DNLI · Denali Therapeutics Inc. · Shareholders & Ownership
Every disclosed holder of DNLI in one list: 13F institutions, Schedule 13D and 13G filers, insiders (Forms 3, 4 and 5) and registered funds, each from its own latest filing. Holders nest by CIK and SEC reporting links, never by name, and each is summed once.
Who Owns DNLI
13F quarter Sep 30, 2026, combined view: filers that have not filed yet carry their Jun 30, 2026 positionsNo chart: the summed disclosures exceed shares outstanding, so they are not parts of one whole.
| Holder group | Shares | % Out |
|---|---|---|
| Institutional (13F) | 156,264,973 | 97.76% |
| Insiders (Forms 3, 4, 5) | 7,074,471 | 4.43% |
| 13D/G-only blocks | 12,906,125 | 8.07% |
| Remaining | 16,398,166 | -10.26% |
| Registered funds (N-PORT, not summed) | 65,147,756 | 40.76% |
Shares outstanding: 159,847,403, the issuer's latest cover-page count restated onto today's split basis.
Remaining is negative: the summed disclosures exceed shares outstanding, as when filings are dated at different times or holders that do not disclose their links count the same shares. It is flagged, not clipped.
13D/G-only counts each filer's latest 13D or 13G that no 13F report of the listing contains, that names no 13F filer and that no counted insider files. Funds are never summed: their shares sit inside their advisers' 13F reports.
Holders of DNLI
| # | Holder | Shares | As of | Notes |
|---|---|---|---|---|
| 1 |
T. Rowe Price Investment Management, Inc.
13F
CIK 1897612 · Investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
16,178,963 | ||
| 1.1 |
T. Rowe Price Investment Management, Inc.
13G
CIK 1897612 · Investment adviser · Jun 30, 2026
Rule 13d-1(b), qualified institution, not summed
|
16,178,963 | ||
| 2 |
BlackRock, Inc.
13F
CIK 2012383 · Holding company · Jun 30, 2026
carried, not yet filed for the newest quarter
|
14,026,413 | ||
| 2.1 |
iShares Russell 2000 ETF
Fund
S000004344 · Registered fund · Jun 30, 2026
iShares Trust, not summed
|
3,804,692 | ||
| 2.2 |
iShares Russell 2000 Value ETF
Fund
S000004342 · Registered fund · Jun 30, 2026
iShares Trust, not summed
|
1,302,420 | ||
| 2.3 |
iShares Biotechnology ETF
Fund
S000004350 · Registered fund · Jun 30, 2026
iShares Trust, not summed
|
1,054,236 | ||
| 2.4 |
Master Small Cap Index Series
Fund
S000002716 · Registered fund · Jun 30, 2026
Quantitative Master Series LLC, not summed
|
273,130 | ||
| 2.5 |
iShares U.S. Small-Cap Equity Factor ETF
Fund
S000048624 · Registered fund · Jul 31, 2026
iShares Trust, not summed
|
253,103 | ||
| 3 |
BAILLIE GIFFORD & CO
13F
CIK 1088875 · Investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
11,960,230 | ||
| 3.1 |
BAILLIE GIFFORD & CO
13G
CIK 1088875 · Investment adviser · Jun 30, 2026
Rule 13d-1(b), qualified institution, not summed
|
11,960,230 | ||
| 4 |
|
7,155,243 | ||
| 5 |
|
7,049,304 | ||
| 5.1 |
State Street(R) SPDR(R) S&P(R) Biotech ETF
Fund
S000010018 · Registered fund · Jun 30, 2026
SPDR SERIES TRUST, not summed
|
3,475,517 | ||
| 5.2 |
STATE STREET SMALL/MID CAP EQUITY INDEX PORTFOLIO
Fund
S000050332 · Registered fund · Jun 30, 2026
STATE STREET INSTITUTIONAL INVESTMENT TRUST, not summed
|
50,432 | ||
| 5.3 |
State Street Total Return V.I.S. Fund
Fund
S000001637 · Registered fund · Jun 30, 2026
STATE STREET VARIABLE INSURANCE SERIES FUNDS INC, not summed
|
3,713 | ||
| 5.4 |
State Street Balanced Index Fund
Fund
S000082802 · Registered fund · Jun 30, 2026
STATE STREET INSTITUTIONAL INVESTMENT TRUST, not summed
|
750 | ||
| 6 |
Temasek Holdings (Private) Ltd
13F
CIK 1021944 · Holding company · Jun 30, 2026
carried, not yet filed for the newest quarter
|
7,012,974 | ||
| 7 |
VANGUARD PORTFOLIO MANAGEMENT LLC
13F
CIK 2100121 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
6,477,057 | ||
| 8 |
VANGUARD CAPITAL MANAGEMENT LLC
13F
CIK 2100119 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
6,275,729 | ||
| 9 |
|
5,750,882 | ||
| 10 |
FMR LLC
13F
CIK 315066 · Holding company · Jun 30, 2026
carried, not yet filed for the newest quarter
|
4,535,836 | ||
| 10.1 |
Fidelity Small Cap Index Fund
Fund
S000033638 · Registered fund · Jul 31, 2026
Fidelity Salem Street Trust, not summed
|
1,608,716 | ||
| 10.2 |
Fidelity Growth Company Fund
Fund
S000007119 · Registered fund · May 31, 2026
Fidelity Mt. Vernon Street Trust, not summed
|
1,454,450 | ||
| 10.3 |
Fidelity Extended Market Index Fund
Fund
S000006021 · Registered fund · May 31, 2026
Fidelity Concord Street Trust, not summed
|
768,130 | ||
| 10.4 |
Fidelity Growth Company K6 Fund
Fund
S000065696 · Registered fund · May 31, 2026
Fidelity Mt. Vernon Street Trust, not summed
|
413,611 | ||
| 10.5 |
Fidelity Total Market Index Fund
Fund
S000006026 · Registered fund · May 31, 2026
Fidelity Concord Street Trust, not summed
|
293,168 | ||
| 11 |
VANGUARD TOTAL STOCK MARKET INDEX FUND
Fund
S000002848 · Registered fund · Jun 30, 2026
VANGUARD INDEX FUNDS, not summed
|
4,288,391 | ||
| 12 |
GOLDMAN SACHS GROUP INC
13F
CIK 886982 · Holding company · Jun 30, 2026
carried, not yet filed for the newest quarter
|
4,155,895 | ||
| 12.1 |
Goldman Sachs Small Cap Value Insights Fund
Fund
S000014929 · Registered fund · Jul 31, 2026
Goldman Sachs Trust, not summed
|
399,392 | ||
| 12.2 |
Goldman Sachs Small Cap Equity Insights Fund
Fund
S000009329 · Registered fund · Jul 31, 2026
Goldman Sachs Trust, not summed
|
256,894 | ||
| 12.3 |
Goldman Sachs U.S. Tax-Managed Equity Fund
Fund
S000009340 · Registered fund · Jun 30, 2026
Goldman Sachs Trust, not summed
|
77,256 | ||
| 12.4 |
Goldman Sachs Variable Insurance Trust Small Cap Equity Insights Fund
Fund
S000009355 · Registered fund · Jun 30, 2026
Goldman Sachs Variable Insurance Trust, not summed
|
57,608 | ||
| 12.5 |
Goldman Sachs Small Cap Growth Insights Fund
Fund
S000014930 · Registered fund · Jul 31, 2026
Goldman Sachs Trust, not summed
|
49,836 | ||
| 13 |
GEODE CAPITAL MANAGEMENT, LLC
13F
CIK 1214717 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
3,973,342 | ||
| 14 |
Holocene Advisors, LP
13F
CIK 1700574 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
3,905,225 | ||
| 15 |
BAKER BROS. ADVISORS LP
13F
CIK 1263508 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
3,734,336 | ||
| 15.1 |
BAKER BROS. ADVISORS LP
Insider
CIK 1263508 · Jun 3, 2026
Director, 10% Owner, issuer-wide, not summed
|
3,422,366 | ||
| 16 |
UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC
13F
CIK 861177 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
3,161,972 | ||
| 17 |
PRICE T ROWE ASSOCIATES INC /MD/
13F
CIK 80255 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
3,136,149 | ||
| 17.1 |
T. Rowe Price Capital Appreciation Fund
Fund
S000002070 · Registered fund · Jun 30, 2026
T. ROWE PRICE CAPITAL APPRECIATION FUND, INC., not summed
|
4,966,700 | ||
| 17.2 |
T. Rowe Price Small-Cap Stock Fund, Inc.
Fund
S000002120 · Registered fund · Jun 30, 2026
T. ROWE PRICE SMALL-CAP STOCK FUND, INC., not summed
|
2,328,096 | ||
| 17.3 |
T. Rowe Price Institutional Small-Cap Stock Fund
Fund
S000002094 · Registered fund · Jun 30, 2026
T. Rowe Price Equity Funds, Inc., not summed
|
1,274,515 | ||
| 17.4 |
T. Rowe Price Small-Cap Value Fund, Inc.
Fund
S000002121 · Registered fund · Jun 30, 2026
T. ROWE PRICE SMALL-CAP VALUE FUND, INC., not summed
|
1,251,030 | ||
| 17.5 |
T. Rowe Price New Horizons Fund, Inc.
Fund
S000002106 · Registered fund · Jun 30, 2026
T. ROWE PRICE NEW HORIZONS FUND, INC., not summed
|
711,908 | ||
| 18 |
|
3,120,592 | ||
| 19 |
VANGUARD SMALL-CAP INDEX FUND
Fund
S000002845 · Registered fund · Jun 30, 2026
VANGUARD INDEX FUNDS, not summed
|
3,097,183 | ||
| 20 |
VANGUARD U.S. GROWTH FUND
Fund
S000004444 · Registered fund · May 31, 2026
VANGUARD WORLD FUND, not summed
|
3,037,857 | ||
| 21 |
ALLIANCEBERNSTEIN L.P.
13F
CIK 1109448 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
2,984,807 | ||
| 21.1 |
AB Small Cap Growth Portfolio
Fund
S000010309 · Registered fund · Jun 30, 2026
AB CAP FUND, INC., not summed
|
691,817 | ||
| 21.2 |
AB DISCOVERY GROWTH FUND, INC.
Fund
S000010117 · Registered fund · Jun 30, 2026
AB DISCOVERY GROWTH FUND, INC., not summed
|
650,814 | ||
| 21.3 |
Small Cap Core Portfolio
Fund
S000051867 · Registered fund · Jun 30, 2026
Bernstein Fund Inc, not summed
|
179,900 | ||
| 21.4 |
AB Small Cap Growth Portfolio
Fund
S000010435 · Registered fund · Jun 30, 2026
AB VARIABLE PRODUCTS SERIES FUND, INC., not summed
|
13,143 | ||
| 21.5 |
Tax-Aware Overlay A Portfolio
Fund
S000027648 · Registered fund · Jun 30, 2026
BERNSTEIN SANFORD C FUND INC, not summed
|
4,521 | ||
| 22 |
DIMENSIONAL FUND ADVISORS LP
13F
CIK 354204 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
2,860,604 | ||
| 22.1 |
U.S. SMALL CAP PORTFOLIO
Fund
S000000971 · Registered fund · Jul 31, 2026
DFA INVESTMENT DIMENSIONS GROUP INC, not summed
|
679,330 | ||
| 22.2 |
U.S. MICRO CAP PORTFOLIO
Fund
S000000948 · Registered fund · Jul 31, 2026
DFA INVESTMENT DIMENSIONS GROUP INC, not summed
|
506,524 | ||
| 22.3 |
Dimensional U.S. Small Cap ETF
Fund
S000070906 · Registered fund · Jul 31, 2026
Dimensional ETF Trust, not summed
|
456,915 | ||
| 22.4 |
Dimensional U.S. Core Equity 2 ETF
Fund
S000070903 · Registered fund · Jul 31, 2026
Dimensional ETF Trust, not summed
|
238,844 | ||
| 22.5 |
U.S. CORE EQUITY 2 PORTFOLIO
Fund
S000000980 · Registered fund · Jul 31, 2026
DFA INVESTMENT DIMENSIONS GROUP INC, not summed
|
169,195 | ||
| 23 |
NORGES BANK
13F
CIK 1374170 · Non-US institution · Jun 30, 2026
carried, not yet filed for the newest quarter
|
2,824,746 | ||
| 24 |
Pictet Asset Management Holding SA
13F
CIK 1993888 · Unknown · Jun 30, 2026
carried, not yet filed for the newest quarter
|
2,791,206 | ||
| 25 |
Capital Research Global Investors
13F
CIK 1422848 · Investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
2,565,817 | ||
| 26 |
SMALLCAP World Fund Inc
Fund
S000009599 · Registered fund · Jun 30, 2026
SMALLCAP World Fund Inc, not summed
|
2,496,890 | ||
| 27 |
|
2,417,217 | ||
| 28 |
Fisher Asset Management, LLC
13F
CIK 850529 · Investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
2,414,162 | ||
| 29 |
NOMURA ASSET MANAGEMENT INTERNATIONAL INC.
13F
CIK 921739 · Investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
2,208,263 | ||
| 29.1 |
Nomura Small Cap Core Fund
Fund
S000002400 · Registered fund · May 29, 2026
Delaware Group Equity Funds V, not summed
|
727,399 | ||
| 29.2 |
Optimum Small-Mid Cap Growth Fund
Fund
S000002424 · Registered fund · Jun 30, 2026
Optimum Fund Trust, not summed
|
137,860 | ||
| 30 |
Casdin Capital, LLC
13F
CIK 1534261 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
2,045,000 | ||
| 31 |
|
1,965,424 | ||
| 32 |
Crestline Management, L.P.
13F
CIK 1536054 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
1,910,732 | ||
| 33 |
VANGUARD SMALL-CAP GROWTH INDEX FUND
Fund
S000002846 · Registered fund · Jun 30, 2026
VANGUARD INDEX FUNDS, not summed
|
1,784,440 | ||
| 34 |
VANGUARD EXTENDED MARKET INDEX FUND
Fund
S000002841 · Registered fund · Jun 30, 2026
VANGUARD INDEX FUNDS, not summed
|
1,731,822 | ||
| 35 |
PRINCIPAL FINANCIAL GROUP INC
13F
CIK 1126328 · Insurance company · Jun 30, 2026
carried, not yet filed for the newest quarter
|
1,420,083 | ||
| 35.1 |
SmallCap Fund
Fund
S000007214 · Registered fund · Jul 31, 2026
Principal Funds, Inc, not summed
|
561,500 | ||
| 35.2 |
SmallCap Growth Fund I
Fund
S000007175 · Registered fund · Jul 31, 2026
Principal Funds, Inc, not summed
|
209,518 | ||
| 35.3 |
SmallCap Account
Fund
S000007675 · Registered fund · Jun 30, 2026
Principal Variable Contract Funds, Inc, not summed
|
51,300 | ||
| 35.4 |
SmallCap Value Fund II
Fund
S000007169 · Registered fund · Jul 31, 2026
Principal Funds, Inc, not summed
|
9,271 | ||
| 36 |
Foresite Capital Management VI LLC
13F
CIK 2013341 · Unknown · Jun 30, 2026
carried, not yet filed for the newest quarter
|
1,396,771 | ||
| 37 |
Siren, L.L.C.
13F
CIK 2005245 · Unknown · Jun 30, 2026
carried, not yet filed for the newest quarter
|
1,396,513 | ||
| 38 |
NORTHERN TRUST CORP
13F
CIK 73124 · Bank · Jun 30, 2026
carried, not yet filed for the newest quarter
|
1,389,765 | ||
| 38.1 |
Small Cap Index Fund
Fund
S000001245 · Registered fund · Jun 30, 2026
Northern Funds, not summed
|
74,380 | ||
| 38.2 |
Small Cap Core Fund
Fund
S000001244 · Registered fund · Jun 30, 2026
Northern Funds, not summed
|
13,153 | ||
| 38.3 |
FlexShares Morningstar US Market Factor Tilt Index Fund
Fund
S000033312 · Registered fund · Jul 31, 2026
FlexShares Trust, not summed
|
10,428 | ||
| 39 |
BNP PARIBAS ASSET MANAGEMENT Holding S.A.
13F
CIK 1520354 · Non-US institution · Jun 30, 2026
carried, not yet filed for the newest quarter
|
1,355,514 | ||
| 40 |
Aberdeen Group plc
13F
CIK 1716774 · Holding company · Jun 30, 2026
carried, not yet filed for the newest quarter
|
1,345,481 | ||
| 41 |
CHARLES SCHWAB INVESTMENT MANAGEMENT INC
13F
CIK 884546 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
1,244,955 | ||
| 41.1 |
Schwab U.S. Small-Cap ETF
Fund
S000026636 · Registered fund · May 31, 2026
SCHWAB STRATEGIC TRUST, not summed
|
712,909 | ||
| 41.2 |
Schwab Small-Cap Index Fund
Fund
S000005912 · Registered fund · Jul 31, 2026
SCHWAB CAPITAL TRUST, not summed
|
357,236 | ||
| 41.3 |
Schwab Total Stock Market Index Fund
Fund
S000005913 · Registered fund · Jul 31, 2026
SCHWAB CAPITAL TRUST, not summed
|
90,464 | ||
| 41.4 |
Schwab U.S. Broad Market ETF
Fund
S000026631 · Registered fund · May 31, 2026
SCHWAB STRATEGIC TRUST, not summed
|
85,423 | ||
| 42 |
BANK OF AMERICA CORP /DE/
13F
CIK 70858 · Bank · Jun 30, 2026
carried, not yet filed for the newest quarter
|
1,230,957 | ||
| 43 |
Artisan Partners Limited Partnership
13F
CIK 1466153 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
1,224,497 | ||
| 43.1 |
Artisan International Small-Mid Fund
Fund
S000006494 · Registered fund · Jun 30, 2026
Artisan Partners Funds Inc., not summed
|
753,697 | ||
| 44 |
Candriam S.C.A.
13F
CIK 1537014 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
1,189,886 | ||
| 45 |
MILLENNIUM MANAGEMENT LLC
13F
CIK 1273087 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
1,125,748 | ||
| 46 |
JNL/T. Rowe Price Capital Appreciation Fund
Fund
S000068115 · Registered fund · Jun 30, 2026
JNL Series Trust, not summed
|
1,084,400 | ||
| 47 |
ADAR1 Capital Management, LLC
13F
CIK 1940272 · Registered investment adviser · Jun 30, 2026
carried, not yet filed for the newest quarter
|
1,029,080 | ||
| 48 |
VANGUARD FIDUCIARY TRUST CO
13F
CIK 933478 · Unknown · Jun 30, 2026
carried, not yet filed for the newest quarter
|
991,904 | ||
| 49 |
MORGAN STANLEY
13F
CIK 895421 · Holding company · Jun 30, 2026
carried, not yet filed for the newest quarter
|
961,050 | ||
| 49.1 |
E*TRADE No Fee Total Market Index Fund
Fund
S000090722 · Registered fund · Jul 31, 2026
E Trade Trust, not summed
|
383 | ||
| 50 |
|
819,999 |
56 more nested rows are not shown here; Every filing lists them all.
Shares are restated onto today's split basis. Change compares each row with the same holder's previous filing in the same source; a dash means there is none to compare. Nested 13D/G, insider and fund rows are not summed, since a 13F report already holds them.
Showing 1–50 of 426 holders
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