LGND · Ligand Pharmaceuticals Inc · Holdings

$306.40 +9.12 (+3.07%) At close · Oct 20

S&P 600 member Russell 2000 member Rebalance outlook locked

Market Cap

$5.93B

19.94M

Volume · Oct 20 168.16K Avg daily vol (3M) 244.48K

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Use institutional holdings through the 13F API or MCP → See every holder of LGND: 13F, 13D and 13G, insiders and funds →

Institutional holdings are being reconciled with SEC filings. Holder totals and changes may be incomplete until reconciliation finishes.

The Sep 30, 2026 filing window is still open — showing the latest known positions. Funds that have not filed yet carry their prior-quarter positions; a fund that filed without this stock counts as sold out. Each fund's Change is measured against its own previous reported quarter, shown in the Reported column. Pick a completed quarter in the Quarter selector below to put every fund on one quarter and see the stock-wide activity breakdown.

Quarter:

Holders: 424

Total Value: $7,166,481,236

Total Shares: 22,700,549

Showing 1–100 of 432 holdings by value · page 1 of 5

Institutional Ownership Trend

27 quarters

Superinvestors Holding LGND

All superinvestors →

Joel Greenblatt $1,141,084

Holder Concentration

Across 422 13F filers

Top Holder

13.7%

of institutional value

Top 5

32.4%

of institutional value

Top 10

48.2%

of institutional value

HHI

374

Broadly held

Who Holds LGND

Last 8 quarters through Sep 30, 2026 (filing window open)

Each kind of 13F filer's share of the common shares 13F filers report. Holder type comes from regulatory records, never the filer's name; style, turnover and horizon are measured from each filer's own 13F book. The quarter selector does not change this card.

Jun 30, 2026, the newest closed quarter: 418 filers, like-for-like change +8.7%; actively managed books +10.7%; index-like books +4.9%.

Holder type	Share	Filers	Change	New	Exited
Registered adviser	40.9%	229	+7.1%	61	21
Exempt adviser	0.0%	2	-82.2%	2	2
Adviser	13.7%	12	+25.4%	2	1
Bank	8.3%	23	+2.8%	3	0
Insurer	0.7%	2	-4.5%	0	0
Broker-dealer	0.1%	2	+28.2%	0	0
Benefit plan	0.2%	5	-7.2%	0	1
Non-US	2.5%	11	-8.7%	1	1
Holding company	22.5%	12	+13.6%	0	1
Unknown	11.1%	120	+1.0%	25	7

Style	Share	Filers	Change	New	Exited
Index-like	32.6%	40	+4.9%	1	1
Systematic-like	0.6%	14	-34.6%	6	7
Concentrated	0.3%	49	+141.4%	19	10
Diversified active	66.6%	315	+11.1%	68	16
Not measured	0.0%	0	—	0	0
Unknown	0.0%	0	—	0	0

Turnover	Share	Filers	Change	New	Exited
Low turnover	63.2%	261	+9.4%	53	12
Medium turnover	18.5%	100	+14.6%	19	9
High turnover	1.2%	26	-17.2%	12	12
Unknown	17.1%	31	+3.0%	10	1
Not measured	0.0%	0	—	0	0

Holding horizon	Share	Filers	Change	New	Exited
Short horizon	17.9%	44	+5.2%	18	5
Medium horizon	15.1%	74	+5.0%	21	8
Long horizon	62.6%	277	+9.8%	48	15
Unknown	4.4%	23	+24.0%	7	6
Not measured	0.0%	0	—	0	0

13F book size	Share	Filers	Change	New	Exited
Under $1B	2.7%	89	+0.9%	33	11
$1B to $10B	13.5%	139	+12.6%	35	9
$10B to $100B	15.5%	121	+5.1%	22	11
$100B or more	68.3%	69	+9.2%	4	3
Not measured	0.0%	0	—	0	0

Client base	Share	Filers	Change	New	Exited
Individuals and high net worth	6.8%	123	+1.7%	36	13
Pooled investment vehicles	1.0%	20	+107.5%	11	7
Registered funds and BDCs	20.6%	31	+1.8%	3	0
Pension and profit sharing plans	0.2%	5	+26.3%	3	0
Insurance companies	0.6%	3	+806.2%	0	0
Charitable organizations	0.0%	1	0.0%	0	0
Governments and sovereign funds	1.1%	6	+10.8%	1	0
Banks and thrifts	0.0%	1	-1.1%	0	0
Corporations	1.2%	2	-3.7%	0	0
Other investment advisers	0.1%	8	+13.2%	2	0
Other	0.3%	4	+6.8%	2	0
Diversified	9.0%	25	+12.7%	3	1
Unknown	59.1%	189	+9.9%	33	13

Private funds	Share	Filers	Change	New	Exited
Hedge funds	4.1%	42	+12.8%	14	9
Private equity funds	1.3%	11	-13.8%	2	1
Venture capital funds	0.0%	3	+134.0%	1	0
Real estate funds	0.0%	7	+0.7%	2	0
Securitized asset funds	0.9%	1	-1.2%	0	0
Other private funds	5.6%	19	+3.1%	4	1
Mixed private funds	0.4%	3	+2.9%	0	0
Private funds, type not stated	1.9%	4	-1.6%	2	0
No private funds	26.7%	141	+8.6%	38	12
Unknown	59.1%	187	+10.4%	31	11

Location	Share	Filers	Change	New	Exited
United States	93.5%	363	+10.4%	80	28
Outside the US	6.5%	54	-10.7%	14	6
Unknown	0.0%	1	0.0%	0	0

The Sep 30, 2026 filing window is still open, so its bar carries the Jun 30, 2026 positions of filers that have not filed yet.

How the mix is measured

Shares are 13F common shares restated to today's split basis; options and principal amounts are left out. The change compares only filers with a 13F in both quarters and keeps each filer in its bucket of the later quarter on both sides; a filer that continues under a new CIK is compared through its successor. Holder type comes from each filer's current regulatory profile; client base and private funds come from the Form ADV roster current at each quarter end (rosters from December 2019, so an earlier quarter reads Unknown). Style, turnover, horizon and book size are measured from each filer's 13F book in the newest measured quarter, and Not measured means none was. Actively managed books are those measured systematic-like, concentrated or diversified active.

Excluded 2 holders with no observed 2026-03-31 13F and 4 holders with no observed 2026-06-30 13F. Compared across CIK changes: JANUS HENDERSON GROUP PLC continues as Jupiter Topco LLC.

Conviction and Crowding

417 13F filers, quarter ended Jun 30, 2026

Measured across every 13F filer's common-share book once the quarter's filing window closed. Crowding counts filers with high turnover or that advise private funds on Form ADV.

Conviction holders

3.36% of filers hold it in their top 10

Mar 31, 2026: 9

Holders above 5%

median weight 0.008%

Mar 31, 2026: 1

Crowding

—

holders exceed shares outstanding: 13F filers report more shares than the cover-page count, which predates an issuance, is misstated, or misses shares lent and reported twice

Private-fund advisers

—

of shares outstanding

Institutional ownership

—

holders exceed shares outstanding: 13F filers report more shares than the cover-page count, which predates an issuance, is misstated, or misses shares lent and reported twice

Top 10 holders

48.4%

of 13F shares, Herfindahl 0.038

Active holders

+10.7%

like-for-like share change of active filers

5% stakes on record

0 13D, 7 13G (7 institutional, 0 passive, 0 exempt)

How long holders have held it value-weighted 14.7 quarters

Under 1 year 21.3% 1 to 3 years 29.7% 3 years or more 49.0% Since coverage start 0.000%

Managers in the top fifth by current three-year FundScore alpha were net buyers of $0 (0 buying, 0 selling).

Estimated Holder Cost Basis

Quarter ended Jun 30, 2026, on that quarter's share basis

Estimated, never filed: each 13F buy is costed at its quarter's average traded price and each sale keeps the average cost, so this is what holders paid on average if they traded evenly through each quarter.

Estimated avg cost

$149.91

estimated per share

Mar 31, 2026: $123.13

Quarter-end price

$316.09

implied by 13F values

Estimated gain or loss

+110.9%

price against the estimated cost

Estimated at a loss

0.000%

of costed 13F value

Estimated cost covers 28.2% of the 13F shares; positions already held when a filer's 13F history starts, or bought in a quarter without a trusted average price, have no estimate.

Shared Holders

Companies LGND's active holders also own, quarter ended Jun 30, 2026

#	Company	Shared holders	Similarity
1	HRI Herc Holdings Inc	10	0.202
2	FROG JFrog Ltd	14	0.200
3	STRL Sterling Infrastructure, Inc.	23	0.176
4	OFRM Once Upon a Farm, PBC	4	0.169
5	XMTR Xometry, Inc.	10	0.167
6	BLFS Biolife Solutions Inc	7	0.167
7	KNSA Kiniksa Pharmaceuticals International, plc	8	0.166
8	GHM Graham Corp	8	0.166
9	UFPT Ufp Technologies Inc	8	0.166
10	NOVT Novanta Inc	11	0.162

Shareholder Targets

13F filers that own its peers but not LGND, quarter ended Jun 30, 2026

Found through its holder-overlap peers: HRI, FROG, STRL, OFRM, XMTR, BLFS, KNSA, GHM, UFPT, NOVT, AGYS, OUST, LQDT, TVTX, HALO, VSEC, COCO, UCTT, BOOT, KRUS. 1,329 filers in all.

#	Filer	In peers	Of book	Largest peers
1	RENAISSANCE TECHNOLOGIES LLC Registered adviser, Diversified active	$1,021,101,791 12 held	1.42%	STRL, FROG, TVTX
2	T. Rowe Price Investment Management, Inc. Adviser, Diversified active	$982,065,972 7 held	0.650%	VSEC, BLFS, NOVT
3	JANE STREET GROUP, LLC Unknown, Diversified active	$967,872,104 12 held	0.640%	STRL, XMTR, COCO
4	WASATCH ADVISORS LP Registered adviser, Diversified active	$830,972,322 7 held	5.44%	FROG, UFPT, COCO
5	NORGES BANK Non-US, Index-like	$800,410,839 8 held	0.080%	HRI, STRL, FROG
6	PRIMECAP MANAGEMENT CO/CA/ Registered adviser, Diversified active	$659,926,904 3 held	0.390%	XMTR, VSEC, BOOT
7	FULLER & THALER ASSET MANAGEMENT, INC. Registered adviser, Diversified active	$636,447,905 9 held	1.74%	XMTR, VSEC, HALO
8	Capital World Investors Adviser, Diversified active	$633,147,325 3 held	0.080%	UCTT, VSEC, STRL
9	WILLIAM BLAIR INVESTMENT MANAGEMENT, LLC Registered adviser, Diversified active	$599,026,668 5 held	1.79%	HALO, STRL, NOVT
10	ARROWSTREET CAPITAL, LIMITED PARTNERSHIP Registered adviser, Diversified active	$543,075,445 9 held	0.240%	STRL, FROG, HALO

Fund Ownership of LGND by Fund Type

Registered funds' Form N-PORT reports for the quarter ended Jun 30, 2026

Index funds and index ETFs hold 35.85% of shares outstanding.

Fund type	Of shares outstanding	Change	Shares	Funds
Index funds	5.10%	+0.69 pp	1,016,493	82
Index ETFs	30.75%	+7.18 pp	6,129,524	82
Active ETFs	0.86%	-0.34 pp	170,863	22
Active funds	31.01%	+5.19 pp	6,182,237	186
Unclassified	0.42%	+0.32 pp	83,456	15
All registered funds	68.13%	+13.03 pp	13,582,573	387

How fund types are measured

Each fund series counts once, with its latest Form N-PORT report dated inside the quarter; only long share positions count, restated to today's split basis. The fund type is the series' latest Form N-CEN answer on or before the quarter end (the index-fund box and the exchange-traded flag), never its name. Unclassified funds have no usable N-CEN; unit investment trusts are among them, so index ownership is a floor. Funds sit inside their advisers' 13F reports, so these shares overlap the 13F figures and are never added to them.

Holder type: Registered adviser 234 Exempt adviser 2 Adviser 12 Bank 22 Insurer 2 Broker-dealer 2 Benefit plan 5 Non-US 11 Holding company 12 Unknown 122

Showing 100 of 432

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

BlackRock, Inc.

$980,193,819

13.68%

3,100,996

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	15,792	$4,991,693	0.5%	Sole
BlackRock Investment Management (Australia) Ltd	32	$10,114	0.0%	Sole
BlackRock Asset Management Canada Ltd	116	$36,666	0.0%	Sole
BlackRock Investment Management, LLC	62,557	$19,773,642	2.0%	Sole
BLACKROCK ADVISORS LLC	46,798	$14,792,379	1.5%	Sole
BlackRock Fund Advisors	2,224,803	$703,237,980	71.7%	Sole
BlackRock Institutional Trust Company, N.A.	590,800	$186,745,972	19.1%	Sole
BlackRock Fund Managers Ltd.	3,467	$1,095,884	0.1%	Sole
BlackRock Investment Management (UK) Ltd.	6,619	$2,092,199	0.2%	Sole
BlackRock (Netherlands) B.V.	5,417	$1,712,259	0.2%	Sole
BlackRock Asset Management Ireland Ltd	92,820	$29,339,473	3.0%	Sole
BlackRock (Luxembourg) S.A.	67	$21,178	0.0%	Sole
BlackRock Asset Management Schweiz AG	3,311	$1,046,573	0.1%	Sole
Aperio Group, LLC	48,382	$15,293,066	1.6%	Sole
SpiderRock Advisors, LLC	15	$4,741	0.0%	Sole

Invesco Ltd.

$379,827,645

5.30%

1,201,644

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	892,041	$281,965,235	74.2%	Defined
Invesco Trust Co joint	7,993	$2,526,507	0.7%	Defined
Invesco Asset Management Limited joint	28,044	$8,864,427	2.3%	Defined
Invesco Capital Management LLC	273,566	$86,471,476	22.8%	Defined

VANGUARD PORTFOLIO MANAGEMENT LLC

$359,810,304

5.02%

1,138,316

Jupiter Topco LLC

$328,916,298

4.59%

1,040,578

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	91,920	$29,054,992	8.8%	Defined
JANUS HENDERSON INVESTORS US LLC	941,848	$297,708,734	90.5%	Defined
Held directly	6,810	$2,152,572	0.7%	Defined

VANGUARD CAPITAL MANAGEMENT LLC

$275,298,585

3.84%

870,950

STATE STREET CORP

$261,260,712

3.65%

826,539

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	826,539	$261,260,712	100.0%	Defined

FMR LLC

$241,121,985

3.36%

762,827

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	733,137	$231,737,274	96.1%	Defined
STRATEGIC ADVISERS LLC	1,781	$562,956	0.2%	Defined
FIAM LLC	27,909	$8,821,755	3.7%	Defined

FRANKLIN RESOURCES INC

$237,713,585

3.32%

752,044

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FIDUCIARY TRUST CO INTERNATIONAL	96	$30,344	0.0%	Defined
FIDUCIARY TRUST INTERNATIONAL LLC	317	$100,200	0.0%	Defined
FRANKLIN ADVISERS INC	593,005	$187,442,950	78.9%	Defined
Franklin Templeton International Services S.a r.l	80	$25,287	0.0%	Defined
PUTNAM INVESTMENT MANAGEMENT LLC	101,464	$32,071,755	13.5%	Defined
PUTNAM ADVISORY CO LLC	57,082	$18,043,049	7.6%	Defined

CONGRESS ASSET MANAGEMENT CO

$205,713,137

2.87%

658,556

GEODE CAPITAL MANAGEMENT, LLC

$186,183,331

2.60%

589,020

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	73,880	$23,352,729	12.5%	Defined
Held directly	515,140	$162,830,602	87.5%	Defined

NOMURA ASSET MANAGEMENT INTERNATIONAL INC.

$179,656,389

2.51%

568,371

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NOMURA HOLDINGS INC joint	568,371	$179,656,389	100.0%	Defined

Chicago Capital, LLC

$172,186,153

2.40%

551,225

Stephens Investment Management Group LLC

$150,335,248

2.10%

475,609

DIMENSIONAL FUND ADVISORS LP

$135,701,859

1.89%

429,314

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	1,941	$613,530	0.5%	Defined
DFA Australia Ltd.	5,215	$1,648,409	1.2%	Defined
Dimensional Fund Advisors Pte. Ltd.	131	$41,407	0.0%	Defined
Dimensional Ireland Ltd	11,763	$3,718,166	2.7%	Defined
Held directly	410,264	$129,680,347	95.6%	Sole

MASSACHUSETTS FINANCIAL SERVICES CO /MA/

$118,774,610

1.66%

375,762

GOLDMAN SACHS GROUP INC

$102,174,510

1.43%

323,245

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	81,443	$25,743,317	25.2%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	237,838	$75,178,213	73.6%	Defined
GOLDMAN SACHS INTERNATIONAL	1,059	$334,739	0.3%	Defined
Goldman Sachs Trust Company, N.A. joint	2,905	$918,241	0.9%	Defined

LOOMIS SAYLES & CO L P

$92,236,641

1.29%

291,805

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
LOOMIS SAYLES TRUST COMPANY, LLC	49,161	$15,539,300	16.8%	Defined
Held directly	242,644	$76,697,341	83.2%	Defined

RICE HALL JAMES & ASSOCIATES, LLC

$85,360,736

1.19%

270,052

Ranger Investment Management, L.P.

$81,423,519

1.14%

257,596

NORTHERN TRUST CORP

$81,358,086

1.14%

257,389

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	233,141	$73,693,538	90.6%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	14,433	$4,562,126	5.6%	Defined
MASON STREET ADVISORS, LLC joint	6,698	$2,117,170	2.6%	Other
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	3,117	$985,252	1.2%	Other

VICTORY CAPITAL MANAGEMENT INC

$80,212,262

1.12%

253,764

DEUTSCHE BANK AG\

$78,899,539

1.10%

249,611

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Deutsche Bank Trust Co Americas	225	$71,120	0.1%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC	226,446	$71,577,316	90.7%	Defined
DWS Investments UK Ltd	22,364	$7,069,036	9.0%	Defined
DWS International GmbH	576	$182,067	0.2%	Defined

NEUBERGER BERMAN GROUP LLC

$78,502,531

1.10%

248,355

GW&K Investment Management, LLC

$78,037,246

1.09%

246,883

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$66,513,866

0.93%

210,427

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	2,454	$775,684	1.2%	Defined
UBS Asset Management Switzerland AG	102,671	$32,453,276	48.8%	Sole
UBS Asset Management (UK) Ltd.	5,369	$1,697,085	2.6%	Defined
Held directly	99,933	$31,587,821	47.5%	Defined

MORGAN STANLEY

$63,438,943

0.89%

200,699

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	243	$76,809	0.1%	Defined
MORGAN STANLEY & CO. LLC	6,711	$2,121,279	3.3%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	73,525	$23,240,517	36.6%	Defined
MORGAN STANLEY SMITH BARNEY LLC	27,451	$8,676,986	13.7%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	92,769	$29,323,352	46.2%	Defined

NEXT CENTURY GROWTH INVESTORS LLC

$62,757,140

0.88%

198,542

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$61,080,599

0.85%

193,238

Aberdeen Group plc

$54,519,201

0.76%

172,480

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
abrdn Inc.	96,697	$30,564,954	56.1%	Defined
abrdn Investment Management Ltd	74,751	$23,628,043	43.3%	Defined
Held directly	1,032	$326,204	0.6%	Defined

BANK OF AMERICA CORP /DE/

$53,845,614

0.75%

170,349

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	43,494	$13,748,018	25.5%	Defined
MERRILL LYNCH, PIERCE, FENNER & SMITH INC.	12,959	$4,096,210	7.6%	Defined
BOFA SECURITIES, INC.	2,346	$741,547	1.4%	Defined
MANAGED ACCOUNT ADVISORS LLC joint	111,550	$35,259,839	65.5%	Defined

ASHFORD CAPITAL MANAGEMENT INC

$53,809,897

0.75%

170,236

Private Capital Management, LLC

$52,475,997

0.73%

166,016

VILLERE ST DENIS J & CO LLC

$49,153,891

0.69%

155,506

Granahan Investment Management, LLC

$46,538,246

0.65%

147,231

VANGUARD FIDUCIARY TRUST CO

$45,177,479

0.63%

142,926

BESSEMER GROUP INC

$44,708,085

0.62%

141,449

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BESSEMER TRUST CO	2,177	$688,127	1.5%	Defined
BESSEMER TRUST Co NATIONAL ASSOCIATION	119,711	$37,839,449	84.6%	Defined
Bessemer Trust CO of Delaware, N.A.	9,002	$2,845,442	6.4%	Defined
BESSEMER TRUST Co OF FLORIDA	10,130	$3,201,991	7.2%	Defined
Bessemer Trust Co of Nevada, National Association	429	$133,076	0.3%	Defined

NEW YORK STATE COMMON RETIREMENT FUND

$43,385,881

0.61%

137,258

Bank of New York Mellon Corp

$41,429,278

0.58%

131,068

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNY Mellon Trust of Delaware joint	1,220	$385,629	0.9%	Defined
Mellon Investments Corporation joint	63,111	$19,948,755	48.2%	Defined
BNY Mellon Investment Adviser, Inc.	16,265	$5,141,203	12.4%	Defined
The Bank of New York Mellon	37,714	$11,921,018	28.8%	Defined
BNY Mellon, NA joint	7,569	$2,392,484	5.8%	Defined
BNY Mellon Advisors, Inc. joint	4,822	$1,524,185	3.7%	Other
Newton Investment Management North America, LLC	366	$115,688	0.3%	Defined
Held directly	1	$316	0.0%	Defined

Impax Asset Management Group plc

$41,091,700

0.57%

130,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
IMPAX ASSET MANAGEMENT LLC	130,000	$41,091,700	100.0%	Defined

VOYA INVESTMENT MANAGEMENT LLC

$40,165,872

0.56%

127,071

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	127,071	$40,165,872	100.0%	Defined

UBS Group AG

$39,408,203

0.55%

124,674

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	6,840	$2,162,055	5.5%	Defined
UBS AG	97,314	$30,759,982	78.1%	Defined
UBS Switzerland AG	20,520	$6,486,166	16.5%	Defined

Fisher Asset Management, LLC

$38,385,969

0.54%

121,440

Russell Investments Group, Ltd.

$38,026,888

0.53%

120,304

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC	59,650	$18,854,768	49.6%	Defined
Russell Investments Canada Limited	275	$86,924	0.2%	Defined
Russell Investments Capital, LLC	32,844	$10,381,659	27.3%	Defined
Russell Investments France SAS	9,914	$3,133,716	8.2%	Defined
Russell Investments Limited	380	$120,114	0.3%	Defined
Russell Investments Trust Company	17,241	$5,449,707	14.3%	Defined

ROYAL BANK OF CANADA

$37,637,782

0.53%

119,073

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC CMA LTD.	612	$193,447	0.5%	Defined
RBC Capital Markets, LLC	20,810	$6,577,832	17.5%	Defined
RBC Dominion Securities Inc.	78	$24,655	0.1%	Defined
RBC Trust Co (Delaware) Ltd	104	$32,873	0.1%	Defined
RBC Rochdale, LLC	172	$54,367	0.1%	Defined
Held directly	97,297	$30,754,608	81.7%	Sole

Driehaus Capital Management LLC

$37,455,716

0.52%

118,497

Penn Capital Management Company, LLC

$31,715,522

0.44%

100,337

Legal & General Group Plc

$31,212,622

0.44%

98,746

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	30,702	$9,704,595	31.1%	Defined
Legal & General Investment Management Ltd joint	68,044	$21,508,027	68.9%	Defined

PRINCIPAL FINANCIAL GROUP INC

$30,967,653

0.43%

97,971

BNP PARIBAS ARBITRAGE, SA

$29,735,218

0.41%

94,072

PRICE T ROWE ASSOCIATES INC /MD/

$28,915,280

0.40%

91,478

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	28,551	$9,024,685	31.2%	Defined
Held directly	62,927	$19,890,595	68.8%	Sole

FIRST TRUST ADVISORS LP

$28,730,368

0.40%

90,893

OSAIC HOLDINGS, INC.

$28,522,697

0.40%

90,236

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
OSAIC WEALTH, INC.	13,456	$4,253,307	14.9%	Defined
CW Advisors, LLC	76,746	$24,258,643	85.1%	Sole
Osaic Advisory Services, LLC	34	$10,747	0.0%	Defined

SEI INVESTMENTS CO

$28,344,422

0.40%

89,672

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI INVESTMENTS MANAGEMENT CORP	3,259	$1,030,137	3.6%	Sole
SEI TRUST CO	86,413	$27,314,285	96.4%	Sole

Fiera Capital Corp

$24,448,928

0.34%

77,348

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fiera Capital Inc.	75,730	$23,937,495	97.9%	Other
Held directly	1,618	$511,433	2.1%	Sole

AMERIPRISE FINANCIAL INC

$24,085,108

0.34%

76,197

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	74,519	$23,554,710	97.8%	Defined
Ameriprise Financial Services, LLC	1,630	$515,226	2.1%	Defined
Ameriprise Bank, FSB	48	$15,172	0.1%	Defined

JPMORGAN CHASE & CO

$23,091,004

0.32%

73,052

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	10,323	$3,262,996	14.1%	Other
J.P. Morgan Investment Management Inc.	62,161	$19,648,470	85.1%	Defined
J.P. Morgan Securities LLC	466	$147,297	0.6%	Defined
J.P. Morgan Securities plc	102	$32,241	0.1%	Defined

Public Sector Pension Investment Board

$21,774,807

0.30%

68,888

CITIGROUP INC

$21,063,921

0.29%

66,639

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	66,639	$21,063,921	100.0%	Defined

CASTLEARK MANAGEMENT LLC

$19,611,171

0.27%

62,043

NOMURA HOLDINGS INC

$19,577,982

0.27%

61,938

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NOMURA GLOBAL FINANCIAL PRODUCTS INC	61,938	$19,577,982	100.0%	Defined

Dana Investment Advisors, Inc.

$18,181,496

0.25%

57,520

WELLS FARGO & COMPANY/MN

$18,108,161

0.25%

57,288

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	45,952	$14,524,967	80.2%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	2,318	$732,696	4.0%	Defined
WELLS FARGO CLEARING SERVICES, LLC	8,975	$2,836,907	15.7%	Defined
WELLS FARGO INVESTMENT INSTITUTE, INC.	43	$13,591	0.1%	Defined

D. E. Shaw & Co., Inc.

$17,863,826

0.25%

56,515

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	56,515	$17,863,826	100.0%	Defined

PRUDENTIAL FINANCIAL INC

$17,265,784

0.24%

54,623

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PGIM Quantitative Solutions LLC	45,523	$14,389,365	83.3%	Defined
Prudential Trust Co	9,100	$2,876,419	16.7%	Defined

Granite Investment Partners, LLC

$17,109,319

0.24%

54,128

Allspring Global Investments Holdings, LLC

$16,420,243

0.23%

51,948

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Allspring Funds Management, LLC reporting on behalf of Morgan Stanley Smith Barney, LLC	13	$4,109	0.0%	Defined
Allspring Global Investments, LLC reporting on behalf of UBS ASSET MANAGEMENT AMERICAS INC	51,935	$16,416,134	100.0%	Defined

RHUMBLINE ADVISERS

$15,844,011

0.22%

50,125

NORDEA INVESTMENT MANAGEMENT AB

$15,786,482

0.22%

49,943

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nordea Funds Ltd	11,863	$3,749,775	23.8%	Defined
Nordea Investment Funds S.A.	38,080	$12,036,707	76.2%	Defined

Emerald Investment Advisers, LLC

$14,831,258

0.21%

46,921

Nuveen, LLC

$14,744,649

0.21%

46,647

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	3,588	$1,134,130	7.7%	Defined
TEACHERS ADVISORS, LLC joint	43,059	$13,610,519	92.3%	Defined

Context Capital Management, LLC

$14,540,140

0.20%

46,000

SUSQUEHANNA ADVISORS GROUP, INC.

$14,097,614

0.20%

44,600

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CAPITAL VENTURES INTERNATIONAL	44,600	$14,097,614	100.0%	Other

WCM INVESTMENT MANAGEMENT, LLC

$14,088,131

0.20%

44,570

KORNITZER CAPITAL MANAGEMENT INC /KS

$14,076,119

0.20%

44,532

ENVESTNET ASSET MANAGEMENT INC

$12,754,231

0.18%

40,350

Pembroke Management, LTD

$12,731,156

0.18%

40,277

Swiss National Bank

$12,327,510

0.17%

39,000

ESSEX INVESTMENT MANAGEMENT CO LLC

$12,156,503

0.17%

38,917

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 29026	38,917	$12,156,503	100.0%	Sole

ARISTEIA CAPITAL, L.L.C.

$11,774,352

0.16%

37,250

STIFEL FINANCIAL CORP

$11,642,541

0.16%

36,833

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STIFEL NICOLAUS & CO INC \MO\	36,759	$11,619,152	99.8%	Defined
Stifel Independent Advisors, LLC	63	$19,913	0.2%	Defined
Held directly	11	$3,476	0.0%	Defined

CAPTRUST FINANCIAL ADVISORS

$10,938,926

0.15%

34,607

Uniplan Investment Counsel, Inc.

$10,825,766

0.15%

34,249

Universal- Beteiligungs- und Servicegesellschaft mbH

$10,699,962

0.15%

33,851

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Universal-Investment-Gesellschaft mbH joint	33,851	$10,699,962	100.0%	Defined

KENNEDY CAPITAL MANAGEMENT LLC

$10,542,549

0.15%

33,353

Clark Capital Management Group, Inc.

$10,414,533

0.15%

32,948

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$10,304,534

0.14%

32,600

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	32,600	$10,304,534	100.0%	Other

Moody Aldrich Partners LLC

$8,901,094

0.12%

28,160

Linden Advisors LP

$8,669,084

0.12%

27,426

ALLIANCEBERNSTEIN L.P.

$8,564,458

0.12%

27,095

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	27,095	$8,564,458	100.0%	Defined

Hodges Capital Management Inc.

$7,410,413

0.10%

23,444

Squarepoint Ops LLC

$7,206,852

0.10%

22,800

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Squarepoint Capital LLP	22,800	$7,206,852	100.0%	Defined

Calamos Advisors LLC

$7,091,478

0.10%

22,435

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC	5,232	$1,653,782	23.3%	Other
Held directly	17,203	$5,437,696	76.7%	Sole

BANK OF MONTREAL /CAN/

$7,054,494

0.10%

22,318

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BMO Asset Management Inc.	1,399	$442,209	6.3%	Defined
BMO Bank N.A.	710	$224,423	3.2%	Defined
BMO Family Office, LLC	1,738	$549,364	7.8%	Defined
Held directly	18,471	$5,838,498	82.8%	Defined

CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM

$6,970,732

0.10%

22,053

NEW YORK STATE TEACHERS RETIREMENT SYSTEM

$6,622,085

0.09%

20,950

ISTHMUS PARTNERS, LLC

$6,617,344

0.09%

20,935

Moran Wealth Management, LLC

$6,255,420

0.09%

19,790

LAZARD ASSET MANAGEMENT LLC

$6,214,013

0.09%

19,659

Informed Momentum Co LLC

$6,163,122

0.09%

19,498

PALISADE CAPITAL MANAGEMENT, LP

$6,023,727

0.08%

19,057

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Key facts CIK 886163 CUSIP 53220K504 13F (30d) 60 filings 59 filers Visit website Investor relations