VCEL · Vericel Corp · Holdings

$40.58 +0.21 (+0.52%) At close · Oct 20

S&P 600 member Russell 2000 member Rebalance outlook locked

Market Cap

$2.09B

51.21M

Volume · Oct 20 475.27K Avg daily vol (3M) 509.5K

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Use institutional holdings through the 13F API or MCP → See every holder of VCEL: 13F, 13D and 13G, insiders and funds →

Institutional holdings are being reconciled with SEC filings. Holder totals and changes may be incomplete until reconciliation finishes.

The Sep 30, 2026 filing window is still open — showing the latest known positions. Funds that have not filed yet carry their prior-quarter positions; a fund that filed without this stock counts as sold out. Each fund's Change is measured against its own previous reported quarter, shown in the Reported column. Pick a completed quarter in the Quarter selector below to put every fund on one quarter and see the stock-wide activity breakdown.

Quarter:

Holders: 322

Total Value: $2,445,972,405

Total Shares: 55,244,288

Showing 1–100 of 326 holdings by value · page 1 of 4

Institutional Ownership Trend

27 quarters

Superinvestors Holding VCEL

All superinvestors →

Joel Greenblatt $635,895

Holder Concentration

Across 321 13F filers

Top Holder

14.6%

of institutional value

Top 5

33.5%

of institutional value

Top 10

47.9%

of institutional value

HHI

395

Broadly held

Who Holds VCEL

Last 8 quarters through Sep 30, 2026 (filing window open)

Each kind of 13F filer's share of the common shares 13F filers report. Holder type comes from regulatory records, never the filer's name; style, turnover and horizon are measured from each filer's own 13F book. The quarter selector does not change this card.

Jun 30, 2026, the newest closed quarter: 320 filers, like-for-like change +0.3%; actively managed books -1.9%; index-like books +4.5%.

Holder type	Share	Filers	Change	New	Exited
Registered adviser	44.8%	162	+5.4%	34	14
Exempt adviser	0.2%	4	-56.4%	2	0
Adviser	7.9%	11	-5.6%	1	0
Bank	12.9%	19	+1.4%	2	0
Insurer	0.6%	3	+6.5%	1	0
Broker-dealer	0.1%	1	-39.3%	0	0
Benefit plan	0.3%	5	+22.9%	0	0
Non-US	3.7%	9	-2.5%	1	1
Holding company	18.3%	10	-6.7%	1	0
Unknown	11.2%	96	-1.3%	20	7

Style	Share	Filers	Change	New	Exited
Index-like	34.9%	38	+4.5%	3	1
Systematic-like	4.5%	25	+29.6%	11	4
Concentrated	0.3%	31	-35.8%	11	2
Diversified active	60.2%	226	-3.4%	37	15
Not measured	0.0%	0	—	0	0
Unknown	0.0%	0	—	0	0

Turnover	Share	Filers	Change	New	Exited
Low turnover	59.5%	179	+0.4%	23	9
Medium turnover	22.6%	76	+7.0%	15	5
High turnover	9.1%	38	-18.6%	18	7
Unknown	8.7%	27	+7.3%	6	1
Not measured	0.0%	0	—	0	0

Holding horizon	Share	Filers	Change	New	Exited
Short horizon	10.6%	41	-0.9%	15	2
Medium horizon	20.0%	40	-7.3%	9	3
Long horizon	64.6%	219	+0.1%	31	14
Unknown	4.8%	20	+64.2%	7	3
Not measured	0.0%	0	—	0	0

13F book size	Share	Filers	Change	New	Exited
Under $1B	2.4%	50	-17.5%	18	6
$1B to $10B	18.0%	103	-17.2%	28	10
$10B to $100B	29.0%	102	+9.8%	14	4
$100B or more	50.6%	65	+3.9%	2	2
Not measured	0.0%	0	—	0	0

Client base	Share	Filers	Change	New	Exited
Individuals and high net worth	7.1%	73	+3.5%	19	4
Pooled investment vehicles	4.5%	25	-16.9%	9	6
Registered funds and BDCs	14.6%	26	+1.3%	4	2
Pension and profit sharing plans	0.1%	2	+32.5%	0	0
Insurance companies	0.1%	4	-8.5%	0	0
Governments and sovereign funds	1.0%	6	-30.1%	1	0
Other investment advisers	0.1%	6	+19.8%	1	0
Other	0.3%	1	-5.8%	0	0
Diversified	17.0%	18	+23.5%	0	2
Unknown	55.2%	159	-3.6%	28	8

Private funds	Share	Filers	Change	New	Exited
Hedge funds	11.2%	37	-10.3%	10	8
Private equity funds	0.4%	7	+0.1%	1	0
Venture capital funds	0.0%	3	+1.9%	0	0
Real estate funds	0.0%	4	+38.5%	0	0
Securitized asset funds	0.2%	1	-16.3%	0	0
Other private funds	10.6%	17	+9.3%	1	1
Mixed private funds	0.2%	2	+96.5%	0	0
Private funds, type not stated	0.1%	5	+37.1%	2	0
No private funds	22.2%	89	+11.8%	22	5
Unknown	55.0%	155	-3.2%	26	8

Location	Share	Filers	Change	New	Exited
United States	91.3%	271	+0.5%	51	20
Outside the US	8.7%	49	-1.9%	11	2

The Sep 30, 2026 filing window is still open, so its bar carries the Jun 30, 2026 positions of filers that have not filed yet.

How the mix is measured

Shares are 13F common shares restated to today's split basis; options and principal amounts are left out. The change compares only filers with a 13F in both quarters and keeps each filer in its bucket of the later quarter on both sides; a filer that continues under a new CIK is compared through its successor. Holder type comes from each filer's current regulatory profile; client base and private funds come from the Form ADV roster current at each quarter end (rosters from December 2019, so an earlier quarter reads Unknown). Style, turnover, horizon and book size are measured from each filer's 13F book in the newest measured quarter, and Not measured means none was. Actively managed books are those measured systematic-like, concentrated or diversified active.

Excluded 2 holders with no observed 2026-03-31 13F and 2 holders with no observed 2026-06-30 13F. Compared across CIK changes: JANUS HENDERSON GROUP PLC continues as Jupiter Topco LLC.

Conviction and Crowding

320 13F filers, quarter ended Jun 30, 2026

Measured across every 13F filer's common-share book once the quarter's filing window closed. Crowding counts filers with high turnover or that advise private funds on Form ADV.

Conviction holders

0.625% of filers hold it in their top 10

Mar 31, 2026: 4

Holders above 5%

median weight 0.004%

Mar 31, 2026: 1

Crowding

—

holders exceed shares outstanding: 13F filers report more shares than the cover-page count, which predates an issuance, is misstated, or misses shares lent and reported twice

Private-fund advisers

—

of shares outstanding

Institutional ownership

—

holders exceed shares outstanding: 13F filers report more shares than the cover-page count, which predates an issuance, is misstated, or misses shares lent and reported twice

Top 10 holders

47.9%

of 13F shares, Herfindahl 0.040

Active holders

-1.9%

like-for-like share change of active filers

5% stakes on record

0 13D, 6 13G (6 institutional, 0 passive, 0 exempt)

How long holders have held it value-weighted 14.7 quarters

Under 1 year 16.1% 1 to 3 years 29.6% 3 years or more 54.3% Since coverage start 0.000%

Managers in the top fifth by current three-year FundScore alpha were net buyers of $0 (0 buying, 0 selling).

Estimated Holder Cost Basis

Quarter ended Jun 30, 2026, on that quarter's share basis

Estimated, never filed: each 13F buy is costed at its quarter's average traded price and each sale keeps the average cost, so this is what holders paid on average if they traded evenly through each quarter.

Estimated avg cost

$38.27

estimated per share

Mar 31, 2026: $38.53

Quarter-end price

$44.49

implied by 13F values

Estimated gain or loss

+16.2%

price against the estimated cost

Estimated at a loss

7.51%

of costed 13F value

Estimated cost covers 44.6% of the 13F shares; positions already held when a filer's 13F history starts, or bought in a quarter without a trusted average price, have no estimate.

Shared Holders

Companies VCEL's active holders also own, quarter ended Jun 30, 2026

#	Company	Shared holders	Similarity
1	AGYS Agilysys Inc	9	0.278
2	VCYT Veracyte, Inc.	8	0.237
3	LMAT Lemaitre Vascular Inc	7	0.219
4	CWST Casella Waste Systems Inc	9	0.216
5	PNTG Pennant Group, Inc.	6	0.203
6	CVLT Commvault Systems Inc	6	0.200
7	ROAD Construction Partners, Inc.	7	0.197
8	ALHC Alignment Healthcare, Inc.	5	0.186
9	BLFS Biolife Solutions Inc	5	0.183
10	RGEN Repligen Corp	8	0.181

Shareholder Targets

13F filers that own its peers but not VCEL, quarter ended Jun 30, 2026

Found through its holder-overlap peers: AGYS, VCYT, LMAT, CWST, PNTG, CVLT, ROAD, ALHC, BLFS, RGEN, AXGN, KRYS, SUPN, NSSC, TRNS, WLDN, DORM, EXLS, KNSA, CVSA. 1,069 filers in all.

#	Filer	In peers	Of book	Largest peers
1	Avoro Capital Advisors LLC Registered adviser, Concentrated	$1,073,713,002 1 held	9.64%	KRYS
2	T. Rowe Price Investment Management, Inc. Adviser, Diversified active	$964,081,574 5 held	0.630%	ALHC, CWST, RGEN
3	WASATCH ADVISORS LP Registered adviser, Diversified active	$938,051,672 8 held	6.15%	CWST, RGEN, ROAD
4	Capital World Investors Adviser, Diversified active	$632,239,151 4 held	0.070%	KRYS, CVSA, ROAD
5	Hood River Capital Management LLC Registered adviser, Diversified active	$498,903,163 4 held	3.63%	KRYS, ALHC, TRNS
6	NORGES BANK Non-US, Index-like	$408,880,305 10 held	0.040%	KRYS, RGEN, EXLS
7	Artisan Partners Limited Partnership Registered adviser, Diversified active	$397,661,273 2 held	0.650%	VCYT, AGYS
8	UNITED CAPITAL FINANCIAL ADVISORS, LLC Registered adviser, Diversified active	$380,756,286 1 held	2.82%	RGEN
9	MILLENNIUM MANAGEMENT LLC Registered adviser, Systematic-like	$374,321,424 16 held	0.290%	SUPN, EXLS, ROAD
10	Capital Research Global Investors Adviser, Diversified active	$344,727,153 3 held	0.050%	CWST, ALHC, AGYS

Fund Ownership of VCEL by Fund Type

Registered funds' Form N-PORT reports for the quarter ended Jun 30, 2026

Index funds and index ETFs hold 27.25% of shares outstanding.

Fund type	Of shares outstanding	Change	Shares	Funds
Index funds	4.80%	+0.56 pp	2,459,021	75
Index ETFs	22.45%	+0.74 pp	11,494,166	53
Active ETFs	0.61%	+0.07 pp	313,427	11
Active funds	18.16%	-1.55 pp	9,298,690	124
Unclassified	0.33%	+0.32 pp	168,844	6
All registered funds	46.35%	+0.14 pp	23,734,148	269

How fund types are measured

Each fund series counts once, with its latest Form N-PORT report dated inside the quarter; only long share positions count, restated to today's split basis. The fund type is the series' latest Form N-CEN answer on or before the quarter end (the index-fund box and the exchange-traded flag), never its name. Unclassified funds have no usable N-CEN; unit investment trusts are among them, so index ownership is a floor. Funds sit inside their advisers' 13F reports, so these shares overlap the 13F figures and are never added to them.

Holder type: Registered adviser 162 Exempt adviser 4 Adviser 11 Bank 19 Insurer 3 Broker-dealer 1 Benefit plan 5 Non-US 9 Holding company 10 Unknown 98

Showing 100 of 326

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

BlackRock, Inc.

$360,023,128

14.72%

8,092,226

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	58,651	$2,609,382	0.7%	Sole
BlackRock Investment Management (Australia) Ltd	154	$6,851	0.0%	Sole
BlackRock Asset Management Canada Ltd	298	$13,258	0.0%	Sole
BlackRock Investment Management, LLC	157,036	$6,986,531	1.9%	Sole
BLACKROCK ADVISORS LLC	120,379	$5,355,661	1.5%	Sole
BlackRock Fund Advisors	5,860,154	$260,718,251	72.4%	Sole
BlackRock Institutional Trust Company, N.A.	1,594,990	$70,961,105	19.7%	Sole
BlackRock Fund Managers Ltd.	9,876	$439,383	0.1%	Sole
BlackRock Investment Management (UK) Ltd.	12,537	$557,771	0.2%	Sole
BlackRock Asset Management Ireland Ltd	210,087	$9,346,770	2.6%	Sole
BlackRock Life Ltd	293	$13,035	0.0%	Sole
BlackRock Asset Management Schweiz AG	9,436	$419,807	0.1%	Sole
Aperio Group, LLC	58,304	$2,593,944	0.7%	Sole
SpiderRock Advisors, LLC	31	$1,379	0.0%	Sole

STATE STREET CORP

$146,548,947

5.99%

3,293,975

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	3,293,975	$146,548,947	100.0%	Defined

CONGRESS ASSET MANAGEMENT CO

$102,439,929

4.19%

2,552,066

VANGUARD CAPITAL MANAGEMENT LLC

$100,357,338

4.10%

2,255,728

Conestoga Capital Advisors, LLC

$92,709,685

3.79%

2,083,832

WILLIAM BLAIR INVESTMENT MANAGEMENT, LLC

$88,662,118

3.62%

1,992,855

BNP PARIBAS ASSET MANAGEMENT Holding S.A.

$72,963,065

2.98%

1,639,988

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNP PARIBAS ASSET MANAGEMENT Europe	575	$25,581	0.0%	Defined
BNP Paribas Asset Management USA, Inc.	1,639,413	$72,937,484	100.0%	Defined

GW&K Investment Management, LLC

$68,981,610

2.82%

1,550,497

Soleus Capital Management, L.P.

$64,917,672

2.65%

1,459,152

GEODE CAPITAL MANAGEMENT, LLC

$59,671,944

2.44%

1,341,244

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	190,891	$8,492,740	14.2%	Defined
Held directly	1,150,353	$51,179,204	85.8%	Defined

GENEVA CAPITAL MANAGEMENT LLC

$59,577,582

2.44%

1,339,123

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nationwide Fund Advisors	405,739	$18,051,328	30.3%	Other
Held directly	933,384	$41,526,254	69.7%	Sole

LOOMIS SAYLES & CO L P

$53,398,633

2.18%

1,200,239

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
LOOMIS SAYLES TRUST COMPANY, LLC	201,943	$8,984,444	16.8%	Defined
Held directly	998,296	$44,414,189	83.2%	Defined

VANGUARD PORTFOLIO MANAGEMENT LLC

$47,192,900

1.93%

1,060,753

BANK OF AMERICA CORP /DE/

$45,210,024

1.85%

1,016,184

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	340,718	$15,158,543	33.5%	Defined
BOFA SECURITIES, INC.	75,374	$3,353,389	7.4%	Defined
MANAGED ACCOUNT ADVISORS LLC joint	600,092	$26,698,092	59.1%	Defined

DIMENSIONAL FUND ADVISORS LP

$40,316,835

1.65%

906,200

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	7,979	$354,985	0.9%	Defined
DFA Australia Ltd.	9,211	$409,797	1.0%	Defined
Dimensional Fund Advisors Pte. Ltd.	321	$14,281	0.0%	Defined
Dimensional Ireland Ltd	25,210	$1,121,592	2.8%	Defined
Held directly	863,479	$38,416,180	95.3%	Sole

SILVERCREST ASSET MANAGEMENT GROUP LLC

$39,892,002

1.63%

896,651

ROYAL BANK OF CANADA

$34,513,647

1.41%

775,762

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC CMA LTD.	1,548	$68,870	0.2%	Defined
RBC Capital Markets, LLC	142,475	$6,338,712	18.4%	Defined
RBC Global Asset Management (U.S.) Inc.	495,012	$22,023,083	63.8%	Defined
RBC Dominion Securities Inc.	36,199	$1,610,493	4.7%	Defined
RBC Trust Co (Delaware) Ltd	3,500	$155,715	0.5%	Defined
RBC Rochdale, LLC	75	$3,336	0.0%	Defined
Alaska Permanent Fund Corp joint	95,950	$4,268,815	12.4%	Defined
Held directly	1,003	$44,623	0.1%	Sole

Champlain Investment Partners, LLC

$33,626,787

1.37%

755,828

Aberdeen Group plc

$31,598,488

1.29%

710,238

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
abrdn Inc.	710,238	$31,598,488	100.0%	Defined

NOMURA ASSET MANAGEMENT INTERNATIONAL INC.

$29,554,929

1.21%

664,305

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NOMURA HOLDINGS INC joint	664,305	$29,554,929	100.0%	Defined

NEXT CENTURY GROWTH INVESTORS LLC

$27,253,862

1.11%

612,584

NORTHERN TRUST CORP

$26,589,046

1.09%

597,641

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	537,228	$23,901,273	89.9%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	35,270	$1,569,162	5.9%	Defined
MASON STREET ADVISORS, LLC joint	17,157	$763,314	2.9%	Other
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	7,986	$355,297	1.3%	Other

TWO SIGMA INVESTMENTS, LP

$25,847,889

1.06%

580,982

SG Capital Management LLC

$25,415,001

1.04%

571,252

TimesSquare Capital Management, LLC

$24,462,648

1.00%

549,846

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GuideStone Capital Management, LLC joint	549,846	$24,462,648	100.0%	Sole

Nuveen, LLC

$23,388,570

0.96%

525,704

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	35,679	$1,587,358	6.8%	Defined
TIAA CREF INVESTMENT MANAGEMENT LLC joint	73,186	$3,256,045	13.9%	Defined
TEACHERS ADVISORS, LLC joint	416,839	$18,545,167	79.3%	Defined

GOLDMAN SACHS GROUP INC

$23,384,833

0.96%

525,620

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	408,636	$18,180,215	77.7%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	116,984	$5,204,618	22.3%	Defined

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$22,601,498

0.92%

508,013

BROWN CAPITAL MANAGEMENT LLC

$22,425,140

0.92%

504,049

D. E. Shaw & Co., Inc.

$20,122,337

0.82%

452,289

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	452,289	$20,122,337	100.0%	Defined

Point72 Asset Management, L.P.

$19,724,018

0.81%

443,336

RAYMOND JAMES FINANCIAL INC

$19,718,322

0.81%

443,208

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	125,915	$5,601,958	28.4%	Defined
Raymond James Financial Services Advisors, Inc.	3,018	$134,270	0.7%	Defined
ClariVest Asset Management LLC joint	2,611	$116,163	0.6%	Defined
EAGLE ASSET MANAGEMENT INC	311,664	$13,865,931	70.3%	Defined

SEI INVESTMENTS CO

$19,575,154

0.80%

439,990

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI INVESTMENTS MANAGEMENT CORP	15,263	$679,050	3.5%	Sole
SEI TRUST CO	424,727	$18,896,104	96.5%	Sole

MORGAN STANLEY

$17,976,715

0.73%

404,062

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	26	$1,156	0.0%	Defined
MORGAN STANLEY & CO. LLC	18,231	$811,097	4.5%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	209,233	$9,308,776	51.8%	Defined
MORGAN STANLEY SMITH BARNEY LLC	10,430	$464,030	2.6%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	162,142	$7,213,696	40.1%	Defined
EATON VANCE MANAGEMENT	4,000	$177,960	1.0%	Defined

OSTERWEIS CAPITAL MANAGEMENT INC

$17,723,258

0.72%

398,365

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
OSTERWEIS CAPITAL MANAGEMENT LLC/CA	398,365	$17,723,258	100.0%	Sole

VANGUARD FIDUCIARY TRUST CO

$16,262,385

0.66%

365,529

UBS Group AG

$15,683,924

0.64%

352,527

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	18,030	$802,154	5.1%	Defined
UBS AG	303,987	$13,524,381	86.2%	Defined
UBS Switzerland AG	30,510	$1,357,389	8.7%	Defined

Balyasny Asset Management L.P.

$15,382,817

0.63%

345,759

FRONTIER CAPITAL MANAGEMENT CO LLC

$14,696,025

0.60%

330,322

Invesco Ltd.

$13,755,505

0.56%

309,182

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	20,349	$905,326	6.6%	Defined
Invesco Trust Co joint	20,571	$915,203	6.7%	Defined
Invesco Capital Management LLC	268,262	$11,934,976	86.8%	Defined

JPMORGAN CHASE & CO

$13,247,296

0.54%

297,759

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	25,860	$1,150,511	8.7%	Defined
J.P. Morgan Investment Management Inc.	68,521	$3,048,499	23.0%	Defined
J.P. Morgan Securities LLC	23,119	$1,028,564	7.8%	Defined
J.P. Morgan Securities plc	4,149	$184,589	1.4%	Defined
JPMorgan Asset Management (UK) Ltd	176,079	$7,833,754	59.1%	Defined
55I, LLC	31	$1,379	0.0%	Defined

AQR CAPITAL MANAGEMENT LLC

$12,605,258

0.52%

283,328

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Petros Family Wealth, LLC	89	$3,959	0.0%	Other
RF&L WEALTH MANAGEMENT, LLC	14	$622	0.0%	Other
Prism Planning Partners LLC	164	$7,296	0.1%	Other
Compound Planning, Inc.	34	$1,512	0.0%	Other
Arax Advisory Partners	1,850	$82,306	0.7%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	1,079	$48,004	0.4%	Other
Stokes Family Office, LLC	226	$10,054	0.1%	Other
Held directly	279,872	$12,451,505	98.8%	Defined

Bank of New York Mellon Corp

$12,366,348

0.51%

277,958

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Investments Corporation	136,096	$6,054,911	49.0%	Defined
BNY Mellon Investment Adviser, Inc.	41,195	$1,832,765	14.8%	Defined
The Bank of New York Mellon	95,246	$4,237,494	34.3%	Defined
BNY Mellon, NA joint	2,594	$115,406	0.9%	Defined
BNY Mellon Advisors, Inc.	1,288	$57,303	0.5%	Defined
Newton Investment Management North America, LLC	1,530	$68,069	0.6%	Defined
Held directly	9	$400	0.0%	Defined

ENVESTNET ASSET MANAGEMENT INC

$12,231,012

0.50%

274,916

Woodline Partners LP

$12,090,290

0.49%

271,753

PRINCIPAL FINANCIAL GROUP INC

$11,107,951

0.45%

249,673

JANE STREET GROUP, LLC

$11,041,216

0.45%

248,173

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	163,609	$7,278,964	65.9%	Defined
Jane Street Global Trading, LLC	84,564	$3,762,252	34.1%	Defined

FEDERATED HERMES, INC.

$10,304,550

0.42%

231,615

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FEDERATED GLOBAL INVESTMENT MANAGEMENT CORP	201,612	$8,969,717	87.0%	Defined
HERMES INVESTMENT MANAGEMENT LTD	30,003	$1,334,833	13.0%	Defined

Summit Creek Advisors LLC

$9,757,190

0.40%

219,312

AMERIPRISE FINANCIAL INC

$8,806,261

0.36%

197,938

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	197,288	$8,777,343	99.7%	Defined
Ameriprise Financial Services, LLC	650	$28,918	0.3%	Defined

PEREGRINE CAPITAL MANAGEMENT LLC

$8,254,185

0.34%

185,529

Fisher Asset Management, LLC

$8,245,643

0.34%

185,337

FIFTH THIRD BANCORP

$8,232,918

0.34%

185,051

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fifth Third Bank, National Association	185,051	$8,232,918	100.0%	Defined

OSAIC HOLDINGS, INC.

$8,098,869

0.33%

182,038

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
OSAIC WEALTH, INC.	7,046	$313,476	3.9%	Defined
OSAIC INSTITUTIONS, INC.	84	$3,737	0.0%	Defined
CW Advisors, LLC	174,743	$7,774,316	96.0%	Sole
Osaic Advisory Services, LLC	165	$7,340	0.1%	Defined

Public Sector Pension Investment Board

$7,803,946

0.32%

175,409

First Washington CORP

$7,579,093

0.31%

170,355

Thrivent Financial for Lutherans

$7,575,400

0.31%

170,272

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Thrivent Asset Management, LLC	57,342	$2,551,145	33.7%	Defined
Held directly	112,930	$5,024,255	66.3%	Sole

Allspring Global Investments Holdings, LLC

$6,969,179

0.28%

156,646

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Allspring Funds Management, LLC reporting on behalf of Morgan Stanley Smith Barney, LLC	6	$266	0.0%	Defined
Allspring Global Investments, LLC reporting on behalf of UBS ASSET MANAGEMENT AMERICAS INC	156,640	$6,968,913	100.0%	Defined

Russell Investments Group, Ltd.

$6,837,221

0.28%

153,680

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC	93,682	$4,167,911	61.0%	Defined
Russell Investments Capital, LLC	45,355	$2,017,843	29.5%	Defined
Russell Investments Trust Company	14,643	$651,467	9.5%	Defined

SEGALL BRYANT & HAMILL, LLC

$6,761,501

0.28%

151,978

MUTUAL OF AMERICA CAPITAL MANAGEMENT LLC

$6,470,848

0.26%

145,445

GLENMEDE TRUST CO NA

$6,225,084

0.25%

139,921

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Glenmede Investment Management, LP	123,353	$5,487,974	88.2%	Sole
Held directly	16,568	$737,110	11.8%	Sole

RICE HALL JAMES & ASSOCIATES, LLC

$6,176,991

0.25%

138,840

EFG International AG

$5,678,792

0.23%

127,642

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
EFG Asset Management (North America) Corp.	127,588	$5,676,390	100.0%	Defined
EFG Private Wealth Management (Americas) Corp.	54	$2,402	0.0%	Defined

BNP PARIBAS ARBITRAGE, SA

$5,573,395

0.23%

125,273

Granite Investment Partners, LLC

$5,414,255

0.22%

121,696

RHUMBLINE ADVISERS

$5,402,776

0.22%

121,438

Lisanti Capital Growth, LLC

$5,355,706

0.22%

120,380

Legal & General Group Plc

$5,144,378

0.21%

115,630

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	78,372	$3,486,770	67.8%	Defined
Legal & General Investment Management Ltd joint	37,258	$1,657,608	32.2%	Defined

Susquehanna Portfolio Strategies, LLC

$5,047,434

0.21%

113,451

KORNITZER CAPITAL MANAGEMENT INC /KS

$5,033,020

0.21%

113,127

Trexquant Investment LP

$4,698,544

0.19%

105,609

Swiss National Bank

$4,546,878

0.19%

102,200

Jupiter Topco LLC

$4,417,633

0.18%

99,295

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	21,659	$963,608	21.8%	Defined
JANUS HENDERSON INVESTORS US LLC	77,636	$3,454,025	78.2%	Defined

WELLS FARGO & COMPANY/MN

$4,397,078

0.18%

98,833

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	47,061	$2,093,743	47.6%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	17,991	$800,419	18.2%	Defined
WELLS FARGO CLEARING SERVICES, LLC	33,781	$1,502,916	34.2%	Defined

Longaeva Partners L.P.

$4,157,679

0.17%

93,452

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Balyasny Asset Management L.P.	93,452	$4,157,679	100.0%	Other

STATE OF WISCONSIN INVESTMENT BOARD

$4,016,468

0.16%

90,278

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$3,725,722

0.15%

83,743

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	1,185	$52,720	1.4%	Defined
UBS Asset Management Switzerland AG	61,171	$2,721,497	73.0%	Sole
UBS Asset Management (UK) Ltd.	14,986	$666,726	17.9%	Defined
Held directly	6,401	$284,779	7.6%	Defined

SIT INVESTMENT ASSOCIATES INC

$3,700,900

0.15%

83,185

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SIT FIXED INCOME ADVISORS II LLC /ADV	83,185	$3,700,900	100.0%	Sole

Universal- Beteiligungs- und Servicegesellschaft mbH

$3,637,858

0.15%

81,768

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Universal-Investment-Gesellschaft mbH joint	81,768	$3,637,858	100.0%	Defined

MARSHALL WACE, LLP

$3,505,634

0.14%

78,796

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marshall Wace North America L.P. joint	78,796	$3,505,634	100.0%	Other

ALLIANCEBERNSTEIN L.P.

$3,365,757

0.14%

75,652

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	75,652	$3,365,757	100.0%	Defined

PRICE T ROWE ASSOCIATES INC /MD/

$3,356,414

0.14%

75,442

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	75,442	$3,356,414	100.0%	Defined

HARBOR CAPITAL ADVISORS, INC.

$3,241,185

0.13%

72,852

BARCLAYS PLC

$3,235,667

0.13%

72,728

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	68,057	$3,027,855	93.6%	Defined
BARCLAYS CAPITAL INC.	2,278	$101,348	3.1%	Defined
BARCLAYS CAPITAL SECURITIES LTD	2,393	$106,464	3.3%	Defined

Rhenman & Partners Asset Management AB

$3,114,300

0.13%

70,000

Man Group plc

$2,998,314

0.12%

67,393

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Numeric Investors LLC	67,393	$2,998,314	100.0%	Defined

Cerity Partners LLC

$2,989,281

0.12%

67,190

DEUTSCHE BANK AG\

$2,865,332

0.12%

64,404

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Deutsche Bank Trust Co Americas	8,003	$356,053	12.4%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC	967	$43,021	1.5%	Defined
DWS Investments UK Ltd	52,723	$2,345,646	81.9%	Defined
DWS International GmbH	2,711	$120,612	4.2%	Defined

INTECH INVESTMENT MANAGEMENT LLC

$2,857,859

0.12%

64,236

Global Alpha Capital Management Ltd.

$2,853,633

0.12%

64,141

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 1	64,141	$2,853,633	100.0%	Sole

NEW YORK STATE COMMON RETIREMENT FUND

$2,835,792

0.12%

63,740

FMR LLC

$2,801,134

0.11%

62,961

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	62,060	$2,761,049	98.6%	Defined
STRATEGIC ADVISERS LLC	337	$14,993	0.5%	Defined
FIAM LLC	564	$25,092	0.9%	Defined

STIFEL FINANCIAL CORP

$2,710,419

0.11%

60,922

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STIFEL NICOLAUS & CO INC \MO\	60,813	$2,705,570	99.8%	Defined
Stifel Independent Advisors, LLC	43	$1,913	0.1%	Defined
Held directly	66	$2,936	0.1%	Defined

CITIGROUP INC

$2,603,777

0.11%

58,525

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	58,525	$2,603,777	100.0%	Defined

CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM

$2,485,656

0.10%

55,870

MARTINGALE ASSET MANAGEMENT L P

$2,392,449

0.10%

53,775

Bridge City Capital, LLC

$2,381,727

0.10%

53,534

PRUDENTIAL FINANCIAL INC

$2,366,467

0.10%

53,191

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PGIM Quantitative Solutions LLC	53,191	$2,366,467	100.0%	Defined

Legato Capital Management LLC

$2,314,369

0.09%

52,020

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Key facts CIK 887359 CUSIP 92346J108 13F (30d) 31 filings 31 filers Visit website Investor relations