CDXS · Codexis, Inc. · Holdings

$1.36 +0.04 (+3.03%) At close · Sep 18

Russell 2000 member Rebalance outlook locked

Market Cap

$150.57M

109.51M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Use institutional holdings through the 13F API or MCP →

Quarter:

Holders: 148

Total Value: $146,683,445

Total Shares: 64,904,222

Showing 1–100 of 150 holdings by value · page 1 of 2

Institutional Ownership Trend

26 quarters

Superinvestors Holding CDXS

All superinvestors →

Mario Gabelli $22,600

Holder Concentration

Across 148 13F filers · Δ vs Mar 31, 2026

Top Holder

11.2%

−1.8 pts

Top 5

46.6%

+0.5 pts

Top 10

69.6%

+1.1 pts

HHI

598

Broadly held−3

Quarterly Activity

vs Mar 31, 2026

New

Increased

Decreased

Sold Out

Unchanged

Holder count: +13

Institutional value: +35.3%

Shares held: -2.4%

Top Buyers

vs Mar 31, 2026

Institution	Added	Now Holds
Mega MILLENNIUM MANAGEMENT LLC	+1,551,009	5,549,139
Large Assenagon Asset Management S.A.	+527,036	1,278,206
Mega TWO SIGMA INVESTMENTS, LP	+454,644	696,599
Mega VANGUARD PORTFOLIO MANAGEMENT LLC	+427,832	1,307,171
Mega AMERIPRISE FINANCIAL INC	+408,944	3,809,440

Top Sellers

vs Mar 31, 2026

Institution	Sold	Now Holds
Mega UBS Group AG	−1,943,868	239,907
Large Aberdeen Group plc	−1,910,414	6,769,954
Mega JANE STREET GROUP, LLC	−799,149	120,815
Mid Nantahala Capital Management, LLC	−500,000	3,623,613
Mega GOLDMAN SACHS GROUP INC	−389,946	434,924

Top New Positions

vs Mar 31, 2026

Institution	Acquired	Value (USD)
Large Man Group plc	565,593	$1,278,240
Mega ARROWSTREET CAPITAL, LIMITED PARTNERSHIP	148,051	$334,595
Large MARSHALL WACE, LLP	117,135	$264,724
Mid Numerai GP LLC	60,209	$136,072
Large PANAGORA ASSET MANAGEMENT INC	58,200	$131,532

Top Sold-Out Positions

vs Mar 31, 2026

Institution	Previously Held	Last Value (USD)
Small Timothy G. Youngquist 2020 Irrevocable Trust	978,071	$1,594,255
Mid Private Advisor Group, LLC	847,001	$1,380,611
Small Diametric Capital, LP	359,268	$585,606
Large LAZARD ASSET MANAGEMENT LLC	282,380	$460,279
Mid PRELUDE CAPITAL MANAGEMENT, LLC	54,680	$89,128

Filter: New 26 Increased 62 Reduced 36 Sold out 13 Unchanged 24

Showing 100 of 150

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

BlackRock, Inc.

$16,462,631

11.22%

7,284,352

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	287,027	$648,681	3.9%	Sole
BlackRock Asset Management Canada Ltd	860	$1,943	0.0%	Sole
BlackRock Investment Management, LLC	252,470	$570,582	3.5%	Sole
BLACKROCK ADVISORS LLC	162,893	$368,138	2.2%	Sole
BlackRock Fund Advisors	4,049,932	$9,152,846	55.6%	Sole
BlackRock Institutional Trust Company, N.A.	2,421,785	$5,473,234	33.2%	Sole
BlackRock Japan Co. Ltd	8,403	$18,990	0.1%	Sole
BlackRock Investment Management (UK) Ltd.	26,627	$60,177	0.4%	Sole
BlackRock (Netherlands) B.V.	3,046	$6,883	0.0%	Sole
BlackRock Asset Management Ireland Ltd	70,565	$159,476	1.0%	Sole
Aperio Group, LLC	744	$1,681	0.0%	Sole

Opaleye Management Inc.

$15,334,100

10.45%

6,785,000

Aberdeen Group plc

$15,300,096

10.43%

6,769,954

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
abrdn Inc.	6,769,954	$15,300,096	100.0%	Defined

MILLENNIUM MANAGEMENT LLC

$12,541,054

8.55%

5,549,139

Telemark Asset Management, LLC

$8,692,888

5.93%

3,846,411

AMERIPRISE FINANCIAL INC

$8,609,334

5.87%

3,809,440

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	3,809,440	$8,609,334	100.0%	Defined

Nantahala Capital Management, LLC

$8,189,365

5.58%

3,623,613

VANGUARD CAPITAL MANAGEMENT LLC

$8,174,666

5.57%

3,617,109

GEODE CAPITAL MANAGEMENT, LLC

$4,887,007

3.33%

2,162,393

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	302,253	$683,091	14.0%	Defined
Held directly	1,860,140	$4,203,916	86.0%	Defined

STATE STREET CORP

$3,970,399

2.71%

1,756,814

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	1,756,814	$3,970,399	100.0%	Defined

D. E. Shaw & Co., Inc.

$3,032,950

2.07%

1,342,014

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	1,106,296	$2,500,228	82.4%	Defined
D. E. Shaw Investment Management, L.L.C. joint	235,718	$532,722	17.6%	Defined

JACOBS LEVY EQUITY MANAGEMENT, INC

$3,006,263

2.05%

1,330,205

VANGUARD PORTFOLIO MANAGEMENT LLC

$2,954,206

2.01%

1,307,171

Assenagon Asset Management S.A.

$2,888,745

1.97%

1,278,206

AQR CAPITAL MANAGEMENT LLC

$1,910,670

1.30%

845,431

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RF&L WEALTH MANAGEMENT, LLC	41	$92	0.0%	Other
Prism Planning Partners LLC	569	$1,285	0.1%	Other
Compound Planning, Inc.	54	$122	0.0%	Other
Arax Advisory Partners	11,402	$25,768	1.3%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	4,667	$10,547	0.6%	Other
Stokes Family Office, LLC	3,560	$8,045	0.4%	Other
Held directly	825,138	$1,864,811	97.6%	Defined

TWO SIGMA INVESTMENTS, LP

$1,574,313

1.07%

696,599

NORTHERN TRUST CORP

$1,543,444

1.05%

682,940

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	670,554	$1,515,452	98.2%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	12,386	$27,992	1.8%	Other

FRANKLIN RESOURCES INC

$1,533,726

1.05%

678,640

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PUTNAM INVESTMENT MANAGEMENT LLC	678,640	$1,533,726	100.0%	Defined

MORGAN STANLEY

$1,467,121

1.00%

649,170

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO. LLC	23,651	$53,451	3.6%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	245,692	$555,263	37.8%	Defined
MORGAN STANLEY SMITH BARNEY LLC	34,275	$77,461	5.3%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	345,552	$780,946	53.2%	Defined

VANGUARD FIDUCIARY TRUST CO

$1,313,191

0.90%

581,058

Man Group plc

$1,278,240

0.87%

565,593

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Numeric Investors LLC	565,593	$1,278,240	100.0%	Defined

Post Resch Tallon Group Inc.

$1,273,460

0.87%

563,478

PERKINS CAPITAL MANAGEMENT INC

$1,188,986

0.81%

526,100

FMR LLC

$1,081,370

0.74%

478,483

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	202,434	$457,500	42.3%	Defined
STRATEGIC ADVISERS LLC	1,951	$4,409	0.4%	Defined
FIAM LLC	274,098	$619,461	57.3%	Defined

GOLDMAN SACHS GROUP INC

$982,926

0.67%

434,924

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	172,445	$389,725	39.6%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	251,369	$568,093	57.8%	Defined
Goldman Sachs Trust Company, N.A.	11,110	$25,108	2.6%	Defined

ALGERT GLOBAL LLC

$973,856

0.66%

430,910

Nuveen, LLC

$886,521

0.60%

392,267

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	20,352	$45,994	5.2%	Defined
TIAA CREF INVESTMENT MANAGEMENT LLC joint	76,850	$173,681	19.6%	Defined
TEACHERS ADVISORS, LLC joint	295,065	$666,846	75.2%	Defined

ACADIAN ASSET MANAGEMENT LLC

$809,724

0.55%

358,285

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Acadian Inc	358,285	$809,724	100.0%	Defined

TUDOR INVESTMENT CORP ET AL

$808,910

0.55%

357,925

DIMENSIONAL FUND ADVISORS LP

$708,251

0.48%

313,386

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	6,002	$13,564	1.9%	Defined
Dimensional Ireland Ltd	49,121	$111,013	15.7%	Defined
Held directly	258,263	$583,674	82.4%	Sole

Russell Investments Group, Ltd.

$696,291

0.47%

308,095

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC joint	53,005	$119,790	17.2%	Defined
Russell Investments Capital, LLC joint	237,284	$536,260	77.0%	Other
Russell Investments Trust Company joint	17,806	$40,241	5.8%	Other

PLATINUM INVESTMENT MANAGEMENT LTD

$597,688

0.41%

264,464

ROYAL BANK OF CANADA

$594,212

0.41%

262,926

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC CMA LTD.	2,612	$5,903	1.0%	Defined
RBC Capital Markets, LLC	260,313	$588,307	99.0%	Defined
Held directly	1	$2	0.0%	Sole

Potomac Capital Management, Inc.

$587,600

0.40%

260,000

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$547,396

0.37%

242,211

XTX Topco Ltd

$543,817

0.37%

240,627

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	240,627	$543,817	100.0%	Defined

BANK OF AMERICA CORP /DE/

$543,568

0.37%

240,517

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	64,417	$145,582	26.8%	Defined
BOFA SECURITIES, INC.	176,100	$397,986	73.2%	Defined

UBS Group AG

$542,189

0.37%

239,907

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS AG	239,906	$542,187	100.0%	Defined
UBS Switzerland AG	1	$2	0.0%	Defined

Bank of New York Mellon Corp

$510,846

0.35%

226,039

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Investments Corporation	77,070	$174,178	34.1%	Defined
The Bank of New York Mellon	143,334	$323,934	63.4%	Defined
BNY Mellon Advisors, Inc. joint	5,634	$12,732	2.5%	Other
Held directly	1	$2	0.0%	Defined

CITADEL ADVISORS LLC

$432,969

0.30%

191,580

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	91,007	$205,675	47.5%	Defined
Held directly	100,573	$227,294	52.5%	Defined

Invesco Ltd.

$366,731

0.25%

162,271

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	13,793	$31,171	8.5%	Defined
Invesco Trust Co joint	34,151	$77,181	21.0%	Defined
Invesco Capital Management LLC	114,327	$258,379	70.5%	Defined

BARCLAYS PLC

$350,918

0.24%

155,274

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	153,867	$347,739	99.1%	Defined
BARCLAYS CAPITAL INC.	1,407	$3,179	0.9%	Defined

ARROWSTREET CAPITAL, LIMITED PARTNERSHIP

$334,595

0.23%

148,051

ALLIANCEBERNSTEIN L.P.

$285,731

0.19%

126,430

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	126,430	$285,731	100.0%	Defined

BRIDGEWAY CAPITAL MANAGEMENT, LLC

$284,552

0.19%

125,908

STRS OHIO

$273,460

0.19%

121,000

JANE STREET GROUP, LLC

$273,041

0.19%

120,815

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	115,866	$261,857	95.9%	Defined
Jane Street Global Trading, LLC	4,949	$11,184	4.1%	Defined

MARSHALL WACE, LLP

$264,724

0.18%

117,135

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marshall Wace North America L.P. joint	117,135	$264,724	100.0%	Other

Stonepine Capital Management, LLC

$247,836

0.17%

109,662

RHUMBLINE ADVISERS

$233,460

0.16%

103,301

WELLS FARGO & COMPANY/MN

$230,352

0.16%

101,926

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	77,926	$176,112	76.5%	Defined
WELLS FARGO CLEARING SERVICES, LLC	24,000	$54,240	23.5%	Defined

DEUTSCHE BANK AG\

$217,477

0.15%

96,229

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DWS INVESTMENT MANAGEMENT AMERICAS, INC	1,647	$3,722	1.7%	Defined
DWS Investments UK Ltd	94,582	$213,755	98.3%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$186,501

0.13%

82,523

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	10,895	$24,622	13.2%	Other
G1 EXECUTION SERVICES, LLC	71,628	$161,879	86.8%	Other

LPL Financial LLC

$173,025

0.12%

76,560

PRICE T ROWE ASSOCIATES INC /MD/

$151,248

0.10%

66,924

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	66,924	$151,248	100.0%	Defined

PUBLIC EMPLOYEES RETIREMENT SYSTEM OF OHIO

$144,081

0.10%

63,753

Numerai GP LLC

$136,072

0.09%

60,209

PANAGORA ASSET MANAGEMENT INC

$131,532

0.09%

58,200

Mariner, LLC

$123,479

0.08%

54,637

Cerity Partners LLC

$120,507

0.08%

53,322

MetLife Investment Management, LLC

$117,544

0.08%

52,011

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
METLIFE INC	52,011	$117,544	100.0%	Defined

ENVESTNET ASSET MANAGEMENT INC

$116,107

0.08%

51,375

LOS ANGELES CAPITAL MANAGEMENT LLC

$108,762

0.07%

48,125

Virtu Financial LLC

$105,318

0.07%

46,601

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Virtu Americas LLC	46,601	$105,318	100.0%	Defined

IEQ CAPITAL, LLC

$104,558

0.07%

46,265

CITIGROUP INC

$97,931

0.07%

43,333

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	43,333	$97,931	100.0%	Defined

JPMORGAN CHASE & CO

$94,605

0.06%

41,861

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	35,832	$80,980	85.6%	Defined
J.P. Morgan Securities plc	6,029	$13,625	14.4%	Defined

GSA CAPITAL PARTNERS LLP

$94,492

0.06%

41,811

RENAISSANCE TECHNOLOGIES LLC

$93,790

0.06%

41,500

MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.

$89,957

0.06%

39,804

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Global X Management CO LLC joint	39,804	$89,957	100.0%	Defined

Frec Markets, Inc.

$84,227

0.06%

37,269

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Frec Advisers LLC	37,269	$84,227	100.0%	Defined

OSAIC HOLDINGS, INC.

$82,187

0.06%

36,366

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
OSAIC WEALTH, INC.	36,366	$82,187	100.0%	Defined

COMMONWEALTH EQUITY SERVICES, LLC

$81,821

0.06%

36,204

BANK OF MONTREAL /CAN/

$79,095

0.05%

34,998

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BMO Family Office, LLC	34,998	$79,095	100.0%	Defined

Krilogy Financial LLC

$78,704

0.05%

34,825

Dynamic Technology Lab Private Ltd

$74,580

0.05%

33,000

Vanguard Global Advisers, LLC

$73,680

0.05%

32,602

MERCER GLOBAL ADVISORS INC /ADV

$73,111

0.05%

32,350

AMERICAN CENTURY COMPANIES INC

$68,932

0.05%

30,501

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AMERICAN CENTURY INVESTMENT MANAGEMENT INC	30,501	$68,932	100.0%	Defined

BLAIR WILLIAM & CO/IL

$68,294

0.05%

30,219

EVERGREEN CAPITAL MANAGEMENT LLC

$65,540

0.04%

29,000

Capital Advisors, Ltd. LLC

$65,291

0.04%

28,890

PRUDENTIAL FINANCIAL INC

$59,618

0.04%

26,380

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PGIM Quantitative Solutions LLC	26,380	$59,618	100.0%	Defined

GUGGENHEIM CAPITAL LLC

$59,116

0.04%

26,158

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GUGGENHEIM FUNDS DISTRIBUTORS, LLC	24,620	$55,641	94.1%	Defined
SECURITY INVESTORS LLC	832	$1,880	3.2%	Defined
Guggenheim Partners Investment Management, LLC joint	706	$1,595	2.7%	Defined

WEALTH ENHANCEMENT ADVISORY SERVICES, LLC

$57,121

0.04%

25,275

MANUFACTURERS LIFE INSURANCE COMPANY, THE

$56,963

0.04%

25,205

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MANULIFE FINANCIAL CORP joint	25,205	$56,963	100.0%	Defined

INTECH INVESTMENT MANAGEMENT LLC

$48,486

0.03%

21,454

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$45,426

0.03%

20,100

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	20,100	$45,426	100.0%	Other

Police & Firemen's Retirement System of New Jersey

$43,936

0.03%

19,441

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Capital	19,441	$43,936	100.0%	Defined

VOYA INVESTMENT MANAGEMENT LLC

$41,947

0.03%

18,561

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	18,561	$41,947	100.0%	Defined

Vanguard Personalized Indexing Management, LLC

$40,562

0.03%

17,948

Simplicity Wealth,LLC

$39,409

0.03%

17,438

NEW YORK STATE COMMON RETIREMENT FUND

$38,166

0.03%

16,888

Corient Private Wealth LLC

$37,827

0.03%

16,738

CITADEL ADVISORS LLC

$35,708

0.02%

15,800

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	15,800	$35,708	100.0%	Defined

Creative Planning

$35,002

0.02%

15,488

BNP PARIBAS ARBITRAGE, SA

$34,941

0.02%

15,461

Navalign, LLC

$34,668

0.02%

15,340

EP Wealth Advisors, LLC

$34,526

0.02%

15,277

China Universal Asset Management Co., Ltd.

$32,693

0.02%

14,466

Key facts CIK 1200375 CUSIP 192005106 13F (30d) 5 filings 5 filers Visit website Investor relations