CDXS · Codexis, Inc.

$1.45 +0.02 (+1.40%) At close · Aug 5

Market Cap

$131.35M

90.90M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

The Jun 30, 2026 filing window is still open — showing the latest known positions. Funds that have not filed yet carry their prior-quarter positions; a fund that filed without this stock counts as sold out. Pick a completed quarter in the Quarter selector below to see a single quarter with position changes.

Quarter:

Holders: 134

Total Value: $107,057,335

Total Shares: 62,463,925

Showing 1–100 of 135 holdings by value · page 1 of 2

Institutional Ownership Trend

26 quarters

Superinvestors Holding CDXS

All superinvestors →

Mario Gabelli $16,300

Holder Concentration

Across 133 13F filers

Top Holder

13.5%

of institutional value

Top 5

47.8%

of institutional value

Top 10

71.2%

of institutional value

HHI

642

Broadly held

Showing 100 of 135

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

Large Aberdeen Group plc

$15,300,096

14.29%

6,769,954

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
abrdn Inc.	6,769,954	$15,300,096	100.0%	Defined

Mega BlackRock, Inc.

$11,456,895

10.70%

7,028,773

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	285,796	$465,847	4.1%	Sole
BlackRock Asset Management Canada Ltd	860	$1,401	0.0%	Sole
BlackRock Investment Management, LLC	259,559	$423,081	3.7%	Sole
BLACKROCK ADVISORS LLC	162,893	$265,515	2.3%	Sole
BlackRock Fund Advisors	3,866,596	$6,302,551	55.0%	Sole
BlackRock Institutional Trust Company, N.A.	2,380,193	$3,879,714	33.9%	Sole
BlackRock (Netherlands) B.V.	3,046	$4,964	0.0%	Sole
BlackRock Asset Management Ireland Ltd	69,086	$112,610	1.0%	Sole
Aperio Group, LLC	744	$1,212	0.0%	Sole

Small Opaleye Management Inc.

$11,059,550

10.33%

6,785,000

Mid Nantahala Capital Management, LLC

$6,721,489

6.28%

4,123,613

Mega MILLENNIUM MANAGEMENT LLC

$6,516,951

6.09%

3,998,130

Mid Telemark Asset Management, LLC

$6,269,649

5.86%

3,846,411

Mega VANGUARD CAPITAL MANAGEMENT LLC

$5,979,666

5.59%

3,668,507

Mega AMERIPRISE FINANCIAL INC

$5,542,808

5.18%

3,400,496

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	3,400,496	$5,542,808	100.0%	Defined

Mega UBS Group AG

$3,559,552

3.32%

2,183,775

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS AG	2,183,774	$3,559,551	100.0%	Defined
UBS Switzerland AG	1	$1	0.0%	Defined

Mega GEODE CAPITAL MANAGEMENT, LLC

$3,073,124

2.87%

1,885,353

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	290,566	$473,622	15.4%	Defined
Held directly	1,594,787	$2,599,502	84.6%	Defined

Large Assenagon Asset Management S.A.

$2,888,745

2.70%

1,278,206

Mega STATE STREET CORP

$2,717,135

2.54%

1,666,954

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	1,666,954	$2,717,135	100.0%	Defined

Large JACOBS LEVY EQUITY MANAGEMENT, INC

$1,773,484

1.66%

1,088,027

Mega D. E. Shaw & Co., Inc.

$1,597,150

1.49%

979,847

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	689,631	$1,124,098	70.4%	Defined
D. E. Shaw Investment Management, L.L.C. joint	290,216	$473,052	29.6%	Defined

Mega JANE STREET GROUP, LLC

$1,499,541

1.40%

919,964

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	760,689	$1,239,923	82.7%	Defined
Jane Street Global Trading, LLC	159,275	$259,618	17.3%	Defined

Mega GOLDMAN SACHS GROUP INC

$1,344,537

1.26%

824,870

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	678,680	$1,106,248	82.3%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	135,080	$220,180	16.4%	Defined
Goldman Sachs Trust Company, N.A.	11,110	$18,109	1.3%	Defined

Micro Post Resch Tallon Group Inc.

$1,273,460

1.19%

563,478

Small PERKINS CAPITAL MANAGEMENT INC

$1,188,986

1.11%

526,100

Mega FRANKLIN RESOURCES INC

$1,121,782

1.05%

688,210

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PUTNAM INVESTMENT MANAGEMENT LLC	688,210	$1,121,782	100.0%	Defined

Mega NORTHERN TRUST CORP Bank

$1,071,136

1.00%

657,139

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	644,623	$1,050,735	98.1%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	12,516	$20,401	1.9%	Other

Mega AQR CAPITAL MANAGEMENT LLC

$992,014

0.93%

608,601

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RF&L WEALTH MANAGEMENT, LLC	3	$4	0.0%	Other
Prism Planning Partners LLC	430	$700	0.1%	Other
Compound Planning, Inc.	28	$45	0.0%	Other
Arax Advisory Partners	17,414	$28,384	2.9%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	3,625	$5,908	0.6%	Other
Stokes Family Office, LLC	2,665	$4,343	0.4%	Other
Held directly	584,436	$952,630	96.0%	Defined

Mega VANGUARD FIDUCIARY TRUST CO Bank

$898,143

0.84%

551,008

Large TUDOR INVESTMENT CORP ET AL

$876,667

0.82%

537,833

Mega MORGAN STANLEY

$770,728

0.72%

472,841

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	77	$125	0.0%	Defined
MORGAN STANLEY & CO. LLC	20,980	$34,197	4.4%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	76,575	$124,817	16.2%	Defined
MORGAN STANLEY SMITH BARNEY LLC	34,275	$55,868	7.2%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	340,934	$555,721	72.1%	Defined

Small Stonepine Capital Management, LLC

$681,330

0.64%

417,994

Mega FMR LLC

$672,532

0.63%

412,597

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	148,348	$241,807	36.0%	Defined
STRATEGIC ADVISERS LLC	1,951	$3,180	0.5%	Defined
FIAM LLC	262,298	$427,545	63.6%	Defined

Mega Nuveen, LLC

$598,930

0.56%

367,443

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	8,883	$14,479	2.4%	Defined
TIAA CREF INVESTMENT MANAGEMENT LLC joint	63,495	$103,496	17.3%	Defined
TEACHERS ADVISORS, LLC joint	295,065	$480,955	80.3%	Defined

Small Diametric Capital, LP

$585,606

0.55%

359,268

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Thakore Nainoor	359,268	$585,606	100.0%	Defined

Mega CITADEL ADVISORS LLC

$468,779

0.44%

287,595

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	218,325	$355,869	75.9%	Defined
Held directly	69,270	$112,910	24.1%	Defined

Mega DIMENSIONAL FUND ADVISORS LP

$463,956

0.43%

284,636

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	6,002	$9,783	2.1%	Defined
Dimensional Ireland Ltd	49,121	$80,067	17.3%	Defined
Held directly	229,513	$374,106	80.6%	Sole

Large LAZARD ASSET MANAGEMENT LLC

$460,279

0.43%

282,380

Small Potomac Capital Management, Inc.

$446,946

0.42%

274,200

Small PLATINUM INVESTMENT MANAGEMENT LTD

$431,076

0.40%

264,464

Mega CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$394,803

0.37%

242,211

Mega TWO SIGMA INVESTMENTS, LP

$394,386

0.37%

241,955

Mega ROYAL BANK OF CANADA Bank

$351,044

0.33%

215,365

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC Capital Markets, LLC	215,301	$350,940	100.0%	Defined
Held directly	64	$104	0.0%	Sole

Mega Bank of New York Mellon Corp Bank

$341,720

0.32%

209,645

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Investments Corporation	77,070	$125,624	36.8%	Defined
The Bank of New York Mellon	128,722	$209,816	61.4%	Defined
BNY Mellon Advisors, Inc. joint	3,851	$6,277	1.8%	Other
Held directly	2	$3	0.0%	Defined

Large RENAISSANCE TECHNOLOGIES LLC

$323,934

0.30%

198,733

Mega Invesco Ltd.

$294,906

0.28%

180,925

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	14,696	$23,954	8.1%	Defined
Invesco Trust Co joint	11,596	$18,901	6.4%	Defined
Invesco Capital Management LLC	154,633	$252,051	85.5%	Defined

Mega BANK OF AMERICA CORP /DE/ Bank

$283,546

0.26%

173,955

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	92,670	$151,052	53.3%	Defined
BOFA SECURITIES, INC.	81,285	$132,494	46.7%	Defined

Large STRS OHIO

$273,460

0.26%

121,000

Mid XTX Topco Ltd

$273,093

0.26%

167,542

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	167,542	$273,093	100.0%	Defined

Mega BARCLAYS PLC

$252,801

0.24%

155,093

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	154,753	$252,247	99.8%	Defined
BARCLAYS CAPITAL INC.	340	$554	0.2%	Defined

Large LOS ANGELES CAPITAL MANAGEMENT LLC

$217,873

0.20%

133,665

Mid BRIDGEWAY CAPITAL MANAGEMENT, LLC

$205,230

0.19%

125,908

Mega ALLIANCEBERNSTEIN L.P.

$192,551

0.18%

118,130

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	118,130	$192,551	100.0%	Defined

Mega RHUMBLINE ADVISERS

$161,881

0.15%

99,314

Mega DEUTSCHE BANK AG\ Bank

$151,723

0.14%

93,082

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DWS INVESTMENT MANAGEMENT AMERICAS, INC	1,647	$2,684	1.8%	Defined
DWS Investments UK Ltd	91,435	$149,039	98.2%	Defined

Mega LPL Financial LLC

$122,347

0.11%

75,060

Mega ENVESTNET ASSET MANAGEMENT INC

$116,107

0.11%

51,375

Mid Quadrature Capital Ltd

$104,644

0.10%

64,199

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Quadrature US Inc.	64,199	$104,644	100.0%	Other

Large PUBLIC EMPLOYEES RETIREMENT SYSTEM OF OHIO Pension

$103,917

0.10%

63,753

Mega CITIGROUP INC

$97,931

0.09%

43,333

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	43,333	$97,931	100.0%	Defined

Large Russell Investments Group, Ltd.

$95,320

0.09%

58,479

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jacobs Levy Equity Management, Inc. joint	48,636	$79,276	83.2%	Other
Russell Investment Management, LLC	9,843	$16,044	16.8%	Defined

Mega WELLS FARGO & COMPANY/MN

$92,308

0.09%

56,631

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	32,631	$53,188	57.6%	Defined
WELLS FARGO CLEARING SERVICES, LLC	24,000	$39,120	42.4%	Defined

Mid PRELUDE CAPITAL MANAGEMENT, LLC

$89,128

0.08%

54,680

Mega SUSQUEHANNA INTERNATIONAL GROUP, LLP

$86,782

0.08%

53,241

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	53,241	$86,782	100.0%	Other

Large COMMONWEALTH EQUITY SERVICES, LLC

$81,821

0.08%

36,204

Mid Krilogy Financial LLC

$78,704

0.07%

34,825

Large MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd. Mutual Fund

$69,141

0.06%

42,418

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Global X Management CO LLC joint	42,418	$69,141	100.0%	Defined

Mega JPMORGAN CHASE & CO

$67,492

0.06%

41,407

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	36,251	$59,089	87.5%	Defined
J.P. Morgan Securities LLC	17	$27	0.0%	Defined
J.P. Morgan Securities plc	5,139	$8,376	12.4%	Defined

Mid EVERGREEN CAPITAL MANAGEMENT LLC

$65,540

0.06%

29,000

Small Capital Advisors, Ltd. LLC

$65,291

0.06%

28,890

Large CWM, LLC

$63,695

0.06%

39,077

Large OSAIC HOLDINGS, INC.

$59,600

0.06%

36,565

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
OSAIC WEALTH, INC.	36,565	$59,600	100.0%	Defined

Small Frec Markets, Inc.

$59,092

0.06%

36,253

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Frec Advisers LLC	36,253	$59,092	100.0%	Defined

Large WEALTH ENHANCEMENT ADVISORY SERVICES, LLC

$57,121

0.05%

25,275

Mega BANK OF MONTREAL /CAN/ Bank

$57,046

0.05%

34,998

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BMO Family Office, LLC	34,998	$57,046	100.0%	Defined

Large Cerity Partners LLC

$53,612

0.05%

32,892

Mega Vanguard Global Advisers, LLC

$53,141

0.05%

32,602

Large BLAIR WILLIAM & CO/IL

$48,929

0.05%

30,018

Mega CITADEL ADVISORS LLC

$46,781

0.04%

28,700

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	28,700	$46,781	100.0%	Defined

Mega MANUFACTURERS LIFE INSURANCE COMPANY, THE Insurance

$44,725

0.04%

27,439

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MANULIFE FINANCIAL CORP joint	27,439	$44,725	100.0%	Defined

Large Police & Firemen's Retirement System of New Jersey Pension

$43,936

0.04%

19,441

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Capital	19,441	$43,936	100.0%	Defined

Large GUGGENHEIM CAPITAL LLC

$43,926

0.04%

26,950

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GUGGENHEIM FUNDS DISTRIBUTORS, LLC	25,498	$41,561	94.6%	Defined
SECURITY INVESTORS LLC	746	$1,215	2.8%	Defined
Guggenheim Partners Investment Management, LLC joint	706	$1,150	2.6%	Defined

Mid INTECH INVESTMENT MANAGEMENT LLC

$39,522

0.04%

24,247

Large Mariner, LLC

$38,352

0.04%

23,529

Large NEW YORK STATE COMMON RETIREMENT FUND Pension

$38,166

0.04%

16,888

Small Navalign, LLC

$34,668

0.03%

15,340

Mega BNP PARIBAS ARBITRAGE, SA

$33,726

0.03%

20,691

Mid China Universal Asset Management Co., Ltd.

$32,693

0.03%

14,466

Mid Vanguard Personalized Indexing Management, LLC

$29,638

0.03%

18,183

Large MERCER GLOBAL ADVISORS INC /ADV

$28,816

0.03%

17,679

Small Elmwood Wealth Management, Inc.

$28,690

0.03%

12,695

Large ProShare Advisors LLC

$27,670

0.03%

16,976

Small Abel Hall, LLC

$26,930

0.03%

11,916

Large VOYA INVESTMENT MANAGEMENT LLC

$24,150

0.02%

14,816

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	14,816	$24,150	100.0%	Defined

Small StoneX Group Inc.

$22,976

0.02%

14,096

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STONEX FINANCIAL INC.	14,096	$22,976	100.0%	Defined

Mid Fiduciary Trust Co Bank

$22,200

0.02%

13,620

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fiduciary Trust Co of New England	13,620	$22,200	100.0%	Defined

Large PRUDENTIAL FINANCIAL INC

$22,135

0.02%

13,580

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PGIM Quantitative Solutions LLC	13,580	$22,135	100.0%	Defined

Mid Tower Research Capital LLC (TRC)

$22,122

0.02%

13,572

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Latour Trading LLC (Latour)	13,572	$22,122	100.0%	Defined

Small Simplicity Wealth,LLC

$21,692

0.02%

13,308

Small Caerus Investment Advisors, LLC

$17,245

0.02%

10,580

Mid Pallas Capital Advisors LLC

$16,829

0.02%

10,325

Mega VICTORY CAPITAL MANAGEMENT INC

$16,432

0.02%

10,081

Mid GAMCO INVESTORS, INC. ET AL

$16,300

0.02%

10,000

Mid Ameritas Investment Partners, Inc.

$13,737

0.01%

8,428

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
EATON VANCE MANAGEMENT	8,428	$13,737	100.0%	Other

Small Pinnacle Wealth Planning Services, Inc.

$13,484

0.01%

8,273

Small Essential Partners LLC

$13,020

0.01%

7,988

Mega Legal & General Group Plc

$11,641

0.01%

7,142

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	7,142	$11,641	100.0%	Defined

Key facts CIK 1200375 CUSIP 192005106 13F (30d) 33 filings 32 filers Visit website Investor relations