NTRA · Natera, Inc. · Holdings

$395.56 -5.09 (-1.27%) At close · Oct 7

Russell 1000 member Rebalance outlook locked

Market Cap

$57.02B

144.14M

Volume · Oct 7 2.82M Avg daily vol (3M) 1.44M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Use institutional holdings through the 13F API or MCP →

Some SEC filings for this period have not finished importing. Holder totals and changes may be incomplete; these filings remain queued for recovery.

The Sep 30, 2026 filing window is still open — showing the latest known positions. Funds that have not filed yet carry their prior-quarter positions; a fund that filed without this stock counts as sold out. Each fund's Change is measured against its own previous reported quarter, shown in the Reported column. Pick a completed quarter in the Quarter selector below to put every fund on one quarter and see the stock-wide activity breakdown.

Quarter:

Holders: 854

Total Value: $37,879,864,945

Total Shares: 139,541,885

Showing 1–100 of 886 holdings by value · page 1 of 9

Institutional Ownership Trend

27 quarters

Superinvestors Holding NTRA

All superinvestors →

Stanley Druckenmiller $864,922,763 Philippe Laffont $680,006,677 Andreas Halvorsen $106,053,886 Joel Greenblatt $27,795,937 Ray Dalio $1,665,617

Holder Concentration

Across 853 13F filers

Top Holder

9.2%

of institutional value

Top 5

32.1%

of institutional value

Top 10

45.8%

of institutional value

HHI

306

Broadly held

Showing 100 of 886

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

JPMORGAN CHASE & CO

$3,492,298,164

9.22%

12,865,346

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
J.P. Morgan Trust Co of Delaware	1,543	$418,847	0.0%	Other
JPMorgan Chase Bank, N.A.	1,585,790	$430,462,695	12.3%	Other
J.P. Morgan Investment Management Inc.	10,202,726	$2,769,529,972	79.3%	Other
J.P. Morgan Securities LLC	818,568	$222,200,283	6.4%	Defined
J.P. Morgan Securities plc	263	$71,391	0.0%	Defined
JPMorgan Asset Management (UK) Ltd	144,459	$39,213,395	1.1%	Defined
JPMorgan Asset Management (Canada) Inc.	18,130	$4,921,388	0.1%	Defined
J.P. Morgan (Suisse) SA	4,245	$1,152,305	0.0%	Other
JPMorgan Asset Management (Asia Pacific) Ltd	6,382	$1,732,393	0.0%	Defined
JPMorgan Asset Management (Japan) Ltd	53,516	$14,526,918	0.4%	Defined
JPMorgan Asset Management (Taiwan) Ltd	4,500	$1,221,525	0.0%	Defined
55I, LLC	191	$51,846	0.0%	Other
J.P. Morgan SE	17,867	$4,849,996	0.1%	Defined
J.P. MORGAN MANSART MANAGEMENT Ltd	7,166	$1,945,210	0.1%	Defined

BlackRock, Inc.

$2,962,770,335

7.82%

10,914,608

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	225,529	$61,219,847	2.1%	Sole
BlackRock Investment Management (Australia) Ltd	103,508	$28,097,246	0.9%	Sole
BlackRock Asset Management Canada Ltd	23,765	$6,451,009	0.2%	Sole
BLACKROCK (SINGAPORE) LTD.	20,158	$5,471,889	0.2%	Sole
BlackRock Asset Management North Asia Ltd	34,336	$9,320,507	0.3%	Sole
BlackRock Investment Management, LLC	248,760	$67,525,902	2.3%	Sole
BLACKROCK ADVISORS LLC	780,498	$211,866,182	7.2%	Sole
BlackRock Fund Advisors	3,711,930	$1,007,603,398	34.0%	Sole
BlackRock Institutional Trust Company, N.A.	3,606,118	$978,880,731	33.0%	Sole
BlackRock Japan Co. Ltd	148,070	$40,193,601	1.4%	Sole
BlackRock Fund Managers Ltd.	765,567	$207,813,162	7.0%	Sole
BlackRock Investment Management (UK) Ltd.	213,822	$58,041,981	2.0%	Sole
BlackRock (Netherlands) B.V.	206,688	$56,105,457	1.9%	Sole
BlackRock Asset Management Ireland Ltd	589,412	$159,995,887	5.4%	Sole
BlackRock Advisors (UK) Ltd	7,138	$1,937,610	0.1%	Sole
BlackRock (Luxembourg) S.A.	10,113	$2,745,173	0.1%	Sole
BlackRock Life Ltd	26,589	$7,217,584	0.2%	Sole
BlackRock Asset Management Schweiz AG	4,877	$1,323,861	0.0%	Sole
Aperio Group, LLC	171,830	$46,643,253	1.6%	Sole
SpiderRock Advisors, LLC	15,900	$4,316,055	0.1%	Sole

PRICE T ROWE ASSOCIATES INC /MD/

$2,448,993,397

6.47%

9,021,895

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	160,018	$43,436,886	1.8%	Defined
Held directly	8,861,877	$2,405,556,511	98.2%	Sole

VANGUARD CAPITAL MANAGEMENT LLC

$1,690,649,233

4.46%

6,228,216

VANGUARD PORTFOLIO MANAGEMENT LLC

$1,569,697,628

4.14%

5,782,640

FMR LLC

$1,479,781,442

3.91%

5,451,396

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	4,380,308	$1,189,034,606	80.4%	Defined
FIDELITY MANAGEMENT TRUST CO	655,117	$177,831,509	12.0%	Defined
STRATEGIC ADVISERS LLC	66,287	$17,993,606	1.2%	Defined
Fidelity Institutional Asset Management Trust Co	623	$169,113	0.0%	Defined
FIAM LLC	347,481	$94,323,717	6.4%	Defined
Fidelity Diversifying Solutions LLC	1,580	$428,891	0.0%	Defined

WELLINGTON MANAGEMENT GROUP LLP

$1,096,858,326

2.90%

4,040,738

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wellington Management Company LLP	3,558,540	$965,965,683	88.1%	Defined
Wellington Managment Canada LLC	187,708	$50,953,336	4.6%	Defined
Wellington Management Hong Kong Ltd	15,932	$4,324,741	0.4%	Defined
Wellington Management Japan Pte Ltd	4,023	$1,092,043	0.1%	Defined
Wellington Management Singapore Pte. Ltd.	4,048	$1,098,829	0.1%	Defined
Wellington Management International Ltd	18,964	$5,147,777	0.5%	Defined
Wellington Trust Company, NA joint	251,051	$68,147,793	6.2%	Defined
Wellington Management Australia Pty Ltd	472	$128,124	0.0%	Defined

STATE STREET CORP

$992,317,506

2.62%

3,655,618

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS (JAPAN) CO., LTD. joint	3,655,618	$992,317,506	100.0%	Defined

Duquesne Family Office LLC

$864,922,763

2.28%

3,186,306

GEODE CAPITAL MANAGEMENT, LLC

$746,491,028

1.97%

2,750,013

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	367,032	$99,630,836	13.3%	Defined
Held directly	2,382,981	$646,860,192	86.7%	Defined

FRANKLIN RESOURCES INC

$739,946,367

1.95%

2,725,903

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FIDUCIARY TRUST CO INTERNATIONAL	198	$53,747	0.0%	Defined
Fiduciary Trust International of the South	45	$12,215	0.0%	Defined
FIDUCIARY TRUST INTERNATIONAL LLC	1,039	$282,036	0.0%	Defined
FRANKLIN ADVISERS INC	2,710,824	$735,853,174	99.4%	Defined
FRANKLIN ADVISORY SERVICES LLC	1,283	$348,270	0.0%	Defined
Franklin Templeton International Services S.a r.l	12,494	$3,391,496	0.5%	Defined
PUTNAM INVESTMENT MANAGEMENT LLC	20	$5,429	0.0%	Defined

Castle Hook Partners LP

$704,759,120

1.86%

2,596,276

COATUE MANAGEMENT LLC

$680,006,677

1.80%

2,505,090

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
COATUE OFFSHORE MASTER FUND LP	811,253	$220,214,626	32.4%	Defined
Coatue Strategic Maple Long Only Fund LP	104,996	$28,501,164	4.2%	Defined
Coatue Long Only Offshore Master Fund Ltd	674,722	$183,153,286	26.9%	Defined
Coatue F1 LP	34,770	$9,438,316	1.4%	Defined
Coatue Opportunity Fund II LP	549,954	$149,285,013	22.0%	Defined
Coatue Cardinal Main Fund LP	32,605	$8,850,627	1.3%	Defined
Coatue Innovative Strategies Fund joint	296,790	$80,563,645	11.8%	Defined

AQR CAPITAL MANAGEMENT LLC

$636,433,522

1.68%

2,344,570

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Petros Family Wealth, LLC	32	$8,686	0.0%	Other
KEB ASSET MANAGEMENT, LLC	48	$13,029	0.0%	Other
RF&L WEALTH MANAGEMENT, LLC	17	$4,614	0.0%	Other
Prism Planning Partners LLC	205	$55,647	0.0%	Other
Compound Planning, Inc.	14	$3,800	0.0%	Other
DOPKINS WEALTH MANAGEMENT, LLC	74	$20,087	0.0%	Other
Arax Advisory Partners	12,801	$3,474,831	0.5%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	8,149	$2,212,046	0.3%	Other
Stokes Family Office, LLC	4,022	$1,091,771	0.2%	Other
NATIONAL MUTUAL FUNDS MANAGEMENT LTD	26,000	$7,057,700	1.1%	Other
Held directly	2,293,208	$622,491,311	97.8%	Defined

FARALLON CAPITAL MANAGEMENT LLC

$602,175,450

1.59%

2,218,366

MASSACHUSETTS FINANCIAL SERVICES CO /MA/

$535,134,085

1.41%

1,971,391

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MFS Institutional Advisors, Inc.	135,094	$36,671,266	6.9%	Defined
MFS Investment Management CO (LUX) S.A.R.L.	12,287	$3,335,306	0.6%	Defined
MFS Investment Management K.K.	11,257	$3,055,712	0.6%	Defined
MFS Heritage Trust CO	311,240	$84,486,098	15.8%	Defined
MFS INVESTMENT MANAGEMENT CANADA LTD	3,237	$878,683	0.2%	Defined
Held directly	1,498,276	$406,707,020	76.0%	Sole

NORGES BANK

$496,243,716

1.31%

1,828,122

FRED ALGER MANAGEMENT, LLC

$482,640,542

1.27%

1,778,009

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Weatherbie Capital, LLC	56,565	$15,354,569	3.2%	Sole
Held directly	1,721,444	$467,285,973	96.8%	Sole

GOLDMAN SACHS GROUP INC

$466,935,259

1.23%

1,720,152

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	1,146,066	$311,099,615	66.6%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	572,921	$155,519,405	33.3%	Defined
Goldman Sachs Trust Company, N.A.	1,165	$316,239	0.1%	Defined

Sofinnova Investments, Inc.

$420,823,506

1.11%

1,550,280

Capital World Investors

$408,365,851

1.08%

1,504,387

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CAPITAL GROUP COMPANIES INC joint	1,504,387	$408,365,851	100.0%	Defined

ARROWSTREET CAPITAL, LIMITED PARTNERSHIP

$408,180,993

1.08%

1,503,706

MAVERICK CAPITAL LTD

$368,266,171

0.97%

1,356,663

D. E. Shaw & Co., Inc.

$347,253,769

0.92%

1,279,255

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	1,186,577	$322,096,326	92.8%	Defined
D. E. Shaw Investment Management, L.L.C. joint	92,678	$25,157,443	7.2%	Defined

MORGAN STANLEY

$345,905,201

0.91%

1,274,287

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	4,368	$1,185,693	0.3%	Defined
MORGAN STANLEY & CO. LLC	132,837	$36,058,603	10.4%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	96,627	$26,229,399	7.6%	Defined
MORGAN STANLEY INVESTMENT MANAGEMENT INC	8,350	$2,266,607	0.7%	Defined
MORGAN STANLEY SMITH BARNEY LLC	440,117	$119,469,759	34.5%	Defined
Morgan Stanley Wealth Management Australia Pty Ltd	436	$118,352	0.0%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	508,134	$137,932,973	39.9%	Defined
EATON VANCE MANAGEMENT	22,074	$5,991,987	1.7%	Defined
Calvert Research & Management	26,061	$7,074,258	2.0%	Defined
BOSTON MANAGEMENT & RESEARCH	35,283	$9,577,570	2.8%	Defined

FEDERATED HERMES, INC.

$309,172,862

0.82%

1,138,968

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FEDERATED INVESTMENT COUNSELING	3,871	$1,050,782	0.3%	Defined
FEDERATED GLOBAL INVESTMENT MANAGEMENT CORP	530,516	$144,008,568	46.6%	Defined
FEDERATED MDTA LLC	604,581	$164,113,512	53.1%	Defined

T. Rowe Price Investment Management, Inc.

$305,039,494

0.81%

1,123,741

Clearbridge Investments, LLC

$276,424,321

0.73%

1,018,325

RTW INVESTMENTS, LP

$271,450,000

0.72%

1,000,000

JANE STREET GROUP, LLC

$251,630,349

0.66%

926,986

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	444,949	$120,781,406	48.0%	Defined
Jane Street Global Trading, LLC	482,037	$130,848,943	52.0%	Defined

VANGUARD FIDUCIARY TRUST CO

$249,356,413

0.66%

918,609

Invesco Ltd.

$237,065,967

0.63%

873,332

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	661,757	$179,633,935	75.8%	Defined
Invesco Trust Co joint	55,763	$15,136,866	6.4%	Defined
Invesco Asset Management (Japan) Limited	3,039	$824,936	0.3%	Defined
Invesco Capital Management LLC	152,773	$41,470,230	17.5%	Defined

NORTHERN TRUST CORP

$227,070,092

0.60%

836,508

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	687,221	$186,546,138	82.2%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	143,875	$39,054,868	17.2%	Defined
Northern Trust Asset Management Australia Pty Ltd	1,621	$440,020	0.2%	Defined
Northern Trust Co of Hong Kong Ltd	3,791	$1,029,066	0.5%	Defined

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$226,219,910

0.60%

833,376

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	72,659	$19,723,285	8.7%	Defined
UBS Asset Management Switzerland AG	263,684	$71,577,020	31.6%	Defined
UBS Asset Management (UK) Ltd.	238,858	$64,838,003	28.7%	Defined
UBS Asset Management Life Ltd.	6,184	$1,678,646	0.7%	Defined
Held directly	251,991	$68,402,956	30.2%	Defined

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$218,934,196

0.58%

806,536

NWI MANAGEMENT LP

$207,008,584

0.55%

762,603

BANK OF AMERICA CORP /DE/

$199,960,109

0.53%

736,637

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	119,151	$32,343,538	16.2%	Defined
MERRILL LYNCH, PIERCE, FENNER & SMITH INC.	4,682	$1,270,928	0.6%	Defined
BOFA SECURITIES, INC.	21,411	$5,812,015	2.9%	Defined
MERRILL LYNCH INTERNATIONAL	9,176	$2,490,825	1.2%	Defined
MANAGED ACCOUNT ADVISORS LLC joint	582,217	$158,042,803	79.0%	Defined

WESTFIELD CAPITAL MANAGEMENT CO LP

$197,243,170

0.52%

726,628

ARK Investment Management LLC

$186,989,689

0.49%

688,855

Sumitomo Mitsui Trust Group, Inc.

$174,716,892

0.46%

643,643

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUMITOMO MITSUI TRUST BANK Ltd joint	643,643	$174,716,892	100.0%	Defined

Chevy Chase Trust Holdings, LLC

$162,830,911

0.43%

599,856

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CHEVY CHASE TRUST Co	599,856	$162,830,911	100.0%	Defined

RAYMOND JAMES FINANCIAL INC

$160,992,378

0.43%

593,083

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	131,794	$35,775,481	22.2%	Defined
Raymond James Financial Services Advisors, Inc.	9,490	$2,576,060	1.6%	Defined
EAGLE ASSET MANAGEMENT INC joint	429,495	$116,586,417	72.4%	Defined
CHARTWELL INVESTMENT PARTNERS, LLC joint	21,258	$5,770,484	3.6%	Defined
RAYMOND JAMES TRUST N.A.	1,046	$283,936	0.2%	Defined

WORLDQUANT MILLENNIUM ADVISORS LLC

$160,551,817

0.42%

591,460

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MILLENNIUM MANAGEMENT LLC	591,460	$160,551,817	100.0%	Defined

UBS Group AG

$150,312,449

0.40%

553,739

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	110,482	$29,990,338	20.0%	Defined
UBS AG	431,195	$117,047,882	77.9%	Defined
UBS Switzerland AG	5,423	$1,472,073	1.0%	Defined
UBS Europe SE	5,300	$1,438,685	1.0%	Defined
UBS Securities LLC	1,339	$363,471	0.2%	Defined

Legal & General Group Plc

$150,142,794

0.40%

553,114

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	107,945	$29,301,670	19.5%	Defined
Legal & General Investment Management Ltd joint	445,169	$120,841,124	80.5%	Defined

Bank of New York Mellon Corp

$149,277,677

0.39%

549,927

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNY Mellon Trust of Delaware	34	$9,229	0.0%	Defined
Mellon Investments Corporation joint	260,479	$70,707,023	47.4%	Defined
BNY Mellon Securities Corporation	741	$201,144	0.1%	Defined
The Bank of New York Mellon	119,718	$32,497,451	21.8%	Defined
BNY Mellon, NA joint	10,503	$2,851,038	1.9%	Defined
BNY Mellon Advisors, Inc. joint	75,340	$20,451,042	13.7%	Defined
Newton Investment Management North America, LLC joint	83,110	$22,560,208	15.1%	Defined
Held directly	2	$542	0.0%	Defined

FIRST TRUST ADVISORS LP

$138,769,040

0.37%

511,214

WELLS FARGO & COMPANY/MN

$136,489,944

0.36%

502,818

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	43,254	$11,741,298	8.6%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	110,229	$29,921,662	21.9%	Defined
WELLS FARGO CLEARING SERVICES, LLC	344,162	$93,422,774	68.4%	Defined
Wells Fargo Securities, LLC	2,787	$756,531	0.6%	Defined
WELLS FARGO INVESTMENT INSTITUTE, INC.	2,386	$647,679	0.5%	Defined

Nuveen, LLC

$135,477,707

0.36%

499,089

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	42,915	$11,649,276	8.6%	Defined
TIAA CREF INVESTMENT MANAGEMENT LLC joint	220,561	$59,871,283	44.2%	Defined
TEACHERS ADVISORS, LLC joint	235,613	$63,957,148	47.2%	Defined

GILDER GAGNON HOWE & CO LLC

$135,199,472

0.36%

498,064

AMERICAN CENTURY COMPANIES INC

$129,951,801

0.34%

478,732

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AMERICAN CENTURY INVESTMENT MANAGEMENT INC	478,732	$129,951,801	100.0%	Defined

LORD, ABBETT & CO. LLC

$126,705,530

0.33%

466,773

VICTORY CAPITAL MANAGEMENT INC

$125,949,542

0.33%

463,988

Avala Global LP

$124,867,000

0.33%

460,000

Allspring Global Investments Holdings, LLC

$112,469,606

0.30%

414,329

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Allspring Funds Management, LLC reporting on behalf of Morgan Stanley Smith Barney, LLC	16,375	$4,444,993	4.0%	Other
Allspring Global Investments, LLC reporting on behalf of UBS ASSET MANAGEMENT AMERICAS INC	397,954	$108,024,613	96.0%	Defined

Robeco Institutional Asset Management B.V.

$110,918,813

0.29%

408,616

Man Group plc

$110,413,101

0.29%

406,753

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AHL Partners LLP	47,735	$12,957,665	11.7%	Defined
Numeric Investors LLC	359,018	$97,455,436	88.3%	Defined

AMERIPRISE FINANCIAL INC

$108,130,475

0.29%

398,344

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	277,451	$75,314,073	69.7%	Defined
Threadneedle Asset Management Holdings LTD	2,334	$633,564	0.6%	Defined
Ameriprise Financial Services, LLC	38,882	$10,554,518	9.8%	Defined
Ameriprise Bank, FSB	231	$62,704	0.1%	Defined
Columbia Threadneedle Management Ltd	79,446	$21,565,616	19.9%	Defined

VIKING GLOBAL INVESTORS LP

$106,053,886

0.28%

390,694

SUMMIT PARTNERS PUBLIC ASSET MANAGEMENT, LLC

$105,594,050

0.28%

389,000

Driehaus Capital Management LLC

$104,202,054

0.28%

383,872

GOLDMAN SACHS GROUP INC

$103,829,625

0.27%

382,500

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	382,500	$103,829,625	100.0%	Defined

Swiss National Bank

$103,096,710

0.27%

379,800

JANE STREET GROUP, LLC

$100,300,775

0.26%

369,500

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	369,500	$100,300,775	100.0%	Defined

DIMENSIONAL FUND ADVISORS LP

$100,196,808

0.26%

369,117

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	3,510	$952,789	1.0%	Defined
DFA Australia Ltd.	35,299	$9,581,913	9.6%	Defined
Dimensional Fund Advisors Pte. Ltd.	1,040	$282,308	0.3%	Defined
Dimensional Ireland Ltd	24,192	$6,566,918	6.6%	Defined
Held directly	305,076	$82,812,880	82.7%	Sole

Amova Asset Management Americas, Inc.

$99,911,515

0.26%

368,066

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Sumitomo Mitsui Trust Group, Inc. joint	368,066	$99,911,515	100.0%	Sole

California Public Employees Retirement System

$93,399,430

0.25%

344,076

Squarepoint Ops LLC

$91,279,948

0.24%

336,268

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Squarepoint Capital LLP	336,268	$91,279,948	100.0%	Defined

MAPLELANE CAPITAL, LLC

$88,221,250

0.23%

325,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NRMA, LLC joint	126,500	$34,338,425	38.9%	Defined
Maplelane Master Fund, Ltd.	136,700	$37,107,215	42.1%	Defined
Maplelane 2 Fund, LP	51,200	$13,898,240	15.8%	Defined
Held directly	10,600	$2,877,370	3.3%	Sole

DEUTSCHE BANK AG\

$86,874,041

0.23%

320,037

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Deutsche Bank Trust Co Americas	7	$1,900	0.0%	Defined
DWS Investment GmbH	21,298	$5,781,342	6.7%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC	6,943	$1,884,677	2.2%	Defined
DWS Investments UK Ltd	235,257	$63,860,512	73.5%	Defined
DBX Advisors LLC	9,305	$2,525,842	2.9%	Defined
DWS Investments Hong Kong Ltd	6,784	$1,841,516	2.1%	Defined
DWS International GmbH	40,443	$10,978,252	12.6%	Defined

Amundi

$85,684,277

0.23%

315,654

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Amundi Asset Management joint	315,654	$85,684,277	100.0%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$82,819,395

0.22%

305,100

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	305,100	$82,819,395	100.0%	Other

NOMURA ASSET MANAGEMENT CO LTD

$82,022,145

0.22%

302,163

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nomura Asset Management U.K. Ltd	10,602	$2,877,912	3.5%	Sole
Held directly	291,561	$79,144,233	96.5%	Sole

Capital Impact Advisors, LLC

$80,411,090

0.21%

296,228

ROYAL BANK OF CANADA

$80,120,905

0.21%

295,159

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC Global Asset Management Inc.	143,535	$38,962,575	48.6%	Defined
RBC Capital Markets, LLC	102,683	$27,873,300	34.8%	Defined
RBC Dominion Securities Inc.	41,049	$11,142,751	13.9%	Defined
RBC Phillips, Hager & North Investment Counsel Inc.	3,682	$999,478	1.2%	Defined
RBC Private Counsel (USA) Inc.	124	$33,659	0.0%	Defined
RBC Trust Co (Delaware) Ltd	262	$71,119	0.1%	Defined
CITY NATIONAL BANK	17	$4,614	0.0%	Defined
RBC Rochdale, LLC	1,910	$518,469	0.6%	Defined
Held directly	1,897	$514,940	0.6%	Sole

ALLIANCEBERNSTEIN L.P.

$79,281,858

0.21%

292,068

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	292,068	$79,281,858	100.0%	Defined

ENVESTNET ASSET MANAGEMENT INC

$74,937,572

0.20%

276,064

CITADEL ADVISORS LLC

$72,099,020

0.19%

265,607

VANGUARD ASSET MANAGEMENT, Ltd

$71,251,553

0.19%

262,485

ORBIMED ADVISORS LLC

$71,119,900

0.19%

262,000

PRINCIPAL FINANCIAL GROUP INC

$70,377,484

0.19%

259,265

PointState Capital LP

$70,037,357

0.18%

258,012

ZEVENBERGEN CAPITAL INVESTMENTS LLC

$69,097,326

0.18%

254,549

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
VIRTUS CAPITAL ADVISERS, LLC	48,500	$13,165,325	19.1%	Other
Held directly	206,049	$55,932,001	80.9%	Sole

Point72 Asset Management, L.P.

$68,655,134

0.18%

252,920

VOYA INVESTMENT MANAGEMENT LLC

$68,570,983

0.18%

252,610

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	252,610	$68,570,983	100.0%	Defined

Assenagon Asset Management S.A.

$68,098,390

0.18%

250,869

GW&K Investment Management, LLC

$62,788,555

0.17%

231,308

National Pension Service

$61,139,769

0.16%

225,234

MAPLELANE CAPITAL, LLC

$60,261,899

0.16%

222,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NRMA, LLC joint	86,400	$23,453,280	38.9%	Defined
Maplelane Master Fund, Ltd.	93,383	$25,348,815	42.1%	Defined
Maplelane 2 Fund, LP	34,927	$9,480,934	15.7%	Defined
Held directly	7,290	$1,978,870	3.3%	Sole

CITIGROUP INC

$58,735,264

0.16%

216,376

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citibank (Switzerland) AG joint	173	$46,960	0.1%	Defined
Citibank, N.A. joint	10,685	$2,900,443	4.9%	Defined
Citigroup Financial Products Inc. joint	205,518	$55,787,861	95.0%	Defined

NEUBERGER BERMAN GROUP LLC

$57,474,650

0.15%

211,732

Handelsbanken Fonder AB

$56,407,310

0.15%

207,800

CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM

$56,324,789

0.15%

207,496

TD Asset Management Inc

$56,207,522

0.15%

207,064

CITADEL ADVISORS LLC

$54,561,450

0.14%

201,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	201,000	$54,561,450	100.0%	Defined

Artemis Investment Management LLP

$54,313,616

0.14%

200,087

Mitsubishi UFJ Asset Management Co., Ltd.

$50,424,280

0.13%

185,759

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MITSUBISHI UFJ FINANCIAL GROUP INC	185,759	$50,424,280	100.0%	Defined

BNP PARIBAS ARBITRAGE, SA

$49,380,283

0.13%

181,913

SEI INVESTMENTS CO

$49,099,332

0.13%

180,878

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI INVESTMENTS MANAGEMENT CORP	98,748	$26,805,144	54.6%	Sole
SEI TRUST CO	82,130	$22,294,188	45.4%	Sole

Eversept Partners, LP

$47,722,810

0.13%

175,807

Key facts CIK 1604821 CUSIP 632307104 13F (30d) 31 filings 25 filers Visit website Investor relations