INO · Inovio Pharmaceuticals, Inc. · Holdings

$1.30 +0.02 (+1.56%)

Market Cap

$132.35M

103.40M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Use institutional holdings through the 13F API or MCP →

Quarter:

Holders: 105

Total Value: $39,416,007

Total Shares: 35,832,780

Showing 1–100 of 111 holdings by value · page 1 of 2

Institutional Ownership Trend

26 quarters

Holder Concentration

Across 105 13F filers · Δ vs Mar 31, 2026

Top Holder

23.6%

−5.6 pts

Top 5

62.1%

−0.7 pts

Top 10

79.1%

−1.0 pts

HHI

1,054

Broadly held−175

Quarterly Activity

vs Mar 31, 2026

New

Increased

Decreased

Sold Out

Unchanged

Holder count: +5

Institutional value: -29.3%

Shares held: +11.9%

Top Buyers

vs Mar 31, 2026

Institution	Added	Now Holds
Mega SUSQUEHANNA INTERNATIONAL GROUP, LLP	+1,117,638	1,578,300
Mega TWO SIGMA INVESTMENTS, LP	+849,394	1,239,778
Mega VANGUARD CAPITAL MANAGEMENT LLC	+749,354	3,650,562
Mid ADAR1 Capital Management, LLC	+673,802	2,143,910
Mid Boxer Capital Management, LLC	+474,872	2,844,994

Top Sellers

vs Mar 31, 2026

Institution	Sold	Now Holds
Mega FMR LLC	−884,817	8,454,797
Mega D. E. Shaw & Co., Inc.	−861,163	1,351,656
Mega MILLENNIUM MANAGEMENT LLC	−614,738	796,107
Small DIADEMA PARTNERS LP	−362,665	275,200
Large RENAISSANCE TECHNOLOGIES LLC	−282,438	361,339

Top New Positions

vs Mar 31, 2026

Institution	Acquired	Value (USD)
Mega Jupiter Topco LLC	5,171,053	$5,688,158
Large Caption Management, LLC	878,950	$966,845
Large TUDOR INVESTMENT CORP ET AL	300,000	$330,000
Large Quadrature Capital Ltd	258,928	$284,820
Large NOMURA HOLDINGS INC	150,000	$165,000

Top Sold-Out Positions

vs Mar 31, 2026

Institution	Previously Held	Last Value (USD)
Mega JANUS HENDERSON GROUP PLC	3,308,406	$5,756,626
Large ADAGE CAPITAL PARTNERS GP, L.L.C.	804,958	$1,400,626
Mid 683 Capital Management, LLC	248,790	$432,894
Large CIBC Bancorp USA Inc.	50,000	$87,000
Mid IMC-Chicago, LLC	37,889	$65,926

Filter: New 23 Increased 29 Reduced 16 Sold out 18 Unchanged 37

Showing 100 of 111

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

FMR LLC

$9,300,276

23.60%

8,454,797

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	8,454,780	$9,300,258	100.0%	Defined
STRATEGIC ADVISERS LLC	17	$18	0.0%	Defined

Jupiter Topco LLC

$5,688,158

14.43%

5,171,053

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS US LLC	5,171,053	$5,688,158	100.0%	Defined

VANGUARD CAPITAL MANAGEMENT LLC

$4,015,618

10.19%

3,650,562

Boxer Capital Management, LLC

$3,129,493

7.94%

2,844,994

ADAR1 Capital Management, LLC

$2,358,301

5.98%

2,143,910

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$1,672,440

4.24%

1,520,400

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	1,520,400	$1,672,440	100.0%	Other

D. E. Shaw & Co., Inc.

$1,486,821

3.77%

1,351,656

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	1,351,656	$1,486,821	100.0%	Defined

TWO SIGMA INVESTMENTS, LP

$1,363,755

3.46%

1,239,778

Eversept Partners, LP

$1,138,896

2.89%

1,035,360

GEODE CAPITAL MANAGEMENT, LLC

$883,025

2.24%

802,751

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	156,867	$172,553	19.5%	Defined
Held directly	645,884	$710,472	80.5%	Defined

MILLENNIUM MANAGEMENT LLC

$875,717

2.22%

796,107

BlackRock, Inc.

$856,960

2.17%

779,057

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BlackRock Investment Management, LLC	18,882	$20,770	2.4%	Sole
BlackRock Fund Advisors	280,138	$308,151	36.0%	Sole
BlackRock Institutional Trust Company, N.A.	480,029	$528,031	61.6%	Sole
Aperio Group, LLC	8	$8	0.0%	Sole

VANGUARD FIDUCIARY TRUST CO

$528,845

1.34%

480,769

Caption Management, LLC

$526,845

1.34%

478,950

Caption Management, LLC

$440,000

1.12%

400,000

RENAISSANCE TECHNOLOGIES LLC

$397,472

1.01%

361,339

STATE STREET CORP

$370,271

0.94%

336,610

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS TRUST COMPANY	336,610	$370,271	100.0%	Defined

CITADEL ADVISORS LLC

$351,450

0.89%

319,500

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	319,500	$351,450	100.0%	Defined

TUDOR INVESTMENT CORP ET AL

$330,000

0.84%

300,000

DIADEMA PARTNERS LP

$302,720

0.77%

275,200

GSA CAPITAL PARTNERS LLP

$291,512

0.74%

265,011

UBS Group AG

$288,550

0.73%

262,320

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	128	$140	0.0%	Defined
UBS AG	141,014	$155,115	53.8%	Defined
UBS Switzerland AG	121,178	$133,295	46.2%	Defined

Quadrature Capital Ltd

$284,820

0.72%

258,928

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Quadrature US Inc.	258,928	$284,820	100.0%	Other

CITADEL ADVISORS LLC

$265,952

0.67%

241,775

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	212,653	$233,918	88.0%	Defined
Held directly	29,122	$32,034	12.0%	Defined

NORTHERN TRUST CORP

$191,833

0.49%

174,394

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	174,394	$191,833	100.0%	Defined

BRIDGEWAY CAPITAL MANAGEMENT, LLC

$190,300

0.48%

173,000

NOMURA HOLDINGS INC

$165,000

0.42%

150,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NOMURA SECURITIES INTERNATIONAL INC	150,000	$165,000	100.0%	Defined

HRT FINANCIAL LP

$154,475

0.39%

140,432

DZ BANK AG Deutsche Zentral Genossenschafts Bank, Frankfurt am Main

$147,012

0.37%

133,648

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UNION ASSET MANAGEMENT HOLDING AG joint	133,648	$147,012	100.0%	Defined

GMT Capital Corp

$138,730

0.35%

126,119

JANE STREET GROUP, LLC

$136,977

0.35%

124,525

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	123,594	$135,953	99.3%	Defined
Jane Street Global Trading, LLC	931	$1,024	0.7%	Defined

XTX Topco Ltd

$116,416

0.30%

105,833

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	105,833	$116,416	100.0%	Defined

GOLDMAN SACHS GROUP INC

$103,092

0.26%

93,720

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	93,720	$103,092	100.0%	Defined

CITADEL ADVISORS LLC

$92,620

0.23%

84,200

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	84,200	$92,620	100.0%	Defined

Invesco Ltd.

$88,371

0.22%

80,338

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Capital Management LLC	80,338	$88,371	100.0%	Defined

Green Alpha Advisors, LLC

$74,860

0.19%

68,055

JANE STREET GROUP, LLC

$71,940

0.18%

65,400

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	65,400	$71,940	100.0%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$63,690

0.16%

57,900

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	57,900	$63,690	100.0%	Other

MORGAN STANLEY

$55,279

0.14%

50,256

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	19	$20	0.0%	Defined
MORGAN STANLEY & CO. LLC	3,705	$4,075	7.4%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	2,000	$2,200	4.0%	Defined
MORGAN STANLEY SMITH BARNEY LLC	44,532	$48,984	88.6%	Defined

RAYMOND JAMES FINANCIAL INC

$46,513

0.12%

42,286

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	40,278	$44,305	95.3%	Defined
Raymond James Financial Services Advisors, Inc.	2,008	$2,208	4.7%	Defined

Squarepoint Ops LLC

$40,030

0.10%

36,391

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Squarepoint Capital LLP	36,391	$40,030	100.0%	Defined

Engineers Gate Manager LP

$37,060

0.09%

33,691

Greenwich Wealth Management LLC

$35,198

0.09%

31,999

Vanguard Global Advisers, LLC

$33,258

0.08%

30,235

Headlands Technologies LLC

$32,877

0.08%

29,889

ZACKS INVESTMENT MANAGEMENT

$29,078

0.07%

26,435

Prime Capital Investment Advisors, LLC

$18,838

0.05%

17,126

LPL Financial LLC

$18,825

0.05%

17,114

TWO SIGMA SECURITIES, LLC

$17,553

0.04%

15,958

STEPHENS INC /AR/

$17,050

0.04%

15,500

StoneX Group Inc.

$15,677

0.04%

14,252

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
StoneX Advisors Inc. joint	14,252	$15,677	100.0%	Defined

AXQ CAPITAL, LP

$15,249

0.04%

13,863

Tower Research Capital LLC (TRC)

$14,631

0.04%

13,301

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Latour Trading LLC (Latour)	13,301	$14,631	100.0%	Defined

HighTower Advisors, LLC

$14,003

0.04%

12,730

Mayflower Financial Advisors, LLC

$13,814

0.04%

12,559

MCF Advisors LLC

$12,401

0.03%

11,274

SIGNATURE ESTATE & INVESTMENT ADVISORS LLC

$12,130

0.03%

11,028

Caitong International Asset Management Co., Ltd

$11,024

0.03%

10,022

SBI Securities Co., Ltd.

$7,495

0.02%

6,814

Covestor Ltd

$5,632

0.01%

5,120

ROYAL BANK OF CANADA

$4,677

0.01%

4,252

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC Capital Markets, LLC	3,011	$3,312	70.8%	Defined
RBC Dominion Securities Inc.	1,121	$1,233	26.4%	Defined
Held directly	120	$132	2.8%	Sole

BARCLAYS PLC

$3,835

0.01%

3,487

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS CAPITAL INC.	3,487	$3,835	100.0%	Defined

WELLS FARGO & COMPANY/MN

$3,585

0.01%

3,260

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	277	$304	8.5%	Defined
WELLS FARGO CLEARING SERVICES, LLC	2,983	$3,281	91.5%	Defined

TD Waterhouse Canada Inc.

$2,621

0.01%

2,383

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
TORONTO DOMINION BANK	2,383	$2,621	100.0%	Defined

Rockefeller Capital Management L.P.

$2,313

0.01%

2,103

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ROCKEFELLER FINANCIAL LLC	2,103	$2,313	100.0%	Defined

Activest Wealth Management

$1,650

0.00%

1,500

Steward Partners Investment Advisory, LLC

$1,519

0.00%

1,381

Optiver Holding B.V.

$1,457

0.00%

1,325

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Optiver VOF	1,325	$1,457	100.0%	Defined

OSAIC HOLDINGS, INC.

$1,025

0.00%

933

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
OSAIC WEALTH, INC.	529	$581	56.7%	Defined
Osaic Advisory Services, LLC	404	$444	43.3%	Defined

IFG Advisory, LLC

$916

0.00%

833

Persistent Asset Partners Ltd

$771

0.00%

701

Federation des caisses Desjardins du Quebec

$705

0.00%

641

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Desjardins Securities inc.	141	$155	22.0%	Defined
Held directly	500	$550	78.0%	Defined

NFSG Corp

$394

0.00%

359

BNP PARIBAS ARBITRAGE, SA

$387

0.00%

352

MJT & Associates Financial Advisory Group, Inc.

$357

0.00%

325

PNC FINANCIAL SERVICES GROUP, INC.

$330

0.00%

300

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PNC Wealth Management LLC	300	$330	100.0%	Defined

Pioneer Family Office, LLC

$320

0.00%

291

VSM Wealth Advisory, LLC

$275

0.00%

250

JONES FINANCIAL COMPANIES LLLP

$264

0.00%

240

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Edward D. Jones & Co., L.P.	240	$264	100.0%	Sole

HARBOUR INVESTMENTS, INC.

$260

0.00%

237

Aberdeen Wealth Management LLC

$219

0.00%

200

Decker Retirement Planning Inc.

$184

0.00%

168

Farther Finance Advisors, LLC

$122

0.00%

111

Arax Advisory Partners

$113

0.00%

103

Smartleaf Asset Management LLC

$91

0.00%

Blume Capital Management, Inc.

$91

0.00%

PITTENGER & ANDERSON INC

$84

0.00%

Ameritas Investment Partners, Inc.

$70

0.00%

MCDONALD PARTNERS LLC

$63

0.00%

NATIONAL BANK OF CANADA /FI/

$63

0.00%

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
National Bank Financial Inc.	58	$63	100.0%	Defined

Allworth Financial LP

$51

0.00%

JPMORGAN CHASE & CO

$50

0.00%

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
J.P. Morgan Securities LLC	46	$50	100.0%	Defined

1 NORTH WEALTH SERVICES LLC

$45

0.00%

Pinnacle Wealth Planning Services, Inc.

$45

0.00%

INTERNATIONAL ASSETS INVESTMENT MANAGEMENT, LLC

$39

0.00%

Allied Private Wealth LLC

$36

0.00%

VALLEY NATIONAL ADVISERS INC

$26

0.00%

ATWOOD & PALMER INC

$19

0.00%

CoreCap Advisors, LLC

$17

0.00%

Key Financial Inc

$15

0.00%

Key facts CIK 1055726 CUSIP 45773H409 13F (30d) 61 filings 61 filers Visit website Investor relations