BLFS · Biolife Solutions Inc · Holdings

$38.61 +1.31 (+3.51%) At close · Oct 5

Market Cap

$1.89B

48.91M

Volume · Oct 5 4.45M Avg daily vol (3M) 1.12M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Use institutional holdings through the 13F API or MCP → See every holder of BLFS: 13F, 13D and 13G, insiders and funds →

Recent SEC filings are still being reconciled. Holder totals and changes may be incomplete until processing catches up.

The Sep 30, 2026 filing window is still open — showing the latest known positions. Funds that have not filed yet carry their prior-quarter positions; a fund that filed without this stock counts as sold out. Each fund's Change is measured against its own previous reported quarter, shown in the Reported column. Pick a completed quarter in the Quarter selector below to put every fund on one quarter and see the stock-wide activity breakdown.

Quarter:

Holders: 227

Total Value: $1,405,939,287

Total Shares: 49,482,308

Showing 1–100 of 231 holdings by value · page 1 of 3

Institutional Ownership Trend

27 quarters

Holder Concentration

Across 216 13F filers

Top Holder

13.6%

of institutional value

Top 5

44.3%

of institutional value

Top 10

56.3%

of institutional value

HHI

546

Broadly held

Who Holds BLFS

Last 8 quarters through Sep 30, 2026 (filing window open)

Each kind of 13F filer's share of the common shares 13F filers report. Holder type comes from regulatory records, never the filer's name; style, turnover and horizon are measured from each filer's own 13F book. The quarter selector does not change this card.

Jun 30, 2026, the newest closed quarter: 216 filers, like-for-like change -0.8%; actively managed books -3.6%; index-like books +6.7%.

Holder type	Share	Filers	Change	New	Exited
Registered adviser	43.0%	107	-5.0%	36	16
Exempt adviser	0.1%	2	new	2	0
Adviser	19.9%	7	-4.6%	0	0
Bank	11.0%	14	+16.4%	0	1
Insurer	0.5%	3	+17.9%	1	0
Broker-dealer	0.0%	2	-3.1%	1	0
Benefit plan	0.4%	3	+10.5%	1	0
Non-US	0.7%	7	+5.8%	0	1
Holding company	17.2%	7	+1.4%	1	0
Unknown	7.2%	64	+5.0%	11	7

Style	Share	Filers	Change	New	Exited
Index-like	28.5%	29	+6.7%	2	0
Systematic-like	2.2%	14	+159.8%	11	5
Concentrated	10.5%	29	-18.3%	12	5
Diversified active	58.8%	144	-2.7%	28	15
Not measured	0.0%	0	—	0	0
Unknown	0.0%	0	—	0	0

Turnover	Share	Filers	Change	New	Exited
Low turnover	60.1%	127	-10.4%	17	13
Medium turnover	28.6%	44	+11.4%	14	5
High turnover	3.9%	27	+325.5%	21	6
Unknown	7.3%	18	+3.0%	1	1
Not measured	0.0%	0	—	0	0

Holding horizon	Share	Filers	Change	New	Exited
Short horizon	8.1%	27	+11.4%	10	6
Medium horizon	28.2%	22	+4.7%	4	5
Long horizon	59.0%	155	-5.7%	31	13
Unknown	4.6%	12	+14.4%	8	1
Not measured	0.0%	0	—	0	0

13F book size	Share	Filers	Change	New	Exited
Under $1B	3.5%	31	+18.0%	13	11
$1B to $10B	19.5%	57	-17.4%	13	7
$10B to $100B	10.1%	65	+6.0%	17	5
$100B or more	67.0%	63	+3.3%	10	2
Not measured	0.0%	0	—	0	0

Client base	Share	Filers	Change	New	Exited
Individuals and high net worth	3.2%	42	-32.0%	12	10
Pooled investment vehicles	13.8%	18	-2.0%	13	5
Registered funds and BDCs	19.6%	17	+3.3%	4	0
Pension and profit sharing plans	0.0%	1	+25.5%	0	0
Insurance companies	0.3%	4	+9.5%	1	0
Governments and sovereign funds	0.6%	3	-28.3%	1	0
Corporations	0.5%	1	-11.5%	0	0
Other investment advisers	0.1%	4	+5.6%	0	0
Other	0.5%	2	+944.0%	1	0
Diversified	4.4%	15	-23.1%	4	1
Unknown	57.0%	109	+2.5%	17	9

Private funds	Share	Filers	Change	New	Exited
Hedge funds	8.2%	31	+18.7%	19	6
Private equity funds	0.6%	7	+467.9%	3	1
Venture capital funds	0.0%	1	+36.7%	0	0
Real estate funds	0.0%	4	+6.1%	1	1
Securitized asset funds	0.3%	2	+5.0%	0	0
Other private funds	5.3%	11	-9.3%	3	1
Mixed private funds	9.5%	2	-21.6%	0	0
Private funds, type not stated	2.2%	2	-9.3%	1	0
No private funds	17.0%	49	-3.3%	11	7
Unknown	56.9%	107	+2.4%	15	9

Location	Share	Filers	Change	New	Exited
United States	94.4%	184	-2.3%	45	21
Outside the US	5.6%	32	+31.0%	8	4

The Sep 30, 2026 filing window is still open, so its bar carries the Jun 30, 2026 positions of filers that have not filed yet.

How the mix is measured

Shares are 13F common shares restated to today's split basis; options and principal amounts are left out. The change compares only filers with a 13F in both quarters and keeps each filer in its bucket of the later quarter on both sides; a filer that continues under a new CIK is compared through its successor. Holder type comes from each filer's current regulatory profile; client base and private funds come from the Form ADV roster current at each quarter end (rosters from December 2019, so an earlier quarter reads Unknown). Style, turnover, horizon and book size are measured from each filer's 13F book in the newest measured quarter, and Not measured means none was. Actively managed books are those measured systematic-like, concentrated or diversified active.

Excluded 2 holders with no observed 2026-03-31 13F and 1 holder with no observed 2026-06-30 13F. Compared across CIK changes: JANUS HENDERSON GROUP PLC continues as Jupiter Topco LLC.

Conviction and Crowding

216 13F filers, quarter ended Jun 30, 2026

Measured across every 13F filer's common-share book once the quarter's filing window closed. Crowding counts filers with high turnover or that advise private funds on Form ADV.

Conviction holders

1.85% of filers hold it in their top 10

Mar 31, 2026: 1

Holders above 5%

median weight 0.002%

Mar 31, 2026: 1

Crowding

—

holders exceed shares outstanding: 13F filers report more shares than the cover-page count, which predates an issuance, is misstated, or misses shares lent and reported twice

Private-fund advisers

—

of shares outstanding

Institutional ownership

—

holders exceed shares outstanding: 13F filers report more shares than the cover-page count, which predates an issuance, is misstated, or misses shares lent and reported twice

Top 10 holders

56.5%

of 13F shares, Herfindahl 0.055

Active holders

-3.6%

like-for-like share change of active filers

5% stakes on record

0 13D, 4 13G (2 institutional, 2 passive, 0 exempt)

How long holders have held it value-weighted 15.2 quarters

Under 1 year 13.9% 1 to 3 years 34.4% 3 years or more 51.7% Since coverage start 0.000%

Managers in the top fifth by current three-year FundScore alpha were net buyers of $0 (0 buying, 0 selling).

Estimated Holder Cost Basis

Quarter ended Jun 30, 2026, on that quarter's share basis

Estimated, never filed: each 13F buy is costed at its quarter's average traded price and each sale keeps the average cost, so this is what holders paid on average if they traded evenly through each quarter.

Estimated avg cost

$24.64

estimated per share

Mar 31, 2026: $24.96

Quarter-end price

$28.23

implied by 13F values

Estimated gain or loss

+14.6%

price against the estimated cost

Estimated at a loss

9.86%

of costed 13F value

Estimated cost covers 48.0% of the 13F shares; positions already held when a filer's 13F history starts, or bought in a quarter without a trusted average price, have no estimate.

Shared Holders

Companies BLFS's active holders also own, quarter ended Jun 30, 2026

#	Company	Shared holders	Similarity
1	IRMD Iradimed Corp	5	0.243
2	AGYS Agilysys Inc	7	0.237
3	NCNO nCino, Inc.	5	0.229
4	WLDN Willdan Group, Inc.	5	0.224
5	MAMA Mama's Creations, Inc.	5	0.196
6	CVLG Covenant Logistics Group, Inc.	3	0.190
7	MLAB Mesa Laboratories Inc	3	0.190
8	VCEL Vericel Corp	5	0.183
9	NVEC Nve Corp	3	0.181
10	CWST Casella Waste Systems Inc	7	0.181

Shareholder Targets

13F filers that own its peers but not BLFS, quarter ended Jun 30, 2026

Found through its holder-overlap peers: IRMD, AGYS, NCNO, WLDN, MAMA, CVLG, MLAB, VCEL, NVEC, CWST, BWMN, ELVA, LMAT, VERX, LGND, DV, PKE, COCO, KIDS, AMPL. 948 filers in all.

#	Filer	In peers	Of book	Largest peers
1	WASATCH ADVISORS LP Registered adviser, Diversified active	$666,013,018 7 held	4.36%	CWST, COCO, AGYS
2	CONGRESS ASSET MANAGEMENT CO Registered adviser, Diversified active	$510,652,184 4 held	3.15%	LGND, VCEL, LMAT
3	WELLINGTON MANAGEMENT GROUP LLP Holding company, Diversified active	$490,709,709 11 held	0.080%	CWST, COCO, AMPL
4	FRANKLIN RESOURCES INC Adviser, Diversified active	$414,595,660 16 held	0.090%	LGND, AGYS, CWST
5	LOOMIS SAYLES & CO L P Registered adviser, Diversified active	$295,357,930 5 held	0.370%	LGND, COCO, CWST
6	Capital Research Global Investors Adviser, Diversified active	$239,979,130 2 held	0.030%	CWST, AGYS
7	AMERICAN CENTURY COMPANIES INC Unknown, Diversified active	$239,416,115 16 held	0.110%	AGYS, CWST, CVLG
8	NEUBERGER BERMAN GROUP LLC Holding company, Diversified active	$219,468,794 7 held	0.150%	CWST, LGND, LMAT
9	GW&K Investment Management, LLC Registered adviser, Diversified active	$204,092,190 4 held	1.60%	LGND, VCEL, COCO
10	Capital International Investors Adviser, Diversified active	$199,215,071 1 held	0.040%	CWST

Fund Ownership of BLFS by Fund Type

Registered funds' Form N-PORT reports for the quarter ended Jun 30, 2026

Index funds and index ETFs hold 20.54% of shares outstanding.

Fund type	Of shares outstanding	Change	Shares	Funds
Index funds	4.13%	+0.42 pp	2,020,988	75
Index ETFs	16.41%	+0.63 pp	8,024,285	50
Active ETFs	0.80%	+0.15 pp	390,012	12
Active funds	27.41%	+3.21 pp	13,404,536	109
Unclassified	0.07%	+0.02 pp	33,377	6
All registered funds	48.81%	+4.44 pp	23,873,198	252

How fund types are measured

Each fund series counts once, with its latest Form N-PORT report dated inside the quarter; only long share positions count, restated to today's split basis. The fund type is the series' latest Form N-CEN answer on or before the quarter end (the index-fund box and the exchange-traded flag), never its name. Unclassified funds have no usable N-CEN; unit investment trusts are among them, so index ownership is a floor. Funds sit inside their advisers' 13F reports, so these shares overlap the 13F figures and are never added to them.

Holder type: Registered adviser 117 Exempt adviser 2 Adviser 8 Bank 14 Insurer 3 Broker-dealer 2 Benefit plan 3 Non-US 7 Holding company 7 Unknown 64

Showing 100 of 231

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

T. Rowe Price Investment Management, Inc.

$190,652,052

13.56%

6,751,135

BlackRock, Inc.

$180,045,384

12.81%

6,375,545

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	67,514	$1,906,595	1.1%	Sole
BlackRock Investment Management (Australia) Ltd	93	$2,626	0.0%	Sole
BlackRock Asset Management Canada Ltd	196	$5,535	0.0%	Sole
BlackRock Investment Management, LLC	141,780	$4,003,867	2.2%	Sole
BLACKROCK ADVISORS LLC	94,783	$2,676,671	1.5%	Sole
BlackRock Fund Advisors	4,491,733	$126,846,539	70.5%	Sole
BlackRock Institutional Trust Company, N.A.	1,218,673	$34,415,325	19.1%	Sole
BlackRock Fund Managers Ltd.	6,973	$196,917	0.1%	Sole
BlackRock Investment Management (UK) Ltd.	19,033	$537,491	0.3%	Sole
BlackRock (Netherlands) B.V.	14,639	$413,405	0.2%	Sole
BlackRock Asset Management Ireland Ltd	214,475	$6,056,774	3.4%	Sole
BlackRock (Luxembourg) S.A.	43,305	$1,222,933	0.7%	Sole
BlackRock Asset Management Schweiz AG	8,112	$229,082	0.1%	Sole
Aperio Group, LLC	54,236	$1,531,624	0.9%	Sole

Casdin Capital, LLC

$131,457,200

9.35%

4,655,000

Invesco Ltd.

$63,269,543

4.50%

2,240,423

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	2,161,715	$61,046,830	96.5%	Defined
Invesco Trust Co joint	17,016	$480,531	0.8%	Defined
Invesco Capital Management LLC	61,692	$1,742,182	2.8%	Defined

VANGUARD CAPITAL MANAGEMENT LLC

$55,465,732

3.95%

1,964,084

STATE STREET CORP

$46,763,378

3.33%

1,655,927

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	1,655,927	$46,763,378	100.0%	Defined

GEODE CAPITAL MANAGEMENT, LLC

$30,693,067

2.18%

1,086,865

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	151,781	$4,286,295	14.0%	Defined
Held directly	935,084	$26,406,772	86.0%	Defined

MASSACHUSETTS FINANCIAL SERVICES CO /MA/

$30,478,584

2.17%

1,079,270

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MFS Heritage Trust CO	3,568	$100,760	0.3%	Defined
Held directly	1,075,702	$30,377,824	99.7%	Sole

FRED ALGER MANAGEMENT, LLC

$30,430,774

2.16%

1,077,577

DIMENSIONAL FUND ADVISORS LP

$29,546,154

2.10%

1,046,252

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	5,772	$163,001	0.6%	Defined
DFA Australia Ltd.	12,657	$357,433	1.2%	Defined
Dimensional Fund Advisors Pte. Ltd.	518	$14,628	0.0%	Defined
Dimensional Ireland Ltd	32,112	$906,842	3.1%	Defined
Held directly	995,193	$28,104,250	95.1%	Sole

FMR LLC

$28,045,312

1.99%

993,106

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	984,408	$27,799,681	99.1%	Defined
STRATEGIC ADVISERS LLC	698	$19,711	0.1%	Defined
FIAM LLC	8,000	$225,920	0.8%	Defined

BANK OF AMERICA CORP /DE/

$26,832,489

1.91%

950,159

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	268,842	$7,592,098	28.3%	Defined
MERRILL LYNCH, PIERCE, FENNER & SMITH INC.	11,127	$314,226	1.2%	Defined
BOFA SECURITIES, INC.	87,013	$2,457,247	9.2%	Defined
MANAGED ACCOUNT ADVISORS LLC joint	583,177	$16,468,918	61.4%	Defined

VANGUARD PORTFOLIO MANAGEMENT LLC

$25,789,502

1.83%

913,226

ROYAL BANK OF CANADA

$25,159,239

1.79%

890,908

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC CMA LTD.	1,294	$36,542	0.1%	Defined
RBC Capital Markets, LLC	30,748	$868,323	3.5%	Defined
RBC Global Asset Management (U.S.) Inc.	706,168	$19,942,184	79.3%	Defined
RBC Rochdale, LLC	26	$734	0.0%	Defined
Alaska Permanent Fund Corp joint	151,890	$4,289,373	17.0%	Defined
Held directly	782	$22,083	0.1%	Sole

UBS Group AG

$23,499,717

1.67%

832,143

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	13,217	$373,248	1.6%	Defined
UBS AG	495,576	$13,995,066	59.6%	Defined
UBS Switzerland AG	321,035	$9,066,028	38.6%	Defined
UBS Securities LLC	2,315	$65,375	0.3%	Defined

Stephens Investment Management Group LLC

$20,012,727

1.42%

708,666

MILLENNIUM MANAGEMENT LLC

$17,668,581

1.26%

625,658

ROYCE & ASSOCIATES LP

$17,618,230

1.25%

623,875

GENEVA CAPITAL MANAGEMENT LLC

$16,439,661

1.17%

582,141

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nationwide Fund Advisors	175,828	$4,965,382	30.2%	Other
Held directly	406,313	$11,474,279	69.8%	Sole

CRAMER ROSENTHAL MCGLYNN LLC

$16,260,366

1.16%

575,792

NEXT CENTURY GROWTH INVESTORS LLC

$15,781,330

1.12%

558,829

GAGNON SECURITIES LLC

$15,133,985

1.08%

535,906

GOLDMAN SACHS GROUP INC

$15,011,875

1.07%

531,582

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	136,192	$3,846,062	25.6%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	387,609	$10,946,078	72.9%	Defined
Goldman Sachs Trust Company, N.A. joint	7,781	$219,735	1.5%	Other

Granahan Investment Management, LLC

$14,725,183

1.05%

521,430

NORTHERN TRUST CORP

$13,548,900

0.96%

479,777

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	430,155	$12,147,576	89.7%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	29,748	$840,083	6.2%	Defined
MASON STREET ADVISORS, LLC joint	13,335	$376,580	2.8%	Other
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	6,539	$184,661	1.4%	Other

MORGAN STANLEY

$13,484,709

0.96%

477,504

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	111	$3,134	0.0%	Defined
MORGAN STANLEY & CO. LLC	12,586	$355,428	2.6%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	238,908	$6,746,761	50.0%	Defined
MORGAN STANLEY INVESTMENT MANAGEMENT INC	794	$22,422	0.2%	Defined
MORGAN STANLEY SMITH BARNEY LLC	12,622	$356,445	2.6%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	206,957	$5,844,465	43.3%	Defined
EATON VANCE MANAGEMENT	5,526	$156,054	1.2%	Defined

Rockefeller Capital Management L.P.

$12,567,364

0.89%

445,020

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ROCKEFELLER & CO. LLC	382,992	$10,815,694	86.1%	Defined
ROCKEFELLER TRUST COMPANY, N.A.	506	$14,289	0.1%	Defined
Rockefeller Trust Co (Delaware)	15,796	$446,079	3.5%	Defined
ROCKEFELLER FINANCIAL LLC	45,726	$1,291,302	10.3%	Defined

RAYMOND JAMES FINANCIAL INC

$12,361,549

0.88%

437,732

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	2,404	$67,888	0.5%	Defined
Raymond James Financial Services Advisors, Inc.	6,361	$179,634	1.5%	Defined
ClariVest Asset Management LLC joint	45,894	$1,296,046	10.5%	Defined
EAGLE ASSET MANAGEMENT INC	383,073	$10,817,981	87.5%	Defined

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$11,377,783

0.81%

402,896

PALISADE CAPITAL MANAGEMENT, LP

$9,961,292

0.71%

352,737

PEREGRINE CAPITAL MANAGEMENT LLC

$9,188,787

0.65%

325,382

Roubaix Capital, LLC

$8,743,612

0.62%

309,618

NEW YORK STATE COMMON RETIREMENT FUND

$8,537,460

0.61%

302,318

Russell Investments Group, Ltd.

$7,751,030

0.55%

274,470

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC	150,773	$4,257,828	54.9%	Defined
Russell Investments Capital, LLC	66,297	$1,872,227	24.2%	Defined
Russell Investments France SAS	22,119	$624,640	8.1%	Defined
Russell Investments Trust Company	35,281	$996,335	12.9%	Defined

Gagnon Advisors, LLC

$7,743,888

0.55%

274,217

VANGUARD FIDUCIARY TRUST CO

$7,690,401

0.55%

272,323

ABC ARBITRAGE SA

$7,347,312

0.52%

189,120

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ABC ARBITRAGE ASSET MANAGEMENT joint	189,120	$7,347,312	100.0%	Other

Thrivent Financial for Lutherans

$6,771,245

0.48%

239,775

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Thrivent Asset Management, LLC	89,629	$2,531,122	37.4%	Defined
Held directly	150,146	$4,240,123	62.6%	Sole

Heights Point Management, LP

$6,745,124

0.48%

238,850

Ranger Investment Management, L.P.

$6,735,833

0.48%

238,521

MUTUAL OF AMERICA CAPITAL MANAGEMENT LLC

$6,653,528

0.47%

171,262

JPMORGAN CHASE & CO

$6,501,608

0.46%

230,227

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	17,694	$499,678	7.7%	Defined
J.P. Morgan Investment Management Inc.	2,043	$57,694	0.9%	Defined
J.P. Morgan Securities LLC	1,230	$34,735	0.5%	Defined
J.P. Morgan Securities plc	66	$1,863	0.0%	Defined
JPMorgan Asset Management (UK) Ltd	209,194	$5,907,638	90.9%	Defined

Bank of New York Mellon Corp

$6,470,034

0.46%

229,109

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Investments Corporation	104,616	$2,954,355	45.7%	Defined
BNY Mellon Investment Adviser, Inc.	32,330	$912,999	14.1%	Defined
The Bank of New York Mellon	81,035	$2,288,428	35.4%	Defined
BNY Mellon, NA joint	9,429	$266,273	4.1%	Defined
BNY Mellon Advisors, Inc.	1,696	$47,895	0.7%	Defined
Held directly	3	$84	0.0%	Defined

TD Asset Management Inc

$5,434,505

0.39%

192,440

PRINCIPAL FINANCIAL GROUP INC

$5,415,753

0.39%

191,776

Uniplan Investment Counsel, Inc.

$5,318,439

0.38%

188,330

JANE STREET GROUP, LLC

$5,169,642

0.37%

183,061

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	94,271	$2,662,213	51.5%	Defined
Jane Street Global Trading, LLC	88,790	$2,507,429	48.5%	Defined

Summit Creek Advisors LLC

$5,098,562

0.36%

180,544

ENVESTNET ASSET MANAGEMENT INC

$4,559,545

0.32%

161,457

CITADEL ADVISORS LLC

$4,460,338

0.32%

157,944

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	64,971	$1,834,781	41.1%	Defined
Held directly	92,973	$2,625,557	58.9%	Defined

DIAMOND HILL CAPITAL MANAGEMENT INC

$4,345,768

0.31%

153,887

Man Group plc

$4,109,004

0.29%

145,503

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Numeric Investors LLC	145,503	$4,109,004	100.0%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$4,029,283

0.29%

142,680

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	142,680	$4,029,283	100.0%	Other

Public Sector Pension Investment Board

$3,840,414

0.27%

135,992

Squarepoint Ops LLC

$3,754,282

0.27%

132,942

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Squarepoint Capital LLP	132,942	$3,754,282	100.0%	Defined

BNP PARIBAS ARBITRAGE, SA

$3,752,841

0.27%

132,891

SEI INVESTMENTS CO

$3,563,040

0.25%

126,170

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI INVESTMENTS MANAGEMENT CORP	9,760	$275,622	7.7%	Sole
SEI TRUST CO	116,410	$3,287,418	92.3%	Sole

BARINGS LLC

$3,547,626

0.25%

91,316

Conestoga Capital Advisors, LLC

$3,435,452

0.24%

121,652

STATE OF WISCONSIN INVESTMENT BOARD

$3,405,179

0.24%

120,580

WELLS FARGO & COMPANY/MN

$3,246,977

0.23%

114,978

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	36,369	$1,027,060	31.6%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	29,231	$825,483	25.4%	Defined
WELLS FARGO CLEARING SERVICES, LLC	49,378	$1,394,434	42.9%	Defined

NOMURA ASSET MANAGEMENT INTERNATIONAL INC.

$3,034,981

0.22%

107,471

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NOMURA HOLDINGS INC joint	107,471	$3,034,981	100.0%	Defined

Trium Capital LLP

$2,961,613

0.21%

76,232

RHUMBLINE ADVISERS

$2,936,254

0.21%

103,975

Legal & General Group Plc

$2,615,898

0.19%

92,631

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	60,933	$1,720,747	65.8%	Defined
Legal & General Investment Management Ltd joint	31,698	$895,151	34.2%	Defined

Versor Investments LP

$2,373,657

0.17%

61,098

Swiss National Bank

$2,326,976

0.17%

82,400

Nuveen, LLC

$2,165,866

0.15%

76,695

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	602	$17,000	0.8%	Defined
TEACHERS ADVISORS, LLC joint	76,093	$2,148,866	99.2%	Defined

NEW YORK STATE TEACHERS RETIREMENT SYSTEM

$2,162,429

0.15%

55,661

ExodusPoint Capital Management, LP

$2,149,628

0.15%

76,120

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$2,140,307

0.15%

75,790

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	2,532	$71,503	3.3%	Defined
UBS Asset Management Switzerland AG	64,048	$1,808,715	84.5%	Sole
Held directly	9,210	$260,089	12.2%	Defined

Hodges Capital Management Inc.

$2,061,520

0.15%

73,000

Verition Fund Management LLC

$1,957,709

0.14%

69,324

ZACKS INVESTMENT MANAGEMENT

$1,906,369

0.14%

67,506

AMERIPRISE FINANCIAL INC

$1,886,742

0.13%

66,811

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	66,811	$1,886,742	100.0%	Defined

XTX Topco Ltd

$1,801,231

0.13%

63,783

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	63,783	$1,801,231	100.0%	Defined

Numerai GP LLC

$1,758,109

0.13%

62,256

JACOBS LEVY EQUITY MANAGEMENT, INC

$1,746,700

0.12%

61,852

PRUDENTIAL FINANCIAL INC

$1,716,766

0.12%

60,792

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PGIM Quantitative Solutions LLC	60,792	$1,716,766	100.0%	Defined

Cerity Partners LLC

$1,623,969

0.12%

57,506

Vestmark Advisory Solutions, Inc.

$1,565,625

0.11%

55,440

CenterBook Partners LP

$1,518,408

0.11%

53,768

TIG Advisors, LLC

$1,513,974

0.11%

53,611

Penbrook Management LLC

$1,433,370

0.10%

36,895

ALLIANCEBERNSTEIN L.P.

$1,430,356

0.10%

50,650

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	50,650	$1,430,356	100.0%	Defined

MILLENNIUM MANAGEMENT LLC

$1,412,000

0.10%

50,000

DEUTSCHE BANK AG\

$1,292,486

0.09%

45,768

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Deutsche Bank Trust Co Americas	23	$649	0.1%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC	166	$4,687	0.4%	Defined
DWS Investments UK Ltd	45,579	$1,287,150	99.6%	Defined

PRICE T ROWE ASSOCIATES INC /MD/

$1,290,935

0.09%

45,713

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	45,713	$1,290,935	100.0%	Defined

CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM

$1,215,251

0.09%

43,033

KORNITZER CAPITAL MANAGEMENT INC /KS

$1,167,695

0.08%

41,349

CIBC Bancorp USA Inc.

$1,132,140

0.08%

40,090

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CIBC Private Wealth Advisors, Inc.	37,727	$1,065,410	94.1%	Defined
CIBC National Trust Co	7	$197	0.0%	Defined
CIBC WORLD MARKETS CORP	2,356	$66,533	5.9%	Defined

HSBC HOLDINGS PLC

$1,108,730

0.08%

39,261

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
HSBC BANK PLC	39,261	$1,108,730	100.0%	Defined

GOLDMAN SACHS GROUP INC

$1,059,000

0.08%

37,500

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	37,500	$1,059,000	100.0%	Defined

VANGUARD ASSET MANAGEMENT, Ltd

$1,057,446

0.08%

37,445

FOX RUN MANAGEMENT, L.L.C.

$1,036,662

0.07%

36,709

CITADEL ADVISORS LLC

$1,010,992

0.07%

35,800

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	35,800	$1,010,992	100.0%	Defined

Point72 Asset Management, L.P.

$988,936

0.07%

35,019

ISTHMUS PARTNERS, LLC

$976,067

0.07%

25,124

D.A. DAVIDSON & CO.

$958,804

0.07%

33,952

CANADA LIFE ASSURANCE Co

$903,934

0.06%

32,009

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Key facts CIK 834365 CUSIP 09062W204 13F (30d) 38 filings 37 filers Visit website Investor relations