AORT · Artivion, Inc. · Holdings

$23.78 +0.88 (+3.82%)

S&P 600 member Russell 2000 member Rebalance outlook locked

Market Cap

$1.12B

48.76M

Volume · Oct 22 80.24K Avg daily vol (3M) 633.97K

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Use institutional holdings through the 13F API or MCP → See every holder of AORT: 13F, 13D and 13G, insiders and funds →

Institutional holdings are being reconciled with SEC filings. Holder totals and changes may be incomplete until reconciliation finishes.

The Sep 30, 2026 filing window is still open — showing the latest known positions. Funds that have not filed yet carry their prior-quarter positions; a fund that filed without this stock counts as sold out. Each fund's Change is measured against its own previous reported quarter, shown in the Reported column. Pick a completed quarter in the Quarter selector below to put every fund on one quarter and see the stock-wide activity breakdown.

Quarter:

Holders: 210

Total Value: $1,032,052,398

Total Shares: 45,929,152

Showing 1–100 of 210 holdings by value · page 1 of 3

Institutional Ownership Trend

27 quarters

Holder Concentration

Across 206 13F filers

Top Holder

14.5%

of institutional value

Top 5

35.8%

of institutional value

Top 10

55.4%

of institutional value

HHI

459

Broadly held

Who Holds AORT

Last 8 quarters through Sep 30, 2026 (filing window open)

Each kind of 13F filer's share of the common shares 13F filers report. Holder type comes from regulatory records, never the filer's name; style, turnover and horizon are measured from each filer's own 13F book. The quarter selector does not change this card.

Jun 30, 2026, the newest closed quarter: 206 filers, like-for-like change +3.5%; actively managed books +2.3%; index-like books +5.5%.

Holder type	Share	Filers	Change	New	Exited
Registered adviser	41.9%	103	-2.4%	18	43
Exempt adviser	1.2%	3	+106.8%	2	1
Adviser	12.2%	7	+4.2%	2	3
Bank	9.0%	17	+14.2%	2	1
Insurer	0.7%	3	+4.8%	1	0
Benefit plan	0.1%	2	-23.2%	0	2
Non-US	0.8%	6	+9.4%	0	1
Holding company	28.1%	6	+0.6%	0	2
Broker-dealer	0.0%	0	-100.0%	0	1
Unknown	6.0%	59	+49.1%	8	12

Style	Share	Filers	Change	New	Exited
Index-like	36.5%	30	+5.5%	0	3
Systematic-like	6.7%	21	+101.3%	8	8
Concentrated	5.9%	16	+32.2%	6	11
Diversified active	50.9%	139	-6.2%	19	44
Not measured	0.0%	0	—	0	0
Unknown	0.0%	0	—	0	0

Turnover	Share	Filers	Change	New	Exited
Low turnover	59.8%	120	+3.9%	13	29
Medium turnover	13.6%	42	-34.7%	6	23
High turnover	12.2%	31	+123.1%	13	10
Unknown	14.3%	13	+12.5%	1	4
Not measured	0.0%	0	—	0	0

Holding horizon	Share	Filers	Change	New	Exited
Short horizon	18.9%	26	+24.7%	7	9
Medium horizon	16.2%	23	-1.7%	8	14
Long horizon	56.8%	144	+3.6%	15	36
Unknown	8.1%	13	-20.7%	3	7
Not measured	0.0%	0	—	0	0

13F book size	Share	Filers	Change	New	Exited
Under $1B	6.5%	23	+20.0%	8	19
$1B to $10B	14.7%	63	-5.7%	16	18
$10B to $100B	20.7%	65	+1.4%	9	23
$100B or more	58.2%	55	+5.2%	0	6
Not measured	0.0%	0	—	0	0

Client base	Share	Filers	Change	New	Exited
Individuals and high net worth	6.9%	48	+20.9%	11	20
Pooled investment vehicles	17.3%	22	+31.1%	7	7
Registered funds and BDCs	16.0%	15	-12.8%	0	5
Pension and profit sharing plans	0.0%	2	+33.7%	0	0
Insurance companies	0.1%	2	-8.7%	0	1
Governments and sovereign funds	0.5%	3	-38.8%	0	2
Other investment advisers	0.0%	3	-1.5%	0	0
Other	0.1%	1	-9.2%	0	0
Diversified	1.0%	7	-78.0%	0	8
Unknown	58.1%	103	+8.1%	15	23

Private funds	Share	Filers	Change	New	Exited
Hedge funds	15.3%	32	+1.6%	9	11
Private equity funds	4.7%	9	+63.2%	1	0
Venture capital funds	0.0%	1	-55.6%	0	0
Real estate funds	0.0%	2	-54.3%	1	2
Securitized asset funds	0.1%	1	-30.2%	0	0
Other private funds	5.3%	11	-34.9%	0	6
Mixed private funds	0.1%	2	+59.6%	1	0
Private funds, type not stated	0.7%	2	-0.5%	0	1
No private funds	16.8%	46	+2.2%	8	24
Unknown	56.9%	100	+7.1%	13	22

Location	Share	Filers	Change	New	Exited
United States	91.7%	177	+0.4%	30	57
Outside the US	8.3%	29	+54.8%	3	9

The Sep 30, 2026 filing window is still open, so its bar carries the Jun 30, 2026 positions of filers that have not filed yet.

How the mix is measured

Shares are 13F common shares restated to today's split basis; options and principal amounts are left out. The change compares only filers with a 13F in both quarters and keeps each filer in its bucket of the later quarter on both sides; a filer that continues under a new CIK is compared through its successor. Holder type comes from each filer's current regulatory profile; client base and private funds come from the Form ADV roster current at each quarter end (rosters from December 2019, so an earlier quarter reads Unknown). Style, turnover, horizon and book size are measured from each filer's 13F book in the newest measured quarter, and Not measured means none was. Actively managed books are those measured systematic-like, concentrated or diversified active.

Excluded 2 holders with no observed 2026-03-31 13F and 2 holders with no observed 2026-06-30 13F. Compared across CIK changes: JANUS HENDERSON GROUP PLC continues as Jupiter Topco LLC.

Conviction and Crowding

206 13F filers, quarter ended Jun 30, 2026

Measured across every 13F filer's common-share book once the quarter's filing window closed. Crowding counts filers with high turnover or that advise private funds on Form ADV.

Conviction holders

0.971% of filers hold it in their top 10

Mar 31, 2026: 3

Holders above 5%

median weight 0.001%

Mar 31, 2026: 2

Crowding

31.2%

of shares outstanding, 83rd percentile

Mar 31, 2026: 28.1%

Private-fund advisers

24.8%

of shares outstanding

Institutional ownership

94.6%

100.0% of the estimated free float

Top 10 holders

55.4%

of 13F shares, Herfindahl 0.046

Active holders

+2.3%

like-for-like share change of active filers

5% stakes on record

0 13D, 3 13G (3 institutional, 0 passive, 0 exempt)

How long holders have held it value-weighted 13.7 quarters

Under 1 year 28.9% 1 to 3 years 24.6% 3 years or more 46.5% Since coverage start 0.000%

Managers in the top fifth by current three-year FundScore alpha were net buyers of $23,237,791 (1 buying, 0 selling).

Estimated Holder Cost Basis

Quarter ended Jun 30, 2026, on that quarter's share basis

Estimated, never filed: each 13F buy is costed at its quarter's average traded price and each sale keeps the average cost, so this is what holders paid on average if they traded evenly through each quarter.

Estimated avg cost

$29.30

estimated per share

Mar 31, 2026: $31.09

Quarter-end price

$22.47

implied by 13F values

Estimated gain or loss

-23.3%

price against the estimated cost

Estimated at a loss

80.0%

of costed 13F value

Estimated cost covers 37.2% of the 13F shares; positions already held when a filer's 13F history starts, or bought in a quarter without a trusted average price, have no estimate.

Shared Holders

Companies AORT's active holders also own, quarter ended Jun 30, 2026

#	Company	Shared holders	Similarity
1	SION Sionna Therapeutics, Inc.	4	0.218
2	AVAH Aveanna Healthcare Holdings, Inc.	3	0.215
3	ANRO Alto Neuroscience, Inc.	3	0.191
4	TRNS Transcat Inc	3	0.186
5	VIR Vir Biotechnology, Inc.	3	0.186
6	EWTX Edgewise Therapeutics, Inc.	4	0.182
7	DRUG Bright Minds Biosciences Inc.	3	0.177
8	RLMD Relmada Therapeutics, Inc.	3	0.166
9	EYPT EyePoint, Inc.	3	0.163
10	PALI Palisade Bio, Inc.	3	0.160

Shareholder Targets

13F filers that own its peers but not AORT, quarter ended Jun 30, 2026

Found through its holder-overlap peers: SION, AVAH, ANRO, TRNS, VIR, EWTX, DRUG, RLMD, EYPT, PALI, IRTC, URGN, VCEL, BCRX, ANIP, SPRY, MBX, AVTX, SNDX, ERAS. 770 filers in all.

#	Filer	In peers	Of book	Largest peers
1	RA CAPITAL MANAGEMENT, L.P. Registered adviser, Concentrated	$1,073,960,235 7 held	9.46%	SION, EWTX, BCRX
2	Frazier Life Sciences Management, L.P. Registered adviser, Concentrated	$864,828,961 3 held	18.8%	ERAS, MBX, BCRX
3	Paradigm Biocapital Advisors LP Registered adviser, Concentrated	$769,785,041 4 held	16.7%	EWTX, ERAS, MBX
4	RTW INVESTMENTS, LP Registered adviser, Diversified active	$758,857,826 4 held	6.48%	ERAS, IRTC, URGN
5	Bain Capital Investors LLC Unknown, Concentrated	$699,319,541 1 held	14.5%	AVAH
6	MILLENNIUM MANAGEMENT LLC Registered adviser, Systematic-like	$410,049,315 16 held	0.310%	IRTC, SNDX, ANIP
7	Cormorant Asset Management, LP Registered adviser, Diversified active	$405,541,300 5 held	20.6%	ERAS, EWTX, EYPT
8	Venrock Adviser, LLC Registered adviser, Concentrated	$388,335,585 3 held	11.8%	ERAS, RLMD, ANRO
9	T. Rowe Price Investment Management, Inc. Adviser, Diversified active	$388,124,570 1 held	0.260%	ERAS
10	Siren, L.L.C. Unknown, Diversified active	$369,235,377 4 held	7.50%	ERAS, EWTX, SION

Fund Ownership of AORT by Fund Type

Registered funds' Form N-PORT reports for the quarter ended Jun 30, 2026

Index funds and index ETFs hold 22.37% of shares outstanding.

Fund type	Of shares outstanding	Change	Shares	Funds
Index funds	4.21%	+0.44 pp	2,051,667	73
Index ETFs	18.16%	+0.39 pp	8,853,054	54
Active ETFs	1.21%	+0.11 pp	590,027	19
Active funds	16.95%	+1.19 pp	8,265,386	84
Unclassified	0.23%	+0.20 pp	110,118	5
All registered funds	40.75%	+2.33 pp	19,870,252	235

How fund types are measured

Each fund series counts once, with its latest Form N-PORT report dated inside the quarter; only long share positions count, restated to today's split basis. The fund type is the series' latest Form N-CEN answer on or before the quarter end (the index-fund box and the exchange-traded flag), never its name. Unclassified funds have no usable N-CEN; unit investment trusts are among them, so index ownership is a floor. Funds sit inside their advisers' 13F reports, so these shares overlap the 13F figures and are never added to them.

Holder type: Registered adviser 107 Exempt adviser 3 Adviser 7 Bank 17 Insurer 3 Benefit plan 2 Non-US 6 Holding company 6 Unknown 59

Showing 100 of 210

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

BlackRock, Inc.

$149,965,153

14.53%

6,674,017

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	35,134	$789,460	0.5%	Sole
BlackRock Investment Management (Australia) Ltd	11	$247	0.0%	Sole
BlackRock Asset Management Canada Ltd	220	$4,943	0.0%	Sole
BlackRock Investment Management, LLC	139,271	$3,129,419	2.1%	Sole
BLACKROCK ADVISORS LLC	102,485	$2,302,837	1.5%	Sole
BlackRock Fund Advisors	4,900,759	$110,120,054	73.4%	Sole
BlackRock Institutional Trust Company, N.A.	1,305,487	$29,334,292	19.6%	Sole
BlackRock Fund Managers Ltd.	6,099	$137,044	0.1%	Sole
BlackRock Investment Management (UK) Ltd.	21,840	$490,744	0.3%	Sole
BlackRock Asset Management Ireland Ltd	118,721	$2,667,660	1.8%	Sole
BlackRock Asset Management Schweiz AG	6,153	$138,257	0.1%	Sole
Aperio Group, LLC	37,814	$849,680	0.6%	Sole
SpiderRock Advisors, LLC	23	$516	0.0%	Sole

NOMURA ASSET MANAGEMENT INTERNATIONAL INC.

$71,566,365

6.93%

3,184,974

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NOMURA HOLDINGS INC joint	3,184,974	$71,566,365	100.0%	Defined

MORGAN STANLEY

$57,477,380

5.57%

2,557,961

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	16	$359	0.0%	Defined
MORGAN STANLEY & CO. LLC	18,500	$415,695	0.7%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	199,472	$4,482,135	7.8%	Defined
MORGAN STANLEY SMITH BARNEY LLC	2,114,398	$47,510,522	82.7%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	225,575	$5,068,669	8.8%	Defined

Juniper Investment Company, LLC

$45,386,815

4.40%

2,019,885

VANGUARD CAPITAL MANAGEMENT LLC

$45,043,788

4.36%

2,004,619

FMR LLC

$44,119,191

4.27%

1,963,471

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	1,863,440	$41,871,496	94.9%	Defined
STRATEGIC ADVISERS LLC	872	$19,593	0.0%	Defined
Fidelity Institutional Asset Management Trust Co	89,231	$2,005,020	4.5%	Defined
FIAM LLC	9,928	$223,082	0.5%	Defined

STATE STREET CORP

$41,750,405

4.05%

1,858,051

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	1,858,051	$41,750,405	100.0%	Defined

Ophir Asset Management Pty Ltd

$40,446,179

3.92%

1,800,008

PERCEPTIVE ADVISORS LLC

$39,036,906

3.78%

1,737,290

Conestoga Capital Advisors, LLC

$36,484,539

3.54%

1,623,700

WELLINGTON MANAGEMENT GROUP LLP

$28,318,466

2.74%

1,260,279

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wellington Management Company LLP	520,795	$11,702,263	41.3%	Defined
Wellington Managment Canada LLC	9,388	$210,948	0.7%	Defined
Wellington Management International Ltd	22,319	$501,507	1.8%	Defined
Wellington Trust Company, NA joint	650,831	$14,624,172	51.6%	Defined
Wellington Management Australia Pty Ltd	35,466	$796,921	2.8%	Defined
Wellington Management Europe GmbH	21,480	$482,655	1.7%	Defined

GEODE CAPITAL MANAGEMENT, LLC

$25,038,253

2.43%

1,114,297

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	160,932	$3,616,142	14.4%	Defined
Held directly	953,365	$21,422,111	85.6%	Defined

SILVERCREST ASSET MANAGEMENT GROUP LLC

$23,011,324

2.23%

1,024,091

VANGUARD PORTFOLIO MANAGEMENT LLC

$22,205,280

2.15%

988,219

DIMENSIONAL FUND ADVISORS LP

$21,300,231

2.06%

947,941

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	8,385	$188,410	0.9%	Defined
DFA Australia Ltd.	14,036	$315,388	1.5%	Defined
Dimensional Fund Advisors Pte. Ltd.	358	$8,044	0.0%	Defined
Dimensional Ireland Ltd	32,289	$725,533	3.4%	Defined
Held directly	892,873	$20,062,856	94.2%	Sole

CITADEL ADVISORS LLC

$18,596,037

1.80%

827,594

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	39,796	$894,216	4.8%	Defined
Held directly	787,798	$17,701,821	95.2%	Defined

GW&K Investment Management, LLC

$17,851,919

1.73%

794,478

ROYCE & ASSOCIATES LP

$14,364,666

1.39%

639,282

YARGER WEALTH STRATEGIES, LLC

$14,250,990

1.38%

634,223

NORTHERN TRUST CORP

$12,370,450

1.20%

550,532

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	500,772	$11,252,345	91.0%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	28,593	$642,484	5.2%	Defined
MASON STREET ADVISORS, LLC joint	14,197	$319,006	2.6%	Other
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	6,970	$156,615	1.3%	Other

BANK OF AMERICA CORP /DE/

$11,716,687

1.14%

521,437

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	85,115	$1,912,534	16.3%	Defined
MERRILL LYNCH, PIERCE, FENNER & SMITH INC.	11,154	$250,630	2.1%	Defined
BOFA SECURITIES, INC.	43,853	$985,376	8.4%	Defined
MANAGED ACCOUNT ADVISORS LLC joint	381,315	$8,568,147	73.1%	Defined

Qube Research & Technologies Ltd

$10,704,820

1.04%

476,405

FRED ALGER MANAGEMENT, LLC

$10,406,104

1.01%

463,111

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Weatherbie Capital, LLC	463,111	$10,406,104	100.0%	Sole

UBS Group AG

$10,210,389

0.99%

454,401

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	268,204	$6,026,543	59.0%	Defined
UBS AG	184,535	$4,146,501	40.6%	Defined
UBS Switzerland AG	1,662	$37,345	0.4%	Defined

Invesco Ltd.

$9,956,837

0.96%

443,117

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	83,772	$1,882,355	18.9%	Defined
Invesco Trust Co joint	18,230	$409,628	4.1%	Defined
Invesco Capital Management LLC	341,115	$7,664,854	77.0%	Defined

GOLDMAN SACHS GROUP INC

$9,857,903

0.96%

438,714

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	344,152	$7,733,095	78.4%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	94,562	$2,124,808	21.6%	Defined

D. E. Shaw & Co., Inc.

$9,846,713

0.95%

438,216

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	438,216	$9,846,713	100.0%	Defined

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$9,183,578

0.89%

408,704

Soleus Capital Management, L.P.

$8,718,360

0.84%

388,000

TWO SIGMA INVESTMENTS, LP

$8,702,159

0.84%

387,279

Balyasny Asset Management L.P.

$8,584,685

0.83%

382,051

First Eagle Investment Management, LLC

$7,787,494

0.75%

346,573

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FIRST EAGLE SEPARATE ACCOUNT MANAGEMENT, LLC	131	$2,943	0.0%	Sole
Held directly	346,442	$7,784,551	100.0%	Sole

VANGUARD FIDUCIARY TRUST CO

$6,978,103

0.68%

310,552

KENNEDY CAPITAL MANAGEMENT LLC

$6,611,370

0.64%

294,231

Legal & General Group Plc

$5,826,425

0.56%

259,298

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	65,374	$1,468,953	25.2%	Defined
Legal & General Investment Management Ltd joint	193,924	$4,357,472	74.8%	Defined

PRINCIPAL FINANCIAL GROUP INC

$5,705,177

0.55%

253,902

Bank of New York Mellon Corp

$5,671,108

0.55%

252,386

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNY Mellon Trust of Delaware joint	25	$561	0.0%	Defined
Mellon Investments Corporation	112,257	$2,522,414	44.5%	Defined
BNY Mellon Investment Adviser, Inc.	34,602	$777,506	13.7%	Defined
The Bank of New York Mellon	94,288	$2,118,651	37.4%	Defined
BNY Mellon, NA joint	9,905	$222,564	3.9%	Defined
BNY Mellon Advisors, Inc.	1,307	$29,368	0.5%	Defined
Held directly	2	$44	0.0%	Defined

Quantinno Capital Management LP

$4,497,190

0.44%

200,142

AMERIPRISE FINANCIAL INC

$4,348,977

0.42%

193,546

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	193,461	$4,347,068	100.0%	Defined
Ameriprise Financial Services, LLC	85	$1,909	0.0%	Defined

HRT FINANCIAL LP

$4,187,284

0.41%

186,350

Tidal Investments LLC

$4,184,520

0.41%

186,227

FIFTH THIRD BANCORP

$4,170,903

0.40%

185,621

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fifth Third Bank, National Association	185,621	$4,170,903	100.0%	Defined

ORBIMED ADVISORS LLC

$3,932,250

0.38%

175,000

Corient Private Wealth LLC

$3,580,549

0.35%

159,348

Public Sector Pension Investment Board

$3,298,303

0.32%

146,787

Parkman Healthcare Partners LLC

$3,121,981

0.30%

138,940

Universal- Beteiligungs- und Servicegesellschaft mbH

$2,879,148

0.28%

128,133

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Universal-Investment-Gesellschaft mbH	128,133	$2,879,148	100.0%	Defined

VOLORIDGE INVESTMENT MANAGEMENT, LLC

$2,857,262

0.28%

127,159

Summit Creek Advisors LLC

$2,704,354

0.26%

120,354

CONNORS INVESTOR SERVICES INC

$2,700,444

0.26%

120,180

BNP PARIBAS ARBITRAGE, SA

$2,693,815

0.26%

119,885

Russell Investments Group, Ltd.

$2,469,609

0.24%

109,907

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC	18,260	$410,302	16.6%	Defined
Russell Investments Capital, LLC	57,994	$1,303,125	52.8%	Defined
Russell Investments France SAS	10,940	$245,821	10.0%	Defined
Russell Investments Trust Company	22,713	$510,361	20.7%	Defined

Nuveen, LLC

$2,446,824

0.24%

108,893

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	17,906	$402,347	16.4%	Defined
TEACHERS ADVISORS, LLC joint	90,987	$2,044,477	83.6%	Defined

RHUMBLINE ADVISERS

$2,373,303

0.23%

105,621

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$2,303,871

0.22%

102,531

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	102,531	$2,303,871	100.0%	Other

XTX Topco Ltd

$2,173,905

0.21%

96,747

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	96,747	$2,173,905	100.0%	Defined

Swiss National Bank

$1,914,444

0.19%

85,200

ROYAL BANK OF CANADA

$1,905,341

0.18%

84,795

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC CMA LTD.	1,382	$31,053	1.6%	Defined
RBC Capital Markets, LLC	80,222	$1,802,588	94.6%	Defined
RBC Dominion Securities Inc.	1,019	$22,896	1.2%	Defined
RBC Rochdale, LLC	1,330	$29,885	1.6%	Defined
Held directly	842	$18,919	1.0%	Sole

JPMORGAN CHASE & CO

$1,903,184

0.18%

84,699

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
J.P. Morgan Trust Co of Delaware	462	$10,381	0.5%	Other
JPMorgan Chase Bank, N.A.	18,957	$425,963	22.4%	Defined
J.P. Morgan Investment Management Inc.	6,282	$141,156	7.4%	Defined
J.P. Morgan Securities plc	817	$18,357	1.0%	Defined
J.P. MORGAN MANSART MANAGEMENT Ltd	58,181	$1,307,327	68.7%	Defined

LAZARD ASSET MANAGEMENT LLC

$1,875,301

0.18%

83,458

PRICE T ROWE ASSOCIATES INC /MD/

$1,608,043

0.16%

71,564

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	71,564	$1,608,043	100.0%	Defined

RENAISSANCE TECHNOLOGIES LLC

$1,524,072

0.15%

67,827

CITIGROUP INC

$1,490,547

0.14%

66,335

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	66,335	$1,490,547	100.0%	Defined

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$1,424,709

0.14%

63,405

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	1,778	$39,951	2.8%	Defined
UBS Asset Management Switzerland AG	61,627	$1,384,758	97.2%	Sole

Squarepoint Ops LLC

$1,424,238

0.14%

63,384

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Squarepoint Capital LLP	63,384	$1,424,238	100.0%	Defined

DEUTSCHE BANK AG\

$1,416,867

0.14%

63,056

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DWS INVESTMENT MANAGEMENT AMERICAS, INC	865	$19,436	1.4%	Defined
DWS Investments UK Ltd	62,191	$1,397,431	98.6%	Defined

NEW YORK STATE TEACHERS RETIREMENT SYSTEM

$1,338,245

0.13%

59,557

EXCHANGE TRADED CONCEPTS, LLC

$1,320,876

0.13%

58,784

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Madison Avenue Financial Solutions, LLC	21,226	$476,948	36.1%	Other
Held directly	37,558	$843,928	63.9%	Sole

SEI INVESTMENTS CO

$1,288,046

0.12%

57,323

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI INVESTMENTS MANAGEMENT CORP	470	$10,560	0.8%	Sole
SEI TRUST CO	56,853	$1,277,486	99.2%	Sole

VICTORY CAPITAL MANAGEMENT INC

$1,193,157

0.12%

53,100

CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM

$1,119,859

0.11%

49,838

ENVESTNET ASSET MANAGEMENT INC

$1,117,972

0.11%

49,754

Sterling Capital Management LLC

$1,098,917

0.11%

48,906

Sio Capital Management, LLC

$1,089,368

0.11%

48,481

WELLS FARGO & COMPANY/MN

$1,067,481

0.10%

47,507

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	41,714	$937,313	87.8%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	3,980	$89,430	8.4%	Defined
WELLS FARGO CLEARING SERVICES, LLC	1,813	$40,738	3.8%	Defined

BOOTHBAY FUND MANAGEMENT, LLC

$1,010,520

0.10%

44,972

PRUDENTIAL FINANCIAL INC

$995,218

0.10%

44,291

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PGIM Quantitative Solutions LLC	44,291	$995,218	100.0%	Defined

ALLIANCEBERNSTEIN L.P.

$934,190

0.09%

41,575

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	41,575	$934,190	100.0%	Defined

FIRST TRUST ADVISORS LP

$857,657

0.08%

38,169

FRANKLIN RESOURCES INC

$819,480

0.08%

36,470

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FIDUCIARY TRUST CO INTERNATIONAL	24	$539	0.1%	Defined
FIDUCIARY TRUST INTERNATIONAL LLC	1,471	$33,053	4.0%	Defined
FRANKLIN ADVISERS INC	32,098	$721,242	88.0%	Defined
FRANKLIN ADVISORY SERVICES LLC	2,877	$64,646	7.9%	Defined

NATIXIS ADVISORS, LLC

$815,458

0.08%

36,291

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Natixis Investment Managers, L.P.	36,291	$815,458	100.0%	Defined

CANADA LIFE ASSURANCE Co

$749,396

0.07%

33,351

Zurcher Kantonalbank (Zurich Cantonalbank)

$738,274

0.07%

32,856

Y-Intercept (Hong Kong) Ltd

$730,764

0.07%

32,023

Verition Fund Management LLC

$729,398

0.07%

32,461

Aquatic Capital Management LLC

$712,770

0.07%

31,721

Jain Global LLC

$680,908

0.07%

30,303

Brevan Howard Capital Management LP

$652,461

0.06%

29,037

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Brevan Howard US Investment Management, LP	29,037	$652,461	100.0%	Defined

VANGUARD ASSET MANAGEMENT, Ltd

$634,193

0.06%

28,224

Thrivent Financial for Lutherans

$630,305

0.06%

28,051

JONES FINANCIAL COMPANIES LLLP

$630,103

0.06%

28,042

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Edward D. Jones & Co., L.P.	28,042	$630,103	100.0%	Sole

MetLife Investment Management, LLC

$601,050

0.06%

26,749

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
METLIFE INC	26,749	$601,050	100.0%	Defined

Mariner, LLC

$570,513

0.06%

25,390

TOCQUEVILLE ASSET MANAGEMENT L.P.

$561,750

0.05%

25,000

Illinois Municipal Retirement Fund

$545,009

0.05%

24,255

MANUFACTURERS LIFE INSURANCE COMPANY, THE

$516,854

0.05%

23,002

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MANULIFE FINANCIAL CORP joint	23,002	$516,854	100.0%	Defined

Walleye Capital LLC

$490,407

0.05%

21,825

Penn Capital Management Company, LLC

$489,688

0.05%

21,793

ALLSTATE CORP

$483,239

0.05%

21,506

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ALLSTATE INVESTMENTS LLC	21,506	$483,239	100.0%	Sole

Louisiana State Employees Retirement System

$474,656

0.05%

20,800

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Key facts CIK 784199 CUSIP 228903100 13F (30d) 23 filings 23 filers Visit website Investor relations