CDXS · Codexis, Inc. · Shareholders & Ownership
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Every disclosed holder of CDXS in one list: 13F institutions, Schedule 13D and 13G filers, insiders (Forms 3, 4 and 5) and registered funds, each from its own latest filing. Holders nest by CIK and SEC reporting links, never by name, and each is summed once.
Who Owns CDXS
13F quarter Sep 30, 2026, combined view: filers that have not filed yet carry their Jun 30, 2026 positions| Holder group | Shares | % Out |
|---|---|---|
| Institutional (13F) | 64,889,511 | 59.26% |
| Insiders (Forms 3, 4, 5) | 2,364,445 | 2.16% |
| 13D/G-only blocks | 13,746,692 | 12.55% |
| Remaining | 28,505,117 | 26.03% |
| Registered funds (N-PORT, not summed) | 24,527,490 | 22.40% |
Shares outstanding: 109,505,765, the issuer's latest cover-page count restated onto today's split basis.
13D/G-only counts each filer's latest 13D or 13G that no 13F report of the listing contains, that names no 13F filer and that no counted insider files. Funds are never summed: their shares sit inside their advisers' 13F reports.
Holders of CDXS
| # | Holder | Shares | As of | Notes |
|---|---|---|---|---|
| 1 |
Aberdeen Group plc
13G
CIK 1716774 · Holding company · Sep 30, 2026
Rule 13d-1(b), qualified institution, not summed
|
10,797,702 | ||
| 1.1 |
Aberdeen Group plc
13F
CIK 1716774 · Holding company · Jun 30, 2026
carried, not yet filed for the newest quarter
|
6,769,954 | ||
| 2 |
Telemark Asset Management, LLC
13G
CIK 1420816 · Unknown · Jul 24, 2026
Rule 13d-1(c), passive investor, not summed
|
7,846,411 | ||
| 2.1 |
Telemark Asset Management, LLC
13F
CIK 1420816 · Unknown · Jun 30, 2026
carried, not yet filed for the newest quarter
|
3,846,411 | ||
| 3 |
BlackRock, Inc.
13F
CIK 2012383 · Holding company · Jun 30, 2026
carried, not yet filed for the newest quarter
|
7,284,352 | ||
| 3.1 |
iShares Russell 2000 ETF
Fund
S000004344 · Registered fund · Jun 30, 2026
iShares Trust, not summed
|
2,286,444 | ||
| 3.2 |
iShares Russell 2000 Growth ETF
Fund
S000004343 · Registered fund · Jun 30, 2026
iShares Trust, not summed
|
696,097 | ||
| 3.3 |
iShares Biotechnology ETF
Fund
S000004350 · Registered fund · Jun 30, 2026
iShares Trust, not summed
|
593,659 | ||
| 3.4 |
iShares Micro-Cap ETF
Fund
S000004439 · Registered fund · Jun 30, 2026
iShares Trust, not summed
|
290,804 | ||
| 3.5 |
Master Small Cap Index Series
Fund
S000002716 · Registered fund · Jun 30, 2026
Quantitative Master Series LLC, not summed
|
137,295 | ||
| 4 |
Opaleye Management Inc.
13F
CIK 1595855 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
6,785,000 | ||
| 4.1 |
|
10,134,989 | ||
| 5 |
|
5,592,972 | ||
| 6 |
MILLENNIUM MANAGEMENT LLC
13F
CIK 1273087 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
5,549,139 | ||
| 7 |
|
4,734,627 | ||
| 8 |
AMERIPRISE FINANCIAL INC
13F
CIK 820027 · Investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
3,809,440 | ||
| 8.1 |
Columbia Small Cap Value Discovery Fund
Fund
S000012102 · Registered fund · Jul 31, 2026
COLUMBIA FUNDS SERIES TRUST I, not summed
|
2,659,635 | ||
| 8.2 |
Columbia Variable Portfolio - Small Cap Value Discovery Fund
Fund
S000012561 · Registered fund · Jun 30, 2026
COLUMBIA FUNDS VARIABLE INSURANCE TRUST, not summed
|
481,660 | ||
| 8.3 |
Columbia Research Enhanced Small Cap ETF
Fund
S000095313 · Registered fund · Jun 30, 2026
Columbia ETF Trust I, not summed
|
144 | ||
| 9 |
Nantahala Capital Management, LLC
13F
CIK 1472322 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
3,623,613 | ||
| 10 |
VANGUARD CAPITAL MANAGEMENT LLC
13F
CIK 2100119 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
3,617,109 | ||
| 11 |
BAILLIE GIFFORD & CO
13G
CIK 1088875 · Investment adviser · Dec 29, 2023
Rule 13d-1(b), qualified institution, unconfirmed, last confirmed Dec 29, 2023
|
3,419,093 | ||
| 12 |
VANGUARD TOTAL STOCK MARKET INDEX FUND
Fund
S000002848 · Registered fund · Jun 30, 2026
VANGUARD INDEX FUNDS, not summed
|
2,633,770 | ||
| 13 |
GEODE CAPITAL MANAGEMENT, LLC
13F
CIK 1214717 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
2,162,393 | ||
| 14 |
|
1,756,814 | ||
| 14.1 |
STATE STREET SMALL/MID CAP EQUITY INDEX PORTFOLIO
Fund
S000050332 · Registered fund · Jun 30, 2026
STATE STREET INSTITUTIONAL INVESTMENT TRUST, not summed
|
33,021 | ||
| 14.2 |
State Street Total Return V.I.S. Fund
Fund
S000001637 · Registered fund · Jun 30, 2026
STATE STREET VARIABLE INSURANCE SERIES FUNDS INC, not summed
|
1,557 | ||
| 14.3 |
State Street Balanced Index Fund
Fund
S000082802 · Registered fund · Jun 30, 2026
STATE STREET INSTITUTIONAL INVESTMENT TRUST, not summed
|
706 | ||
| 15 |
D. E. Shaw & Co., Inc.
13F
CIK 1009207 · Unknown · Jun 30, 2026
carried, not yet filed for the newest quarter
|
1,342,014 | ||
| 16 |
JACOBS LEVY EQUITY MANAGEMENT, INC
13F
CIK 884414 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
1,330,205 | ||
| 17 |
VANGUARD PORTFOLIO MANAGEMENT LLC
13F
CIK 2100121 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
1,307,171 | ||
| 18 |
PERKINS CAPITAL MANAGEMENT INC
13F
CIK 884300 · Registered investment adviser · Sep 30, 2026
|
1,021,850 | ||
| 19 |
VANGUARD EXTENDED MARKET INDEX FUND
Fund
S000002841 · Registered fund · Jun 30, 2026
VANGUARD INDEX FUNDS, not summed
|
895,869 | ||
| 20 |
AQR CAPITAL MANAGEMENT LLC
13F
CIK 1167557 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
845,431 | ||
| 21 |
|
727,053 | ||
| 22 |
TWO SIGMA INVESTMENTS, LP
13F
CIK 1179392 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
696,599 | ||
| 23 |
NORTHERN TRUST CORP
13F
CIK 73124 · Bank · Jun 30, 2026
carried, not yet filed for the newest quarter
|
682,940 | ||
| 23.1 |
Small Cap Index Fund
Fund
S000001245 · Registered fund · Jun 30, 2026
Northern Funds, not summed
|
37,998 | ||
| 23.2 |
Small Cap Core Fund
Fund
S000001244 · Registered fund · Jun 30, 2026
Northern Funds, not summed
|
6,196 | ||
| 24 |
FRANKLIN RESOURCES INC
13F
CIK 38777 · Investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
678,640 | ||
| 24.1 |
Putnam Small Cap Value Fund
Fund
S000003854 · Registered fund · Aug 31, 2026
Putnam Investment Funds, not summed
|
532,703 | ||
| 24.2 |
Putnam VT Small Cap Value Fund
Fund
S000003888 · Registered fund · Jun 30, 2026
Putnam Variable Trust, not summed
|
185,697 | ||
| 25 |
MORGAN STANLEY
13F
CIK 895421 · Holding company · Jun 30, 2026
carried, not yet filed for the newest quarter
|
649,170 | ||
| 26 |
VANGUARD FIDUCIARY TRUST CO
13F
CIK 933478 · Unknown · Jun 30, 2026
carried, not yet filed for the newest quarter
|
581,058 | ||
| 27 |
Post Resch Tallon Group Inc.
13F
CIK 2054476 · Registered investment adviser · Sep 30, 2026
|
573,953 | ||
| 28 |
Man Group plc
13F
CIK 1637460 · Unknown · Jun 30, 2026
carried, not yet filed for the newest quarter
|
565,593 | ||
| 29 |
VANGUARD RUSSELL 2000 INDEX FUND
Fund
S000030003 · Registered fund · Aug 31, 2026
VANGUARD SCOTTSDALE FUNDS, not summed
|
496,765 | ||
| 30 |
FMR LLC
13F
CIK 315066 · Holding company · Jun 30, 2026
carried, not yet filed for the newest quarter
|
478,483 | ||
| 30.1 |
Fidelity Small Cap Index Fund
Fund
S000033638 · Registered fund · Jul 31, 2026
Fidelity Salem Street Trust, not summed
|
942,511 | ||
| 30.2 |
Fidelity Extended Market Index Fund
Fund
S000006021 · Registered fund · Aug 31, 2026
Fidelity Concord Street Trust, not summed
|
438,135 | ||
| 30.3 |
Fidelity Enhanced Small Cap Core ETF
Fund
S000081807 · Registered fund · Jun 30, 2026
Fidelity Covington Trust, not summed
|
148,756 | ||
| 30.4 |
Fidelity Total Market Index Fund
Fund
S000006026 · Registered fund · Aug 31, 2026
Fidelity Concord Street Trust, not summed
|
147,086 | ||
| 30.5 |
Fidelity Series Total Market Index Fund
Fund
S000065248 · Registered fund · Aug 31, 2026
Fidelity Concord Street Trust, not summed
|
120,049 | ||
| 31 |
Assenagon Asset Management S.A.
13F
CIK 1546007 · Unknown · Sep 30, 2026
|
463,055 | ||
| 32 |
GOLDMAN SACHS GROUP INC
13F
CIK 886982 · Holding company · Jun 30, 2026
carried, not yet filed for the newest quarter
|
434,924 | ||
| 32.1 |
Goldman Sachs Absolute Return Tracker Fund
Fund
S000020314 · Registered fund · Aug 31, 2026
GOLDMAN SACHS TRUST, not summed
|
77,025 | ||
| 33 |
ALGERT GLOBAL LLC
13F
CIK 1275218 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
430,910 | ||
| 34 |
Prescott Group Capital Management, L.L.C.
13F
CIK 1166152 · Registered investment adviser · Sep 30, 2026
|
400,000 | ||
| 35 |
SMALL COMPANY GROWTH PORTFOLIO
Fund
S000004394 · Registered fund · Jun 30, 2026
VANGUARD VARIABLE INSURANCE FUNDS, not summed
|
398,987 | ||
| 36 |
|
392,267 | ||
| 36.1 |
Nuveen Equity Index Fund
Fund
S000005360 · Registered fund · Jul 31, 2026
TIAA-CREF Funds, not summed
|
183,067 | ||
| 36.2 |
Nuveen Small Cap Blend Index Fund
Fund
S000005367 · Registered fund · Jul 31, 2026
TIAA-CREF Funds, not summed
|
105,313 | ||
| 36.3 |
College Retirement Equities Fund - Total Global Stock Account
Fund
S000005080 · Registered fund · Jun 30, 2026
College Retirement Equities Fund, not summed
|
76,850 | ||
| 36.4 |
Nuveen Life Stock Index Fund
Fund
S000005094 · Registered fund · Jun 30, 2026
TIAA-CREF Life Funds, not summed
|
5,038 | ||
| 36.5 |
TIAA Separate Account VA-1 - Stock Index Account
Fund
S000005098 · Registered fund · Jun 30, 2026
TIAA Separate Account VA-1, not summed
|
1,647 | ||
| 37 |
ACADIAN ASSET MANAGEMENT LLC
13F
CIK 916542 · Unknown · Jun 30, 2026
carried, not yet filed for the newest quarter
|
358,285 | ||
| 38 |
TUDOR INVESTMENT CORP ET AL
13F
CIK 923093 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
357,925 | ||
| 39 |
VANGUARD EXPLORER FUND
Fund
S000002578 · Registered fund · Jul 31, 2026
VANGUARD EXPLORER FUND, not summed
|
316,606 | ||
| 40 |
Russell Investments Group, Ltd.
13F
CIK 1692234 · Unknown · Jun 30, 2026
carried, not yet filed for the newest quarter
|
308,095 | ||
| 40.1 |
U.S. Small Cap Equity Fund
Fund
S000001586 · Registered fund · Jul 31, 2026
RUSSELL INVESTMENT CO, not summed
|
16,975 | ||
| 40.2 |
U.S. Small Cap Equity Fund
Fund
S000006864 · Registered fund · Jun 30, 2026
RUSSELL INVESTMENT FUNDS, not summed
|
3,534 | ||
| 41 |
PLATINUM INVESTMENT MANAGEMENT LTD
13F
CIK 1256071 · Unknown · Jun 30, 2026
carried, not yet filed for the newest quarter
|
264,464 | ||
| 42 |
ROYAL BANK OF CANADA
13F
CIK 1000275 · Bank · Jun 30, 2026
carried, not yet filed for the newest quarter
|
262,926 | ||
| 43 |
Potomac Capital Management, Inc.
13F
CIK 1418746 · Unknown · Jun 30, 2026
carried, not yet filed for the newest quarter
|
260,000 | ||
| 44 |
|
244,171 | ||
| 45 |
CHARLES SCHWAB INVESTMENT MANAGEMENT INC
13F
CIK 884546 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
242,211 | ||
| 45.1 |
Schwab Small-Cap Index Fund
Fund
S000005912 · Registered fund · Jul 31, 2026
SCHWAB CAPITAL TRUST, not summed
|
196,089 | ||
| 45.2 |
Schwab Total Stock Market Index Fund
Fund
S000005913 · Registered fund · Jul 31, 2026
SCHWAB CAPITAL TRUST, not summed
|
45,863 | ||
| 46 |
XTX Topco Ltd
13F
CIK 1828301 · Unknown · Jun 30, 2026
carried, not yet filed for the newest quarter
|
240,627 | ||
| 47 |
BANK OF AMERICA CORP /DE/
13F
CIK 70858 · Bank · Jun 30, 2026
carried, not yet filed for the newest quarter
|
240,517 | ||
| 48 |
|
239,907 | ||
| 49 |
Bank of New York Mellon Corp
13F
CIK 1390777 · Bank · Jun 30, 2026
carried, not yet filed for the newest quarter
|
226,039 | ||
| 50 |
Large-Cap Growth Portfolio
Fund
S000026003 · Registered fund · Jun 30, 2026
PACIFIC SELECT FUND, not summed
|
213,439 |
16 more nested rows are not shown here; Every filing lists them all.
Shares are restated onto today's split basis. Change compares each row with the same holder's previous filing in the same source; a dash means there is none to compare. Nested 13D/G, insider and fund rows are not summed, since a 13F report already holds them.
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