TWST · Twist Bioscience Corp · Shareholders & Ownership

$146.98 +7.21 (+5.16%)

Russell 2000 member Rebalance outlook locked

Market Cap

$8.53B

62.71M

Volume · Oct 20 1.42M Avg daily vol (3M) 2.16M

Every disclosed holder of TWST in one list: 13F institutions, Schedule 13D and 13G filers, insiders (Forms 3, 4 and 5) and registered funds, each from its own latest filing. Holders nest by CIK and SEC reporting links, never by name, and each is summed once.

How the holder list picks a filing →

Who Owns TWST

13F quarter Sep 30, 2026, combined view: filers that have not filed yet carry their Jun 30, 2026 positions

No chart: the summed disclosures exceed shares outstanding, so they are not parts of one whole.

Holder group	Holders	Shares	% Out	Summed
Institutional (13F)	347	76,419,432	121.87%	Yes
Insiders (Forms 3, 4, 5)	13	1,345,948	2.15%	Yes
13D/G-only blocks	1	1,821,238	2.90%	Yes
Remaining	—	16,879,194	-26.92%	Negative
Registered funds (N-PORT, not summed)	350	33,031,414	52.68%	No

Shares outstanding: 62,707,424, the issuer's latest cover-page count restated onto today's split basis.

Remaining is negative: the summed disclosures exceed shares outstanding, as when filings are dated at different times or holders that do not disclose their links count the same shares. It is flagged, not clipped.

13D/G-only counts each filer's latest 13D or 13G that no 13F report of the listing contains, that names no 13F filer and that no counted insider files. Funds are never summed: their shares sit inside their advisers' 13F reports.

Holders of TWST

One row per holder Every filing By ultimate parent

All 13F Funds Insiders 13D/G Non-US

#	Holder	Shares	% Out	Change	As of	Notes
1	FMR LLC 13G CIK 315066 · Holding company · Aug 31, 2026 Rule 13d-1(b), qualified institution, not summed	8,760,081	13.97%	+4,057,037	Aug 31, 2026	Rule 13d-1(b), qualified institution, not summed
1.1	FMR LLC 13F CIK 315066 · Holding company · Jun 30, 2026 carried, not yet filed for the newest quarter	4,703,044	7.50%	+2,813,099	Jun 30, 2026	carried, not yet filed for the newest quarter
1.2	Fidelity Small Cap Index Fund Fund S000033638 · Registered fund · Jul 31, 2026 Fidelity Salem Street Trust, not summed	679,049	1.08%	+79,516	Jul 31, 2026	Fidelity Salem Street Trust, not summed
1.3	Fidelity Small Cap Growth Fund Fund S000007193 · Registered fund · Jul 31, 2026 Fidelity Securities Fund, not summed	448,130	0.71%	-157,060	Jul 31, 2026	Fidelity Securities Fund, not summed
1.4	Fidelity Stock Selector Small Cap Fund Fund S000005453 · Registered fund · Jul 31, 2026 Fidelity Capital Trust, not summed	383,500	0.61%	+39,700	Jul 31, 2026	Fidelity Capital Trust, not summed
1.5	Fidelity Extended Market Index Fund Fund S000006021 · Registered fund · May 31, 2026 Fidelity Concord Street Trust, not summed	317,308	0.51%	+985	May 31, 2026	Fidelity Concord Street Trust, not summed
2	BlackRock, Inc. 13F CIK 2012383 · Holding company · Jun 30, 2026 carried, not yet filed for the newest quarter	5,833,891	9.30%	+279,529	Jun 30, 2026	carried, not yet filed for the newest quarter
2.1	iShares Russell 2000 ETF Fund S000004344 · Registered fund · Jun 30, 2026 iShares Trust, not summed	1,594,038	2.54%	+110,155	Jun 30, 2026	iShares Trust, not summed
2.2	iShares Russell 2000 Growth ETF Fund S000004343 · Registered fund · Jun 30, 2026 iShares Trust, not summed	573,272	0.91%	+79,108	Jun 30, 2026	iShares Trust, not summed
2.3	iShares Biotechnology ETF Fund S000004350 · Registered fund · Jun 30, 2026 iShares Trust, not summed	422,334	0.67%	-14,461	Jun 30, 2026	iShares Trust, not summed
2.4	Master Small Cap Index Series Fund S000002716 · Registered fund · Jun 30, 2026 Quantitative Master Series LLC, not summed	114,494	0.18%	+14,753	Jun 30, 2026	Quantitative Master Series LLC, not summed
2.5	iShares Genomics Immunology and Healthcare ETF Fund S000065599 · Registered fund · Jul 31, 2026 iShares Trust, not summed	99,937	0.16%	-3,999	Jul 31, 2026	iShares Trust, not summed
3	Artisan Partners Limited Partnership 13F CIK 1466153 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	4,802,017	7.66%	-1,565,760	Jun 30, 2026	carried, not yet filed for the newest quarter
3.1	Artisan Partners Limited Partnership 13G CIK 1466153 · Registered investment adviser · Jun 30, 2026 Rule 13d-1(b), qualified institution, not summed	4,802,017	7.66%	-259,016	Jun 30, 2026	Rule 13d-1(b), qualified institution, not summed
3.2	Artisan Mid Cap Fund Fund S000006496 · Registered fund · Jun 30, 2026 Artisan Partners Funds Inc., not summed	893,167	1.42%	-251,133	Jun 30, 2026	Artisan Partners Funds Inc., not summed
3.3	Artisan Small Cap Fund Fund S000006498 · Registered fund · Jun 30, 2026 Artisan Partners Funds Inc., not summed	516,903	0.82%	-206,169	Jun 30, 2026	Artisan Partners Funds Inc., not summed
3.4	Artisan Global Discovery Fund Fund S000058458 · Registered fund · Jun 30, 2026 Artisan Partners Funds Inc., not summed	28,547	0.05%	-25,123	Jun 30, 2026	Artisan Partners Funds Inc., not summed
4	PRICE T ROWE ASSOCIATES INC /MD/ 13F CIK 80255 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	3,634,652	5.80%	+3,564,347	Jun 30, 2026	carried, not yet filed for the newest quarter
4.1	PRICE T ROWE ASSOCIATES INC /MD/ 13G CIK 80255 · Registered investment adviser · Jun 30, 2026 Rule 13d-1(b), qualified institution, not summed	3,634,470	5.80%	—	Jun 30, 2026	Rule 13d-1(b), qualified institution, not summed
4.2	T. Rowe Price New Horizons Fund, Inc. Fund S000002106 · Registered fund · Jun 30, 2026 T. ROWE PRICE NEW HORIZONS FUND, INC., not summed	959,300	1.53%	—	Jun 30, 2026	T. ROWE PRICE NEW HORIZONS FUND, INC., not summed
4.3	T. Rowe Price Health Sciences Fund, Inc. Fund S000002088 · Registered fund · Jun 30, 2026 T. ROWE PRICE HEALTH SCIENCES FUND, INC., not summed	555,000	0.89%	—	Jun 30, 2026	T. ROWE PRICE HEALTH SCIENCES FUND, INC., not summed
4.4	T. Rowe Price Global Stock Fund Fund S000001497 · Registered fund · Jul 31, 2026 T. ROWE PRICE INTERNATIONAL FUNDS, INC., not summed	462,100	0.74%	—	Jul 31, 2026	T. ROWE PRICE INTERNATIONAL FUNDS, INC., not summed
4.5	T. Rowe Price Integrated U.S. Small-Cap Growth Equity Fund Fund S000002074 · Registered fund · Jun 30, 2026 T. ROWE PRICE INTEGRATED EQUITY FUNDS, INC., not summed	366,600	0.58%	—	Jun 30, 2026	T. ROWE PRICE INTEGRATED EQUITY FUNDS, INC., not summed
5	ARK Investment Management LLC 13G CIK 1697748 · Registered investment adviser · Aug 31, 2026 Rule 13d-1(b), qualified institution, not summed	3,385,850	5.40%	-1,163,953	Aug 31, 2026	Rule 13d-1(b), qualified institution, not summed
5.1	ARK Investment Management LLC 13F CIK 1697748 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	4,549,803	7.26%	-1,792,930	Jun 30, 2026	carried, not yet filed for the newest quarter
5.2	ARK Innovation ETF Fund S000042977 · Registered fund · Jul 31, 2026 ARK ETF Trust, not summed	2,010,667	3.21%	-1,371,997	Jul 31, 2026	ARK ETF Trust, not summed
5.3	ARK Genomic Revolution ETF Fund S000042975 · Registered fund · Jul 31, 2026 ARK ETF Trust, not summed	1,314,736	2.10%	-585,803	Jul 31, 2026	ARK ETF Trust, not summed
6	STATE STREET CORP 13F CIK 93751 · Bank · Jun 30, 2026 carried, not yet filed for the newest quarter	3,232,532	5.15%	-438,150	Jun 30, 2026	carried, not yet filed for the newest quarter
6.1	State Street(R) SPDR(R) S&P(R) Biotech ETF Fund S000010018 · Registered fund · Jun 30, 2026 SPDR SERIES TRUST, not summed	1,469,111	2.34%	-338,925	Jun 30, 2026	SPDR SERIES TRUST, not summed
6.2	State Street(R) SPDR(R) S&P Kensho New Economies Composite ETF Fund S000063362 · Registered fund · Jun 30, 2026 SPDR SERIES TRUST, not summed	278,488	0.44%	-82,120	Jun 30, 2026	SPDR SERIES TRUST, not summed
6.3	STATE STREET SMALL/MID CAP EQUITY INDEX PORTFOLIO Fund S000050332 · Registered fund · Jun 30, 2026 STATE STREET INSTITUTIONAL INVESTMENT TRUST, not summed	21,412	0.03%	+300	Jun 30, 2026	STATE STREET INSTITUTIONAL INVESTMENT TRUST, not summed
6.4	State Street(R) SPDR(R) Portfolio MSCI Global Stock Market ETF Fund S000036082 · Registered fund · Jun 30, 2026 SPDR INDEX SHARES FUNDS, not summed	3,002	0.00%	+357	Jun 30, 2026	SPDR INDEX SHARES FUNDS, not summed
6.5	State Street Total Return V.I.S. Fund Fund S000001637 · Registered fund · Jun 30, 2026 STATE STREET VARIABLE INSURANCE SERIES FUNDS INC, not summed	1,528	0.00%	+102	Jun 30, 2026	STATE STREET VARIABLE INSURANCE SERIES FUNDS INC, not summed
7	VANGUARD PORTFOLIO MANAGEMENT LLC 13F CIK 2100121 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	2,824,989	4.51%	+175,888	Jun 30, 2026	carried, not yet filed for the newest quarter
8	VANGUARD CAPITAL MANAGEMENT LLC 13F CIK 2100119 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	2,699,280	4.30%	+60,236	Jun 30, 2026	carried, not yet filed for the newest quarter
9	EdgePoint Investment Group Inc. 13F CIK 1481669 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	2,521,635	4.02%	-2,860,886	Jun 30, 2026	carried, not yet filed for the newest quarter
9.1	EdgePoint Investment Group Inc. 13G CIK 1481669 · Registered investment adviser · Jun 30, 2026 Rule 13d-1(b), qualified institution, not summed	2,521,635	4.02%	-2,856,356	Jun 30, 2026	Rule 13d-1(b), qualified institution, not summed
10	WILLIAM BLAIR INVESTMENT MANAGEMENT, LLC 13F CIK 1644956 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	2,174,594	3.47%	-3,095,124	Jun 30, 2026	carried, not yet filed for the newest quarter
10.1	WILLIAM BLAIR INVESTMENT MANAGEMENT, LLC 13G CIK 1644956 · Registered investment adviser · Jun 30, 2026 Rule 13d-1(b), qualified institution, not summed	2,174,594	3.47%	-1,632,273	Jun 30, 2026	Rule 13d-1(b), qualified institution, not summed
10.2	William Blair Small-Mid Cap Growth Fund Fund S000008603 · Registered fund · Jun 30, 2026 WILLIAM BLAIR FUNDS, not summed	202,973	0.32%	-309,100	Jun 30, 2026	WILLIAM BLAIR FUNDS, not summed
10.3	William Blair Small Cap Growth Fund Fund S000008602 · Registered fund · Jun 30, 2026 WILLIAM BLAIR FUNDS, not summed	192,214	0.31%	-257,660	Jun 30, 2026	WILLIAM BLAIR FUNDS, not summed
10.4	William Blair Small-Mid Cap Core Fund Fund S000066499 · Registered fund · Jun 30, 2026 WILLIAM BLAIR FUNDS, not summed	34,791	0.06%	-17,797	Jun 30, 2026	WILLIAM BLAIR FUNDS, not summed
10.5	William Blair Growth Fund Fund S000008594 · Registered fund · Jun 30, 2026 WILLIAM BLAIR FUNDS, not summed	18,838	0.03%	-17,013	Jun 30, 2026	WILLIAM BLAIR FUNDS, not summed
11	FARALLON CAPITAL MANAGEMENT LLC 13F CIK 909661 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	2,136,400	3.41%	+708,750	Jun 30, 2026	carried, not yet filed for the newest quarter
12	Invesco Ltd. 13F CIK 914208 · Investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	1,928,229	3.08%	-243,669	Jun 30, 2026	carried, not yet filed for the newest quarter
12.1	Invesco Discovery Fund Fund S000064618 · Registered fund · May 31, 2026 AIM Counselor Series Trust (Invesco Counselor Series Trust), not summed	822,247	1.31%	—	May 31, 2026	AIM Counselor Series Trust (Invesco Counselor Series Trust), not summed
12.2	Invesco Main Street Small Cap Fund Fund S000064650 · Registered fund · Jun 30, 2026 AIM Growth Series (Invesco Growth Series), not summed	288,002	0.46%	-135,870	Jun 30, 2026	AIM Growth Series (Invesco Growth Series), not summed
12.3	Invesco V.I. Main Street Small Cap Fund Fund S000064664 · Registered fund · Jun 30, 2026 AIM Variable Insurance Funds (Invesco Variable Insurance Funds), not summed	160,155	0.26%	-67,196	Jun 30, 2026	AIM Variable Insurance Funds (Invesco Variable Insurance Funds), not summed
12.4	Invesco Health Care Fund Fund S000000235 · Registered fund · Jul 31, 2026 AIM Investment Funds (Invesco Investment Funds), not summed	74,708	0.12%	0	Jul 31, 2026	AIM Investment Funds (Invesco Investment Funds), not summed
12.5	Invesco RAFI US 1500 Small-Mid ETF Fund S000013120 · Registered fund · Jul 31, 2026 Invesco Exchange-Traded Fund Trust, not summed	44,909	0.07%	-8,155	Jul 31, 2026	Invesco Exchange-Traded Fund Trust, not summed
13	VANGUARD TOTAL STOCK MARKET INDEX FUND Fund S000002848 · Registered fund · Jun 30, 2026 VANGUARD INDEX FUNDS, not summed	1,874,473	2.99%	+27,263	Jun 30, 2026	VANGUARD INDEX FUNDS, not summed
14	Millennium Management LLC 13G CIK 1319244 · Unknown · Dec 31, 2023 Rule 13d-1(c), passive investor	1,821,238	2.90%	—	Dec 31, 2023	Rule 13d-1(c), passive investor
15	GEODE CAPITAL MANAGEMENT, LLC 13F CIK 1214717 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	1,619,361	2.58%	+99,648	Jun 30, 2026	carried, not yet filed for the newest quarter
16	Thrivent Financial for Lutherans 13F CIK 314984 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	1,433,503	2.29%	+120,959	Jun 30, 2026	carried, not yet filed for the newest quarter
16.1	THRIVENT SMALL CAP STOCK FUND Fund S000001358 · Registered fund · Jul 31, 2026 Thrivent Mutual Funds, not summed	442,538	0.71%	-374,235	Jul 31, 2026	Thrivent Mutual Funds, not summed
16.2	THRIVENT SMALL CAP STOCK PORTFOLIO Fund S000001445 · Registered fund · Jun 30, 2026 Thrivent Series Fund, Inc., not summed	266,709	0.43%	+1,032	Jun 30, 2026	Thrivent Series Fund, Inc., not summed
16.3	Thrivent Small-Mid Cap Equity ETF Fund S000075337 · Registered fund · Jun 30, 2026 Thrivent ETF Trust, not summed	140,652	0.22%	—	Jun 30, 2026	Thrivent ETF Trust, not summed
16.4	THRIVENT SMALL CAP GROWTH PORTFOLIO Fund S000061813 · Registered fund · Jun 30, 2026 Thrivent Series Fund, Inc., not summed	30,087	0.05%	-1,288	Jun 30, 2026	Thrivent Series Fund, Inc., not summed
16.5	THRIVENT MODERATELY AGGRESSIVE ALLOCATION PORTFOLIO Fund S000001427 · Registered fund · Jun 30, 2026 Thrivent Series Fund, Inc., not summed	27,737	0.04%	+22	Jun 30, 2026	Thrivent Series Fund, Inc., not summed
17	GOLDMAN SACHS GROUP INC 13F CIK 886982 · Holding company · Jun 30, 2026 carried, not yet filed for the newest quarter	1,408,418	2.25%	-297,128	Jun 30, 2026	carried, not yet filed for the newest quarter
17.1	Goldman Sachs Small Cap Equity Insights Fund Fund S000009329 · Registered fund · Jul 31, 2026 Goldman Sachs Trust, not summed	51,433	0.08%	—	Jul 31, 2026	Goldman Sachs Trust, not summed
17.2	Goldman Sachs Small Cap Growth Insights Fund Fund S000014930 · Registered fund · Jul 31, 2026 Goldman Sachs Trust, not summed	28,346	0.05%	+24,447	Jul 31, 2026	Goldman Sachs Trust, not summed
17.3	Goldman Sachs U.S. Tax-Managed Equity Fund Fund S000009340 · Registered fund · Jun 30, 2026 Goldman Sachs Trust, not summed	24,910	0.04%	—	Jun 30, 2026	Goldman Sachs Trust, not summed
17.4	Goldman Sachs Variable Insurance Trust Small Cap Equity Insights Fund Fund S000009355 · Registered fund · Jun 30, 2026 Goldman Sachs Variable Insurance Trust, not summed	13,031	0.02%	—	Jun 30, 2026	Goldman Sachs Variable Insurance Trust, not summed
17.5	Goldman Sachs Small Cap Value Insights Fund Fund S000014929 · Registered fund · Jul 31, 2026 Goldman Sachs Trust, not summed	8,979	0.01%	—	Jul 31, 2026	Goldman Sachs Trust, not summed
18	VANGUARD SMALL-CAP INDEX FUND Fund S000002845 · Registered fund · Jun 30, 2026 VANGUARD INDEX FUNDS, not summed	1,357,205	2.16%	+12,797	Jun 30, 2026	VANGUARD INDEX FUNDS, not summed
19	MORGAN STANLEY 13F CIK 895421 · Holding company · Jun 30, 2026 carried, not yet filed for the newest quarter	1,189,861	1.90%	+643,016	Jun 30, 2026	carried, not yet filed for the newest quarter
19.1	E*TRADE No Fee Total Market Index Fund Fund S000090722 · Registered fund · Jul 31, 2026 E Trade Trust, not summed	134	0.00%	+82	Jul 31, 2026	E Trade Trust, not summed
20	D. E. Shaw & Co., Inc. 13F CIK 1009207 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	1,185,605	1.89%	+1,138,961	Jun 30, 2026	carried, not yet filed for the newest quarter
21	BAILLIE GIFFORD & CO 13F CIK 1088875 · Investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	1,086,779	1.73%	-128,372	Jun 30, 2026	carried, not yet filed for the newest quarter
22	Driehaus Capital Management LLC 13F CIK 938206 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	1,022,526	1.63%	+437,830	Jun 30, 2026	carried, not yet filed for the newest quarter
22.1	Driehaus Small Cap Growth Fund Fund S000058261 · Registered fund · Jun 30, 2026 Driehaus Mutual Funds, not summed	216,331	0.35%	+102,725	Jun 30, 2026	Driehaus Mutual Funds, not summed
23	Sumitomo Mitsui Trust Group, Inc. 13F CIK 1475365 · Bank · Jun 30, 2026 carried, not yet filed for the newest quarter	1,000,344	1.60%	-495,755	Jun 30, 2026	carried, not yet filed for the newest quarter
24	Amova Asset Management Americas, Inc. 13F CIK 1480751 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	999,734	1.59%	-496,365	Jun 30, 2026	carried, not yet filed for the newest quarter
25	BANK OF AMERICA CORP /DE/ 13F CIK 70858 · Bank · Jun 30, 2026 carried, not yet filed for the newest quarter	922,647	1.47%	+338,386	Jun 30, 2026	carried, not yet filed for the newest quarter
26	Bridge Builder Small/Mid Cap Growth Fund Fund S000048336 · Registered fund · Jun 30, 2026 Bridge Builder Trust, not summed	866,923	1.38%	+81,088	Jun 30, 2026	Bridge Builder Trust, not summed
27	CITADEL ADVISORS LLC 13F CIK 1423053 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	862,042	1.37%	-358,358	Jun 30, 2026	carried, not yet filed for the newest quarter
28	LOOMIS SAYLES & CO L P 13F CIK 312348 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	861,309	1.37%	+369,019	Jun 30, 2026	carried, not yet filed for the newest quarter
28.1	Loomis Sayles Small Cap Growth Fund Fund S000006705 · Registered fund · Jun 30, 2026 LOOMIS SAYLES FUNDS II, not summed	360,266	0.57%	+83,828	Jun 30, 2026	LOOMIS SAYLES FUNDS II, not summed
28.2	Loomis Sayles Small/Mid Cap Growth Fund Fund S000049859 · Registered fund · Jun 30, 2026 LOOMIS SAYLES FUNDS II, not summed	26,203	0.04%	—	Jun 30, 2026	LOOMIS SAYLES FUNDS II, not summed
29	VANGUARD SMALL-CAP GROWTH INDEX FUND Fund S000002846 · Registered fund · Jun 30, 2026 VANGUARD INDEX FUNDS, not summed	780,792	1.25%	+29,763	Jun 30, 2026	VANGUARD INDEX FUNDS, not summed
30	Stephens Investment Management Group LLC 13F CIK 1314440 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	767,364	1.22%	+767,364	Jun 30, 2026	carried, not yet filed for the newest quarter
31	First Light Asset Management, LLC 13F CIK 1600004 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	727,933	1.16%	-150,915	Jun 30, 2026	carried, not yet filed for the newest quarter
32	VANGUARD EXTENDED MARKET INDEX FUND Fund S000002841 · Registered fund · Jun 30, 2026 VANGUARD INDEX FUNDS, not summed	714,822	1.14%	+31,687	Jun 30, 2026	VANGUARD INDEX FUNDS, not summed
33	VOYA INVESTMENT MANAGEMENT LLC 13F CIK 1068837 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	671,866	1.07%	+661,478	Jun 30, 2026	carried, not yet filed for the newest quarter
34	FIL Ltd 13F CIK 318989 · Non-US institution · Jun 30, 2026 carried, not yet filed for the newest quarter	641,096	1.02%	+641,096	Jun 30, 2026	carried, not yet filed for the newest quarter
35	VANGUARD EXPLORER FUND Fund S000002578 · Registered fund · Jul 31, 2026 VANGUARD EXPLORER FUND, not summed	604,500	0.96%	—	Jul 31, 2026	VANGUARD EXPLORER FUND, not summed
36	T. Rowe Price Investment Management, Inc. 13F CIK 1897612 · Investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	597,935	0.95%	+597,935	Jun 30, 2026	carried, not yet filed for the newest quarter
37	NORTHERN TRUST CORP 13F CIK 73124 · Bank · Jun 30, 2026 carried, not yet filed for the newest quarter	591,688	0.94%	+63,722	Jun 30, 2026	carried, not yet filed for the newest quarter
37.1	Small Cap Index Fund Fund S000001245 · Registered fund · Jun 30, 2026 Northern Funds, not summed	31,407	0.05%	+3,038	Jun 30, 2026	Northern Funds, not summed
37.2	Small Cap Core Fund Fund S000001244 · Registered fund · Jun 30, 2026 Northern Funds, not summed	5,842	0.01%	—	Jun 30, 2026	Northern Funds, not summed
37.3	FlexShares Morningstar US Market Factor Tilt Index Fund Fund S000033312 · Registered fund · Jul 31, 2026 FlexShares Trust, not summed	2,430	0.00%	+120	Jul 31, 2026	FlexShares Trust, not summed
38	DIMENSIONAL FUND ADVISORS LP 13F CIK 354204 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	582,702	0.93%	-198,916	Jun 30, 2026	carried, not yet filed for the newest quarter
38.1	Dimensional U.S. Small Cap ETF Fund S000070906 · Registered fund · Jul 31, 2026 Dimensional ETF Trust, not summed	71,347	0.11%	-49,924	Jul 31, 2026	Dimensional ETF Trust, not summed
38.2	U.S. MICRO CAP PORTFOLIO Fund S000000948 · Registered fund · Jul 31, 2026 DFA INVESTMENT DIMENSIONS GROUP INC, not summed	61,060	0.10%	-129,502	Jul 31, 2026	DFA INVESTMENT DIMENSIONS GROUP INC, not summed
38.3	Dimensional U.S. Core Equity 2 ETF Fund S000070903 · Registered fund · Jul 31, 2026 Dimensional ETF Trust, not summed	60,037	0.10%	-40,163	Jul 31, 2026	Dimensional ETF Trust, not summed
38.4	U.S. SMALL CAP PORTFOLIO Fund S000000971 · Registered fund · Jul 31, 2026 DFA INVESTMENT DIMENSIONS GROUP INC, not summed	50,311	0.08%	-144,708	Jul 31, 2026	DFA INVESTMENT DIMENSIONS GROUP INC, not summed
39	WELLINGTON MANAGEMENT GROUP LLP 13F CIK 902219 · Holding company · Jun 30, 2026 carried, not yet filed for the newest quarter	568,347	0.91%	-170,282	Jun 30, 2026	carried, not yet filed for the newest quarter
40	UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC 13F CIK 861177 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	563,782	0.90%	-260,194	Jun 30, 2026	carried, not yet filed for the newest quarter
40.1	PACE Small/Medium Co Growth Equity Investments Fund S000002974 · Registered fund · Jul 31, 2026 PACE SELECT ADVISORS TRUST, not summed	3,749	0.01%	—	Jul 31, 2026	PACE SELECT ADVISORS TRUST, not summed
41	Leproust Emily M. Insider CIK 1753655 · Oct 5, 2026 Director, issuer-wide	562,760	0.90%	-4,548	Oct 5, 2026	Director, issuer-wide
42	FRED ALGER MANAGEMENT, LLC 13F CIK 3520 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	558,157	0.89%	+462,374	Jun 30, 2026	carried, not yet filed for the newest quarter
42.1	ALGER SMALL CAP FOCUS FUND Fund S000020958 · Registered fund · Jul 31, 2026 ALGER FUNDS, not summed	222,288	0.35%	0	Jul 31, 2026	ALGER FUNDS, not summed
42.2	ALGER MID CAP FOCUS FUND Fund S000063650 · Registered fund · Jul 31, 2026 ALGER FUNDS, not summed	141,711	0.23%	+28,650	Jul 31, 2026	ALGER FUNDS, not summed
42.3	Alger Mid Cap 40 ETF Fund S000069955 · Registered fund · Jun 30, 2026 Alger ETF Trust, not summed	71,651	0.11%	—	Jun 30, 2026	Alger ETF Trust, not summed
42.4	ALGER SMALL CAP GROWTH FUND Fund S000009160 · Registered fund · Jul 31, 2026 ALGER FUNDS, not summed	50,706	0.08%	-1,599	Jul 31, 2026	ALGER FUNDS, not summed
42.5	Alger Small Cap Growth Portfolio Fund S000009215 · Registered fund · Jun 30, 2026 ALGER PORTFOLIOS, not summed	26,830	0.04%	0	Jun 30, 2026	ALGER PORTFOLIOS, not summed
43	CHARLES SCHWAB INVESTMENT MANAGEMENT INC 13F CIK 884546 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	557,131	0.89%	+20,020	Jun 30, 2026	carried, not yet filed for the newest quarter
43.1	Schwab U.S. Small-Cap ETF Fund S000026636 · Registered fund · May 31, 2026 SCHWAB STRATEGIC TRUST, not summed	293,984	0.47%	+1,389	May 31, 2026	SCHWAB STRATEGIC TRUST, not summed
43.2	Schwab Small-Cap Index Fund Fund S000005912 · Registered fund · Jul 31, 2026 SCHWAB CAPITAL TRUST, not summed	150,971	0.24%	+18,172	Jul 31, 2026	SCHWAB CAPITAL TRUST, not summed
43.3	Schwab Total Stock Market Index Fund Fund S000005913 · Registered fund · Jul 31, 2026 SCHWAB CAPITAL TRUST, not summed	39,315	0.06%	+3,000	Jul 31, 2026	SCHWAB CAPITAL TRUST, not summed
43.4	Schwab U.S. Broad Market ETF Fund S000026631 · Registered fund · May 31, 2026 SCHWAB STRATEGIC TRUST, not summed	35,523	0.06%	+479	May 31, 2026	SCHWAB STRATEGIC TRUST, not summed
43.5	Schwab Small-Cap Equity Fund Fund S000005516 · Registered fund · Jul 31, 2026 SCHWAB CAPITAL TRUST, not summed	29,847	0.05%	-4,981	Jul 31, 2026	SCHWAB CAPITAL TRUST, not summed
44	BNP PARIBAS ARBITRAGE, SA 13F CIK 1166588 · Non-US institution · Jun 30, 2026 carried, not yet filed for the newest quarter	546,647	0.87%	-57,400	Jun 30, 2026	carried, not yet filed for the newest quarter
45	Nuveen, LLC 13F CIK 1871926 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	522,043	0.83%	+228,388	Jun 30, 2026	carried, not yet filed for the newest quarter
45.1	Nuveen Quant Small Cap Equity Fund Fund S000005386 · Registered fund · Jul 31, 2026 TIAA-CREF Funds, not summed	196,810	0.31%	+45,300	Jul 31, 2026	TIAA-CREF Funds, not summed
45.2	College Retirement Equities Fund - Total Global Stock Account Fund S000005080 · Registered fund · Jun 30, 2026 College Retirement Equities Fund, not summed	140,438	0.22%	—	Jun 30, 2026	College Retirement Equities Fund, not summed
45.3	Nuveen Small Cap Blend Index Fund Fund S000005367 · Registered fund · Jul 31, 2026 TIAA-CREF Funds, not summed	81,003	0.13%	+9,833	Jul 31, 2026	TIAA-CREF Funds, not summed
45.4	Nuveen Equity Index Fund Fund S000005360 · Registered fund · Jul 31, 2026 TIAA-CREF Funds, not summed	54,497	0.09%	0	Jul 31, 2026	TIAA-CREF Funds, not summed
45.5	Nuveen Small Cap Growth Opportunities Fund Fund S000005567 · Registered fund · Jul 31, 2026 Nuveen Investment Funds, Inc., not summed	10,332	0.02%	—	Jul 31, 2026	Nuveen Investment Funds, Inc., not summed
46	AMERICAN CENTURY COMPANIES INC 13F CIK 748054 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	487,031	0.78%	+449,193	Jun 30, 2026	carried, not yet filed for the newest quarter
46.1	Small Cap Growth Fund Fund S000006193 · Registered fund · Jul 31, 2026 American Century Mutual Funds, Inc., not summed	319,416	0.51%	—	Jul 31, 2026	American Century Mutual Funds, Inc., not summed
47	Tao Capital Management LP 13F CIK 1802967 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	478,815	0.76%	-350,000	Jun 30, 2026	carried, not yet filed for the newest quarter
48	NEW YORK STATE COMMON RETIREMENT FUND 13F CIK 810265 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	462,145	0.74%	-211,842	Jun 30, 2026	carried, not yet filed for the newest quarter
49	JENNISON ASSOCIATES LLC 13F CIK 53417 · Registered investment adviser · Jun 30, 2026 carried, not yet filed for the newest quarter	453,216	0.72%	-151,972	Jun 30, 2026	carried, not yet filed for the newest quarter
50	VANGUARD FIDUCIARY TRUST CO 13F CIK 933478 · Unknown · Jun 30, 2026 carried, not yet filed for the newest quarter	443,474	0.71%	+20,359	Jun 30, 2026	carried, not yet filed for the newest quarter

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Key facts CIK 1581280 CUSIP 90184D100 13F (30d) 30 filings 30 filers Visit website Investor relations