ATYR · aTYR PHARMA INC

$0.56 +0.01 (+1.16%)

Market Cap

$54.05M

98.09M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Quarter:

Holders: 93

Total Value: $23,669,474

Total Shares: 39,647,456

Institutional Ownership Trend

26 quarters

Holder Concentration

Across 93 13F filers · Δ vs Mar 31, 2026

Top Holder

21.5%

−4.4 pts

Top 5

52.6%

−16.5 pts

Top 10

74.3%

−10.0 pts

HHI

855

Broadly held−435

Quarterly Activity

vs Mar 31, 2026

New

Increased

Decreased

Sold Out

Unchanged

Holder count: -20

Institutional value: -46.7%

Shares held: -30.4%

Top Buyers

vs Mar 31, 2026

Institution	Added	Now Holds
Mega AQR CAPITAL MANAGEMENT LLC	+2,114,530	2,610,548
Mid GSA CAPITAL PARTNERS LLP	+1,770,277	3,454,863
Mega JANE STREET GROUP, LLC	+1,413,495	1,914,897
Mega CITADEL ADVISORS LLC	+1,344,895	1,548,267
Mega GOLDMAN SACHS GROUP INC	+357,972	641,208

Top Sellers

vs Mar 31, 2026

Institution	Sold	Now Holds
Mega FMR LLC	−12,235,425	2,471,418
Mega BlackRock, Inc.	−4,616,572	1,827,332
Mega UBS Group AG	−2,336,062	1,531,151
Large FEDERATED HERMES, INC.	−2,000,000	8,509,678
Mega STATE STREET CORP	−1,259,448	494,951

Top New Positions

vs Mar 31, 2026

Institution	Acquired	Value (USD)
Small Kalehua Capital Management LLC	1,788,167	$1,067,535
Mega Qube Research & Technologies Ltd	126,996	$75,816
Large HRT FINANCIAL LP	126,681	$75,628
Large Mariner, LLC	58,658	$35,018
Large Schonfeld Strategic Advisors LLC	50,880	$30,375

Top Sold-Out Positions

vs Mar 31, 2026

Institution	Previously Held	Last Value (USD)
Mega VANGUARD PORTFOLIO MANAGEMENT LLC	569,227	$443,997
Mega Nuveen, LLC	229,284	$178,840
Mega CHARLES SCHWAB INVESTMENT MANAGEMENT INC	210,737	$164,374
Mega Bank of New York Mellon Corp	193,531	$150,952
Mega DEUTSCHE BANK AG\	103,326	$80,593

Filter: New 17 Increased 29 Reduced 32 Sold out 37 Unchanged 15

Showing 100 of 100

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

FEDERATED HERMES, INC.

$5,080,277

21.46%

8,509,678

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FEDERATED GLOBAL INVESTMENT MANAGEMENT CORP	8,509,678	$5,080,277	100.0%	Defined

VANGUARD CAPITAL MANAGEMENT LLC

$2,273,430

9.60%

3,808,091

GSA CAPITAL PARTNERS LLP

$2,062,553

8.71%

3,454,863

AQR CAPITAL MANAGEMENT LLC

$1,558,495

6.58%

2,610,548

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RF&L WEALTH MANAGEMENT, LLC	7	$4	0.0%	Other
Prism Planning Partners LLC	491	$293	0.0%	Other
Compound Planning, Inc.	45	$26	0.0%	Other
Arax Advisory Partners	14,183	$8,467	0.5%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	1,362	$813	0.1%	Other
Stokes Family Office, LLC	3,063	$1,828	0.1%	Other
Held directly	2,591,397	$1,547,064	99.3%	Defined

FMR LLC

$1,475,436

6.23%

2,471,418

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	2,471,418	$1,475,436	100.0%	Defined

BlackRock, Inc.

$1,090,914

4.61%

1,827,332

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	60,274	$35,983	3.3%	Sole
BlackRock Investment Management, LLC	1,574	$939	0.1%	Sole
BlackRock Fund Advisors	164,380	$98,134	9.0%	Sole
BlackRock Institutional Trust Company, N.A.	1,596,195	$952,928	87.4%	Sole
BlackRock Investment Management (UK) Ltd.	4,909	$2,930	0.3%	Sole

Kalehua Capital Management LLC

$1,067,535

4.51%

1,788,167

JANE STREET GROUP, LLC

$1,027,614

4.34%

1,721,297

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	616,918	$368,300	35.8%	Defined
Jane Street Global Trading, LLC	1,104,379	$659,314	64.2%	Defined

UBS Group AG

$914,096

3.86%

1,531,151

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS AG	1,330,780	$794,475	86.9%	Defined
UBS Switzerland AG	188,389	$112,468	12.3%	Defined
UBS Securities LLC	11,982	$7,153	0.8%	Defined

MORGAN STANLEY

$802,149

3.39%

1,343,636

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	8,438	$5,037	0.6%	Defined
MORGAN STANLEY & CO. LLC	4,977	$2,971	0.4%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	1,325,798	$791,501	98.7%	Defined
MORGAN STANLEY SMITH BARNEY LLC	300	$179	0.0%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	4,123	$2,461	0.3%	Defined

TWO SIGMA INVESTMENTS, LP

$743,352

3.14%

1,245,147

CITADEL ADVISORS LLC

$741,513

3.13%

1,242,067

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	1,207,073	$720,622	97.2%	Defined
Held directly	34,994	$20,891	2.8%	Defined

GEODE CAPITAL MANAGEMENT, LLC

$702,680

2.97%

1,177,020

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	214,416	$128,006	18.2%	Defined
Held directly	962,604	$574,674	81.8%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$445,982

1.88%

747,039

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	747,039	$445,982	100.0%	Other

MILLENNIUM MANAGEMENT LLC

$391,662

1.65%

656,051

GOLDMAN SACHS GROUP INC

$382,800

1.62%

641,208

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	379,788	$226,733	59.2%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	261,420	$156,067	40.8%	Defined

VANGUARD FIDUCIARY TRUST CO

$343,946

1.45%

576,125

STATE STREET CORP

$295,485

1.25%

494,951

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SSGA FUNDS MANAGEMENT, INC. joint	494,951	$295,485	100.0%	Defined

MARSHALL WACE, LLP

$219,766

0.93%

368,120

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marshall Wace North America L.P. joint	368,120	$219,766	100.0%	Other

CITADEL ADVISORS LLC

$148,653

0.63%

249,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	249,000	$148,653	100.0%	Defined

XTX Topco Ltd

$135,326

0.57%

226,678

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	226,678	$135,326	100.0%	Defined

NORTHERN TRUST CORP

$131,424

0.56%

220,141

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	220,141	$131,424	100.0%	Defined

BANK OF AMERICA CORP /DE/

$123,858

0.52%

207,468

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	182,838	$109,154	88.1%	Defined
BOFA SECURITIES, INC.	24,630	$14,704	11.9%	Defined

JANE STREET GROUP, LLC

$106,624

0.45%

178,600

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	178,600	$106,624	100.0%	Defined

RENAISSANCE TECHNOLOGIES LLC

$98,637

0.42%

165,222

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$92,415

0.39%

154,800

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	154,800	$92,415	100.0%	Other

JPMORGAN CHASE & CO

$89,947

0.38%

150,666

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
J.P. Morgan Securities LLC	147,739	$88,200	98.1%	Defined
J.P. Morgan Securities plc	2,927	$1,747	1.9%	Defined

GROUP ONE TRADING LLC

$82,910

0.35%

138,878

Qube Research & Technologies Ltd

$75,816

0.32%

126,996

HRT FINANCIAL LP

$75,628

0.32%

126,681

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$69,550

0.29%

116,500

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	116,500	$69,550	100.0%	Other

DIMENSIONAL FUND ADVISORS LP

$54,243

0.23%

90,861

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	3,665	$2,188	4.0%	Defined
Dimensional Ireland Ltd	59,285	$35,393	65.2%	Defined
Held directly	27,911	$16,662	30.7%	Sole

HSBC HOLDINGS PLC

$39,512

0.17%

66,185

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
HSBC BANK PLC	66,185	$39,512	100.0%	Defined

Mariner, LLC

$35,018

0.15%

58,658

CITADEL ADVISORS LLC

$34,148

0.14%

57,200

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	57,200	$34,148	100.0%	Defined

HOHIMER WEALTH MANAGEMENT, LLC

$32,835

0.14%

55,000

MERCER GLOBAL ADVISORS INC /ADV

$31,958

0.14%

53,531

KINGSWOOD WEALTH ADVISORS, LLC

$31,044

0.13%

52,000

Schonfeld Strategic Advisors LLC

$30,375

0.13%

50,880

ADAR1 Capital Management, LLC

$30,258

0.13%

50,685

Alpha Wealth Funds, LLC

$29,850

0.13%

50,000

Tower Research Capital LLC (TRC)

$29,114

0.12%

48,768

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Tower Research Capital Investments LLC	30,516	$18,218	62.6%	Other
Latour Trading LLC (Latour)	18,252	$10,896	37.4%	Defined

Virtu Financial LLC

$28,011

0.12%

46,921

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Virtu Americas LLC	46,921	$28,011	100.0%	Defined

WEALTH ENHANCEMENT ADVISORY SERVICES, LLC

$26,236

0.11%

43,948

Creative Planning

$24,831

0.10%

41,593

Point72 Asset Management, L.P.

$22,254

0.09%

37,278

Hobbs Group Advisors, LLC

$21,487

0.09%

35,992

LPL Financial LLC

$21,222

0.09%

35,549

CAPTRUST FINANCIAL ADVISORS

$20,265

0.09%

33,946

Vanguard Global Advisers, LLC

$19,948

0.08%

33,414

BARCLAYS PLC

$18,524

0.08%

31,029

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	24,099	$14,387	77.7%	Defined
BARCLAYS CAPITAL INC.	6,930	$4,137	22.3%	Defined

RHUMBLINE ADVISERS

$18,155

0.08%

30,412

Cerity Partners LLC

$17,450

0.07%

29,232

Rockefeller Capital Management L.P.

$16,786

0.07%

28,118

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ROCKEFELLER FINANCIAL LLC	28,118	$16,786	100.0%	Defined

BNP PARIBAS ARBITRAGE, SA

$14,957

0.06%

25,055

Balyasny Asset Management L.P.

$13,739

0.06%

23,015

Y-Intercept (Hong Kong) Ltd

$13,356

0.06%

22,373

Abel Hall, LLC

$13,203

0.06%

22,117

Steward Partners Investment Advisory, LLC

$11,701

0.05%

19,600

MAI Capital Management

$10,057

0.04%

16,846

RAYMOND JAMES FINANCIAL INC

$10,029

0.04%

16,801

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	13,172	$7,863	78.4%	Defined
Raymond James Financial Services Advisors, Inc.	3,629	$2,166	21.6%	Defined

TRUST CO OF TOLEDO NA /OH/

$9,934

0.04%

16,640

Arax Advisory Partners

$9,884

0.04%

16,557

JANE STREET GROUP, LLC

$8,955

0.04%

15,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	15,000	$8,955	100.0%	Defined

Vanguard Personalized Indexing Management, LLC

$8,894

0.04%

14,899

DIADEMA PARTNERS LP

$7,462

0.03%

12,500

TrinityBridge Ltd

$7,343

0.03%

12,300

TORONTO DOMINION BANK

$7,283

0.03%

12,200

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
TD SECURITIES (USA) LLC	12,200	$7,283	100.0%	Defined

BRIGHTON JONES LLC

$6,840

0.03%

11,458

Gladstone Institutional Advisory LLC

$6,746

0.03%

11,300

WELLS FARGO & COMPANY/MN

$3,880

0.02%

6,500

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	5,000	$2,985	76.9%	Defined
WELLS FARGO CLEARING SERVICES, LLC	1,500	$895	23.1%	Defined

Integrated Wealth Concepts LLC

$3,579

0.02%

5,995

ROYAL BANK OF CANADA

$3,578

0.02%

5,996

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC CMA LTD.	41	$24	0.7%	Defined
RBC Capital Markets, LLC	5,166	$3,084	86.2%	Defined
RBC Dominion Securities Inc.	650	$388	10.8%	Defined
Held directly	139	$82	2.3%	Sole

Allworth Financial LP

$2,158

0.01%

3,615

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$1,200

0.01%

2,011

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	2,011	$1,200	100.0%	Defined

Ares Financial Consulting, LLC

$1,194

0.01%

2,000

NFSG Corp

$1,181

0.00%

1,979

Vision Retirement, LLC

$895

0.00%

1,500

JONES FINANCIAL COMPANIES LLLP

$895

0.00%

1,500

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Edward D. Jones & Co., L.P.	1,500	$895	100.0%	Sole

FNY Investment Advisers, LLC

$895

0.00%

1,500

MCF Advisors LLC

$835

0.00%

1,400

PNC FINANCIAL SERVICES GROUP, INC.

$761

0.00%

1,276

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PNC Bank, National Association	1,276	$761	100.0%	Defined

EverSource Wealth Advisors, LLC

$685

0.00%

1,149

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BMSS Wesson Wealth Solutions, LLC	180	$107	15.6%	Sole
Held directly	969	$578	84.4%	Sole

CITIGROUP INC

$537

0.00%

900

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	900	$537	100.0%	Defined

ROTHSCHILD INVESTMENT LLC

$528

0.00%

885

Clearstead Advisors, LLC

$514

0.00%

861

Essential Partners LLC

$486

0.00%

815

GAMMA Investing LLC

$393

0.00%

659

True Wealth Design, LLC

$277

0.00%

464

Farther Finance Advisors, LLC

$222

0.00%

372

SJS Investment Consulting Inc.

$124

0.00%

208

Larson Financial Group LLC

$89

0.00%

150

IFP Advisors, Inc

$85

0.00%

143

CALDWELL SUTTER CAPITAL, INC.

$31

0.00%

IFG Advisory, LLC

$29

0.00%

Persistent Asset Partners Ltd

$16

0.00%

NORTHWESTERN MUTUAL WEALTH MANAGEMENT CO

$14

0.00%

FNY Investment Advisers, LLC

0.00%

NISA INVESTMENT ADVISORS, LLC

0.00%

Caitong International Asset Management Co., Ltd

0.00%

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	33,829	$26,386	14.8%	Defined
TEACHERS ADVISORS, LLC joint	195,455	$152,454	85.2%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Investments Corporation	55,691	$43,438	28.8%	Defined
The Bank of New York Mellon	137,839	$107,514	71.2%	Defined
Held directly	1	$0	0.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DWS INVESTMENT MANAGEMENT AMERICAS, INC	32,075	$25,018	31.0%	Defined
DWS Investments UK Ltd	71,251	$55,575	69.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	64,480	$50,294	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	13,376	$10,433	28.9%	Defined
Invesco Trust Co joint	32,978	$25,722	71.1%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	46,126	$35,978	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Global X Management CO LLC joint	46,073	$35,936	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI TRUST CO	33,829	$26,386	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Capital	26,081	$20,343	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PGIM Quantitative Solutions LLC	14,660	$11,434	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC	11,361	$8,860	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MANULIFE FINANCIAL CORP joint	10,552	$8,230	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	10,338	$8,063	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
EATON VANCE MANAGEMENT	8,881	$6,927	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	7,920	$6,177	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
OSAIC WEALTH, INC.	5,000	$3,900	100.0%	Defined

Key facts CIK 1339970 CUSIP 002120202 13F (30d) 79 filings 78 filers Visit website Investor relations