	VANGUARD CAPITAL MANAGEMENT LLC 13F CIK 2100119 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	3,808,091	3.88%	-4,041	Jun 30, 2026	carried, not yet filed for the newest quarter
5	GSA CAPITAL PARTNERS LLP 13F CIK 1362033 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	3,454,863	3.52%	+1,770,277	Jun 30, 2026	carried, not yet filed for the newest quarter
6	VANGUARD TOTAL STOCK MARKET INDEX FUND Fund S000002848 · Registered fund · Jun 30, 2026 VANGUARD INDEX FUNDS, not summed	2,650,788	2.70%	0	Jun 30, 2026	VANGUARD INDEX FUNDS, not summed
7	AQR CAPITAL MANAGEMENT LLC 13F CIK 1167557 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	2,610,548	2.66%	+2,114,530	Jun 30, 2026	carried, not yet filed for the newest quarter
8	FMR LLC 13F CIK 315066 · Holding company · Jun 30, 2026 carried, not yet filed for the newest quarter	2,471,418	2.52%	-12,235,425	Jun 30, 2026	carried, not yet filed for the newest quarter
8.1	Fidelity Growth Company Fund Fund S000007119 · Registered fund · May 31, 2026 Fidelity Mt. Vernon Street Trust, not summed	4,216,832	4.30%	-12,000	May 31, 2026	Fidelity Mt. Vernon Street Trust, not summed
8.2	FMR LLC 13G CIK 315066 · Holding company · Jun 30, 2026 Rule 13d-1(b), qualified institution, not summed	2,471,418	2.52%	-3,464,482	Jun 30, 2026	Rule 13d-1(b), qualified institution, not summed
8.3	Select Biotechnology Portfolio Fund S000007486 · Registered fund · May 31, 2026 Fidelity Select Portfolios, not summed	1,754,594	1.79%	0	May 31, 2026	Fidelity Select Portfolios, not summed
8.4	Fidelity Extended Market Index Fund Fund S000006021 · Registered fund · May 31, 2026 Fidelity Concord Street Trust, not summed	550,058	0.56%	+18,704	May 31, 2026	Fidelity Concord Street Trust, not summed
8.5	Fidelity Total Market Index Fund Fund S000006026 · Registered fund · May 31, 2026 Fidelity Concord Street Trust, not summed	168,117	0.17%	0	May 31, 2026	Fidelity Concord Street Trust, not summed
9	BlackRock, Inc. 13F CIK 2012383 · Holding company · Jun 30, 2026 carried, not yet filed for the newest quarter	1,827,332	1.86%	-4,616,572	Jun 30, 2026	carried, not yet filed for the newest quarter
9.1	BlackRock, Inc. 13G CIK 2012383 · Holding company · Jun 30, 2026 Rule 13d-1(b), qualified institution, not summed	1,725,072	1.76%	-4,038,534	Jun 30, 2026	Rule 13d-1(b), qualified institution, not summed
9.2	iShares Core S&P Total U.S. Stock Market ETF Fund S000004317 · Registered fund · Jun 30, 2026 iShares Trust, not summed	146,027	0.15%	+3,536	Jun 30, 2026	iShares Trust, not summed
9.3	Diversified Equity Alpha Master Portfolio Fund S000004282 · Registered fund · Jun 30, 2026 Master Investment Portfolio, not summed	18,223	0.02%	-4,404	Jun 30, 2026	Master Investment Portfolio, not summed
10	Kalehua Capital Management LLC 13F CIK 2106181 · Exempt reporting adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	1,788,167	1.82%	+1,788,167	Jun 30, 2026	carried, not yet filed for the newest quarter
11	Logos Global Management LP 13G CIK 1792126 · Registered investment adviser · Sep 30, 2024 Rule 13d-1(b), qualified institution	1,750,000	1.78%	-4,050,000	Sep 30, 2024	Rule 13d-1(b), qualified institution
12	JANE STREET GROUP, LLC 13F CIK 1595888 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	1,721,297	1.75%	+1,672,795	Jun 30, 2026	carried, not yet filed for the newest quarter
13	UBS Group AG 13F CIK 1610520 · Bank · Jun 30, 2026 carried, not yet filed for the newest quarter	1,531,151	1.56%	-2,336,062	Jun 30, 2026	carried, not yet filed for the newest quarter
14	SCHIMMEL PAUL Insider CIK 1236447 · May 27, 2026 Director, issuer-wide	1,421,913	1.45%	—	May 27, 2026	Director, issuer-wide
15	MORGAN STANLEY 13F CIK 895421 · Holding company · Jun 30, 2026 carried, not yet filed for the newest quarter	1,343,636	1.37%	+118,169	Jun 30, 2026	carried, not yet filed for the newest quarter
16	TWO SIGMA INVESTMENTS, LP 13F CIK 1179392 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	1,245,147	1.27%	+100,582	Jun 30, 2026	carried, not yet filed for the newest quarter
17	CITADEL ADVISORS LLC 13F CIK 1423053 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	1,242,067	1.27%	+1,194,695	Jun 30, 2026	carried, not yet filed for the newest quarter
18	GEODE CAPITAL MANAGEMENT, LLC 13F CIK 1214717 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	1,177,020	1.20%	-1,208,465	Jun 30, 2026	carried, not yet filed for the newest quarter
19	VANGUARD EXTENDED MARKET INDEX FUND Fund S000002841 · Registered fund · Jun 30, 2026 VANGUARD INDEX FUNDS, not summed	1,054,556	1.08%	+48,153	Jun 30, 2026	VANGUARD INDEX FUNDS, not summed
20	SUSQUEHANNA INTERNATIONAL GROUP, LLP 13F CIK 1446194 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	747,039	0.76%	-119,705	Jun 30, 2026	carried, not yet filed for the newest quarter
21	MILLENNIUM MANAGEMENT LLC 13F CIK 1273087 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	656,051	0.67%	-985,195	Jun 30, 2026	carried, not yet filed for the newest quarter
22	GOLDMAN SACHS GROUP INC 13F CIK 886982 · Holding company · Jun 30, 2026 carried, not yet filed for the newest quarter	641,208	0.65%	+357,972	Jun 30, 2026	carried, not yet filed for the newest quarter
22.1	Goldman Sachs Absolute Return Tracker Fund Fund S000020314 · Registered fund · Jun 30, 2026 GOLDMAN SACHS TRUST, not summed	66,034	0.07%	+24,585	Jun 30, 2026	GOLDMAN SACHS TRUST, not summed
23	Lytton, Laurence W. 13G CIK 1105806 · Unknown · Dec 31, 2023 Rule 13d-1(c), passive investor	585,013	0.60%	—	Dec 31, 2023	Rule 13d-1(c), passive investor
24	VANGUARD FIDUCIARY TRUST CO 13F CIK 933478 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	576,125	0.59%	-75,707	Jun 30, 2026	carried, not yet filed for the newest quarter
25	STATE STREET CORP 13F CIK 93751 · Bank · Jun 30, 2026 carried, not yet filed for the newest quarter	494,951	0.50%	-1,259,448	Jun 30, 2026	carried, not yet filed for the newest quarter
25.1	State Street Balanced Index Fund Fund S000082802 · Registered fund · Jun 30, 2026 STATE STREET INSTITUTIONAL INVESTMENT TRUST, not summed	3,393	0.00%	0	Jun 30, 2026	STATE STREET INSTITUTIONAL INVESTMENT TRUST, not summed
26	VANGUARD RUSSELL 2000 INDEX FUND Fund S000030003 · Registered fund · May 31, 2026 VANGUARD SCOTTSDALE FUNDS, not summed	488,823	0.50%	+13,051	May 31, 2026	VANGUARD SCOTTSDALE FUNDS, not summed
27	Stonepine Capital Management, LLC 13G CIK 1440771 · Registered investment adviser · Dec 31, 2023 Rule 13d-1(b), qualified institution	482,258	0.49%	—	Dec 31, 2023	Rule 13d-1(b), qualified institution
28	MARSHALL WACE, LLP 13F CIK 1318757 · Exempt reporting adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	368,120	0.38%	+230,002	Jun 30, 2026	carried, not yet filed for the newest quarter
29	XTX Topco Ltd 13F CIK 1828301 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	226,678	0.23%	+115,255	Jun 30, 2026	carried, not yet filed for the newest quarter
30	NORTHERN TRUST CORP 13F CIK 73124 · Bank · Jun 30, 2026 carried, not yet filed for the newest quarter	220,141	0.22%	-490,964	Jun 30, 2026	carried, not yet filed for the newest quarter
31	BANK OF AMERICA CORP /DE/ 13F CIK 70858 · Bank · Jun 30, 2026 carried, not yet filed for the newest quarter	207,468	0.21%	-96,091	Jun 30, 2026	carried, not yet filed for the newest quarter
32	RENAISSANCE TECHNOLOGIES LLC 13F CIK 1037389 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	165,222	0.17%	+124,700	Jun 30, 2026	carried, not yet filed for the newest quarter
33	Shukla Sanjay Insider CIK 1671026 · Feb 4, 2026 Director, President and CEO, issuer-wide	153,553	0.16%	+6,630	Feb 4, 2026	Director, President and CEO, issuer-wide
34	JPMORGAN CHASE & CO 13F CIK 19617 · Bank · Jun 30, 2026 carried, not yet filed for the newest quarter	150,666	0.15%	+108,679	Jun 30, 2026	carried, not yet filed for the newest quarter
35	GROUP ONE TRADING LLC 13F CIK 932540 · Broker-dealer · Jun 30, 2026 carried, not yet filed for the newest quarter	138,878	0.14%	-45,136	Jun 30, 2026	carried, not yet filed for the newest quarter
36	Qube Research & Technologies Ltd 13F CIK 1729829 · Exempt reporting adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	126,996	0.13%	+126,996	Jun 30, 2026	carried, not yet filed for the newest quarter
37	HRT FINANCIAL LP 13F CIK 1475597 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	126,681	0.13%	+126,681	Jun 30, 2026	carried, not yet filed for the newest quarter
38	DIMENSIONAL FUND ADVISORS LP 13F CIK 354204 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	90,861	0.09%	-1,010	Jun 30, 2026	carried, not yet filed for the newest quarter
39	VANGUARD RUSSELL 2000 GROWTH INDEX FUND Fund S000030005 · Registered fund · May 31, 2026 VANGUARD SCOTTSDALE FUNDS, not summed	85,881	0.09%	+2,504	May 31, 2026	VANGUARD SCOTTSDALE FUNDS, not summed
40	Federated Hermes Kaufmann Fund II Fund S000009751 · Registered fund · Jun 30, 2026 Federated Hermes Insurance Series, not summed	81,100	0.08%	-36,701	Jun 30, 2026	Federated Hermes Insurance Series, not summed
41	HSBC HOLDINGS PLC 13F CIK 873630 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	66,185	0.07%	+47,971	Jun 30, 2026	carried, not yet filed for the newest quarter
42	Mariner, LLC 13F CIK 1373442 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	58,658	0.06%	+58,658	Jun 30, 2026	carried, not yet filed for the newest quarter
43	VANGUARD BALANCED INDEX FUND Fund S000002560 · Registered fund · Jun 30, 2026 VANGUARD VALLEY FORGE FUNDS, not summed	56,767	0.06%	-79	Jun 30, 2026	VANGUARD VALLEY FORGE FUNDS, not summed
44	HOHIMER WEALTH MANAGEMENT, LLC 13F CIK 1802533 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	55,000	0.06%	-45,000	Jun 30, 2026	carried, not yet filed for the newest quarter
45	MERCER GLOBAL ADVISORS INC /ADV 13F CIK 853758 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	53,531	0.05%	+38,326	Jun 30, 2026	carried, not yet filed for the newest quarter
46	KINGSWOOD WEALTH ADVISORS, LLC 13F CIK 1987855 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	52,000	0.05%	0	Jun 30, 2026	carried, not yet filed for the newest quarter
47	Schonfeld Strategic Advisors LLC 13F CIK 1665241 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	50,880	0.05%	+50,880	Jun 30, 2026	carried, not yet filed for the newest quarter
48	ADAR1 Capital Management, LLC 13F CIK 1940272 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	50,685	0.05%	0	Jun 30, 2026	carried, not yet filed for the newest quarter
49	Alpha Wealth Funds, LLC 13F CIK 2059327 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	50,000	0.05%	+50,000	Jun 30, 2026	carried, not yet filed for the newest quarter
50	Tower Research Capital LLC (TRC) 13F CIK 1533421 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	48,768	0.05%	+31,342	Jun 30, 2026	carried, not yet filed for the newest quarter

7 more nested rows are not shown here; Every filing lists them all.

Shares are restated onto today's split basis. Change compares each row with the same holder's previous filing in the same source; a dash means there is none to compare. Nested 13D/G, insider and fund rows are not summed, since a 13F report already holds them.

Showing 1–50 of 117 holders

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Key facts CIK 1339970 CUSIP 002120202 13F (30d) 9 filings 9 filers Visit website Investor relations