ABUS · Arbutus Biopharma Corp

$4.77 -0.21 (-4.22%) At close · Aug 20

Market Cap

$943.73M

197.85M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Quarter:

Holders: 176

Total Value: $675,545,933

Total Shares: 140,749,039

Institutional Ownership Trend

26 quarters

Holder Concentration

Across 176 13F filers · Δ vs Mar 31, 2026

Top Holder

13.6%

−4.9 pts

Top 5

52.7%

−3.7 pts

Top 10

71.5%

−2.3 pts

HHI

697

Broadly held−132

Quarterly Activity

vs Mar 31, 2026

New

Increased

Decreased

Sold Out

Unchanged

Holder count: +6

Institutional value: +7.3%

Shares held: +0.6%

Top Buyers

vs Mar 31, 2026

Institution	Added	Now Holds
Mega GOLDMAN SACHS GROUP INC	+2,406,368	3,506,802
Mega STATE STREET CORP	+1,526,669	7,869,179
Mega BANK OF AMERICA CORP /DE/	+962,569	2,646,197
Large Rubric Capital Management LP	+906,548	7,532,937
Small Whitefort Capital Management, LP	+863,376	15,794,261

Top Sellers

vs Mar 31, 2026

Institution	Sold	Now Holds
Mega MORGAN STANLEY	−8,193,175	17,698,305
Mid ADAR1 Capital Management, LLC	−1,865,028	1,268,147
Mega UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC	−577,158	261,052
Mega CITIGROUP INC	−449,184	879,237
Mega CITADEL ADVISORS LLC	−380,542	404,696

Top New Positions

vs Mar 31, 2026

Institution	Acquired	Value (USD)
Large NOMURA HOLDINGS INC	1,637,081	$7,857,988
Mega ARROWSTREET CAPITAL, LIMITED PARTNERSHIP	650,700	$3,123,360
Large MACKENZIE FINANCIAL CORP	613,806	$2,946,268
Large LOS ANGELES CAPITAL MANAGEMENT LLC	602,008	$2,889,638
Mega Jupiter Topco LLC	597,423	$2,867,630

Top Sold-Out Positions

vs Mar 31, 2026

Institution	Previously Held	Last Value (USD)
Large ADAGE CAPITAL PARTNERS GP, L.L.C.	2,664,941	$11,992,234
Mega MILLENNIUM MANAGEMENT LLC	642,448	$2,891,016
Mega RAYMOND JAMES FINANCIAL INC	191,355	$861,097
Large CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM	142,985	$643,432
Large Balyasny Asset Management L.P.	125,521	$564,844

Filter: New 34 Increased 62 Reduced 41 Sold out 28 Unchanged 39

Showing 100 of 179

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

Two Seas Capital LP

$91,658,798

13.57%

19,095,583

MORGAN STANLEY

$84,951,860

12.58%

17,698,305

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	154	$739	0.0%	Defined
MORGAN STANLEY & CO. LLC	43,676	$209,644	0.2%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	277,247	$1,330,785	1.6%	Defined
MORGAN STANLEY INVESTMENT MANAGEMENT INC	17,122,142	$82,186,281	96.7%	Defined
MORGAN STANLEY SMITH BARNEY LLC	5,041	$24,196	0.0%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	250,045	$1,200,215	1.4%	Defined

Whitefort Capital Management, LP

$75,812,452

11.22%

15,794,261

BlackRock, Inc.

$66,077,138

9.78%

13,766,071

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	118,208	$567,398	0.9%	Sole
BlackRock Asset Management Canada Ltd	628	$3,014	0.0%	Sole
BlackRock Investment Management, LLC	470,053	$2,256,254	3.4%	Sole
BLACKROCK ADVISORS LLC	360,831	$1,731,988	2.6%	Sole
BlackRock Fund Advisors	7,767,429	$37,283,659	56.4%	Sole
BlackRock Institutional Trust Company, N.A.	4,448,055	$21,350,664	32.3%	Sole
BlackRock Fund Managers Ltd.	26,652	$127,929	0.2%	Sole
BlackRock Asset Management Ireland Ltd	408,385	$1,960,248	3.0%	Sole
BlackRock Asset Management Schweiz AG	22,815	$109,512	0.2%	Sole
Aperio Group, LLC	143,015	$686,472	1.0%	Sole

STATE STREET CORP

$37,772,059

5.59%

7,869,179

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	7,869,179	$37,772,059	100.0%	Defined

Rubric Capital Management LP

$36,158,097

5.35%

7,532,937

VANGUARD CAPITAL MANAGEMENT LLC

$33,651,259

4.98%

7,010,679

BARCLAYS PLC

$20,197,987

2.99%

4,207,914

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	104,025	$499,320	2.5%	Defined
BARCLAYS CAPITAL INC.	4,103,889	$19,698,667	97.5%	Defined

GEODE CAPITAL MANAGEMENT, LLC

$19,764,916

2.93%

4,117,691

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	580,220	$2,785,056	14.1%	Defined
Held directly	3,537,471	$16,979,860	85.9%	Defined

GOLDMAN SACHS GROUP INC

$16,832,648

2.49%

3,506,802

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	1,780,381	$8,545,828	50.8%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	1,693,019	$8,126,491	48.3%	Defined
Goldman Sachs Trust Company, N.A. joint	33,402	$160,329	1.0%	Other

BANK OF AMERICA CORP /DE/

$12,701,745

1.88%

2,646,197

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	1,493,719	$7,169,851	56.4%	Defined
BOFA SECURITIES, INC.	1,152,478	$5,531,894	43.6%	Defined

BlackBarn Capital Partners LP

$12,000,000

1.78%

2,500,000

Legal & General Group Plc

$11,837,408

1.75%

2,466,127

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	15,631	$75,028	0.6%	Defined
Legal & General Investment Management Ltd joint	2,450,496	$11,762,380	99.4%	Defined

Woodline Partners LP

$10,952,908

1.62%

2,281,856

Jefferies Financial Group Inc.

$8,216,025

1.22%

1,711,672

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jefferies Financial Services, Inc.	1,711,672	$8,216,025	100.0%	Defined

NOMURA HOLDINGS INC

$7,857,988

1.16%

1,637,081

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NOMURA GLOBAL FINANCIAL PRODUCTS INC	1,637,081	$7,857,988	100.0%	Defined

VANGUARD PORTFOLIO MANAGEMENT LLC

$7,398,998

1.10%

1,541,458

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$6,869,846

1.02%

1,431,218

NORTHERN TRUST CORP

$6,811,314

1.01%

1,419,024

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	1,328,149	$6,375,115	93.6%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	65,596	$314,860	4.6%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	25,279	$121,339	1.8%	Other

OSAIC HOLDINGS, INC.

$6,469,756

0.96%

1,347,866

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
OSAIC WEALTH, INC.	1,347,866	$6,469,756	100.0%	Defined

ADAR1 Capital Management, LLC

$6,087,105

0.90%

1,268,147

Deltroit Asset Management (UK) LLP

$5,760,000

0.85%

1,200,000

VANGUARD FIDUCIARY TRUST CO

$5,449,737

0.81%

1,135,362

CITIGROUP INC

$4,220,337

0.62%

879,237

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	879,237	$4,220,337	100.0%	Defined

BANK OF MONTREAL /CAN/

$3,734,102

0.55%

777,938

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BMO NESBITT BURNS INC.	777,938	$3,734,102	100.0%	Defined

ARROWSTREET CAPITAL, LIMITED PARTNERSHIP

$3,123,360

0.46%

650,700

MACKENZIE FINANCIAL CORP

$2,946,268

0.44%

613,806

LOS ANGELES CAPITAL MANAGEMENT LLC

$2,889,638

0.43%

602,008

Jupiter Topco LLC

$2,867,630

0.42%

597,423

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	60,900	$292,320	10.2%	Defined
JANUS HENDERSON INVESTORS US LLC	536,523	$2,575,310	89.8%	Defined

Western Standard LLC

$2,605,377

0.39%

542,787

Rangeley Capital, LLC

$2,400,000

0.36%

500,000

Nuveen, LLC

$2,383,723

0.35%

496,609

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	2,134	$10,243	0.4%	Defined
TIAA CREF INVESTMENT MANAGEMENT LLC joint	211,720	$1,016,256	42.6%	Defined
TEACHERS ADVISORS, LLC joint	282,755	$1,357,224	56.9%	Defined

Man Group plc

$2,272,641

0.34%

473,467

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Numeric Investors LLC	473,467	$2,272,641	100.0%	Defined

WELLINGTON MANAGEMENT GROUP LLP

$2,215,900

0.33%

461,646

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wellington Management Company LLP	146,139	$701,467	31.7%	Defined
Wellington Trust Company, NA joint	294,889	$1,415,467	63.9%	Defined
Wellington Management Australia Pty Ltd	20,618	$98,966	4.5%	Defined

Bank of New York Mellon Corp

$2,179,751

0.32%

454,115

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Investments Corporation	166,889	$801,067	36.8%	Defined
The Bank of New York Mellon	287,226	$1,378,684	63.2%	Defined

CenterBook Partners LP

$2,136,724

0.32%

445,151

Freestone Capital Holdings, LLC

$2,124,000

0.31%

442,500

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FREESTONE CAPITAL MANAGEMENT LLC	442,500	$2,124,000	100.0%	Defined

CITADEL ADVISORS LLC

$1,942,540

0.29%

404,696

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	404,696	$1,942,540	100.0%	Defined

KENNEDY CAPITAL MANAGEMENT LLC

$1,849,430

0.27%

385,298

D. E. Shaw & Co., Inc.

$1,832,942

0.27%

381,863

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	252,003	$1,209,614	66.0%	Defined
D. E. Shaw Investment Management, L.L.C. joint	129,860	$623,328	34.0%	Defined

Swiss National Bank

$1,375,200

0.20%

286,500

Ledgewood Wealth Advisors, LLC

$1,336,492

0.20%

278,436

FMR LLC

$1,282,502

0.19%

267,188

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	266,090	$1,277,232	99.6%	Defined
STRATEGIC ADVISERS LLC	1,058	$5,078	0.4%	Defined
Fidelity Diversifying Solutions LLC	40	$192	0.0%	Defined

Susquehanna Portfolio Strategies, LLC

$1,265,688

0.19%

263,685

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$1,253,048

0.19%

261,052

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	1,354	$6,499	0.5%	Defined
UBS Asset Management Switzerland AG	227,322	$1,091,145	87.1%	Sole
Held directly	32,376	$155,404	12.4%	Defined

Carrera Capital Advisors

$1,234,401

0.18%

257,167

Quinn Opportunity Partners LLC

$1,225,027

0.18%

255,214

Invesco Ltd.

$1,204,352

0.18%

250,907

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	31,226	$149,884	12.4%	Defined
Invesco Trust Co joint	63,670	$305,616	25.4%	Defined
Invesco Capital Management LLC	156,011	$748,852	62.2%	Defined

UBS Group AG

$1,196,198

0.18%

249,208

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	22,040	$105,792	8.8%	Defined
UBS AG	188,668	$905,606	75.7%	Defined
UBS Switzerland AG	38,500	$184,800	15.4%	Defined

STATE OF WISCONSIN INVESTMENT BOARD

$1,161,681

0.17%

242,017

Campbell & CO Investment Adviser LLC

$1,106,563

0.16%

230,534

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$1,080,475

0.16%

225,099

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	225,099	$1,080,475	100.0%	Other

SG Americas Securities, LLC

$1,079,491

0.16%

224,894

Squarepoint Ops LLC

$999,096

0.15%

208,145

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Squarepoint Capital LLP	208,145	$999,096	100.0%	Defined

MARTINGALE ASSET MANAGEMENT L P

$926,880

0.14%

193,100

RHUMBLINE ADVISERS

$909,230

0.13%

189,423

DEUTSCHE BANK AG\

$818,179

0.12%

170,454

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DWS INVESTMENT MANAGEMENT AMERICAS, INC	3,639	$17,467	2.1%	Defined
DWS Investments UK Ltd	166,815	$800,712	97.9%	Defined

STRS OHIO

$810,720

0.12%

168,900

Hudson Bay Capital Management LP

$720,000

0.11%

150,000

Caption Management, LLC

$720,000

0.11%

150,000

Caption Management, LLC

$720,000

0.11%

150,000

Wealth Management Associates, Inc.

$710,376

0.11%

147,995

WELLS FARGO & COMPANY/MN

$710,370

0.11%

147,994

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	146,722	$704,265	99.1%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	72	$345	0.0%	Defined
WELLS FARGO CLEARING SERVICES, LLC	1,200	$5,760	0.8%	Defined

ALLIANCEBERNSTEIN L.P.

$673,152

0.10%

140,240

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	140,240	$673,152	100.0%	Defined

PRICE T ROWE ASSOCIATES INC /MD/

$642,374

0.10%

133,828

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	133,828	$642,374	100.0%	Defined

Connor, Clark & Lunn Investment Management Ltd.

$624,109

0.09%

130,023

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 1	129,101	$619,684	99.3%	Sole
Held directly	922	$4,425	0.7%	Sole

Front Row Advisors LLC

$530,400

0.08%

110,500

Rafferty Asset Management, LLC

$490,416

0.07%

102,170

JPMORGAN CHASE & CO

$471,474

0.07%

98,224

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	66,680	$320,064	67.9%	Defined
J.P. Morgan Investment Management Inc.	12,931	$62,068	13.2%	Defined
J.P. Morgan Securities plc	18,613	$89,342	18.9%	Defined

MetLife Investment Management, LLC

$452,884

0.07%

94,351

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
METLIFE INC	94,351	$452,884	100.0%	Defined

VANGUARD ASSET MANAGEMENT, Ltd

$429,993

0.06%

89,582

MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.

$412,507

0.06%

85,939

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Global X Management CO LLC joint	74,070	$355,536	86.2%	Defined
Global X Management (AUS) Ltd joint	11,869	$56,971	13.8%	Defined

TWO SIGMA INVESTMENTS, LP

$407,932

0.06%

84,986

JBF Capital, Inc.

$385,348

0.06%

80,281

Integrated Quantitative Investments LLC

$383,697

0.06%

79,937

Virtus Investment Advisers, LLC

$368,673

0.05%

76,807

XTX Topco Ltd

$365,808

0.05%

76,210

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	76,210	$365,808	100.0%	Defined

AMERICAN CENTURY COMPANIES INC

$317,832

0.05%

66,215

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AMERICAN CENTURY INVESTMENT MANAGEMENT INC	66,215	$317,832	100.0%	Defined

Rockefeller Capital Management L.P.

$302,400

0.04%

63,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ROCKEFELLER FINANCIAL LLC	63,000	$302,400	100.0%	Defined

Creative Planning

$298,132

0.04%

62,111

Russell Investments Group, Ltd.

$283,972

0.04%

59,161

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC	59,161	$283,972	100.0%	Defined

Quantinno Capital Management LP

$273,412

0.04%

56,961

Zurcher Kantonalbank (Zurich Cantonalbank)

$265,118

0.04%

55,233

Vanguard Global Advisers, LLC

$264,729

0.04%

55,152

Mitsubishi UFJ Asset Management Co., Ltd.

$255,936

0.04%

53,320

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MITSUBISHI UFJ FINANCIAL GROUP INC	53,320	$255,936	100.0%	Defined

Jump Financial, LLC

$255,177

0.04%

53,162

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Leap GP LLC joint	53,162	$255,177	100.0%	Defined

PROFUND ADVISORS LLC

$254,481

0.04%

53,017

Police & Firemen's Retirement System of New Jersey

$248,203

0.04%

51,709

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Capital	51,709	$248,203	100.0%	Defined

Exome Asset Management LLC

$240,796

0.04%

50,166

Magnetar Financial LLC

$225,921

0.03%

47,067

State of New Jersey Common Pension Fund D

$221,601

0.03%

46,167

MANUFACTURERS LIFE INSURANCE COMPANY, THE

$212,664

0.03%

44,305

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MANULIFE FINANCIAL CORP joint	44,305	$212,664	100.0%	Defined

ALLSTATE CORP

$212,284

0.03%

44,226

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ALLSTATE INVESTMENTS LLC	44,226	$212,284	100.0%	Sole

GSA CAPITAL PARTNERS LLP

$211,593

0.03%

44,082

Quadrature Capital Ltd

$201,619

0.03%

42,004

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Quadrature US Inc.	42,004	$201,619	100.0%	Other

Universal- Beteiligungs- und Servicegesellschaft mbH

$189,475

0.03%

39,474

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Universal-Investment-Gesellschaft mbH	39,474	$189,475	100.0%	Defined

ARIZONA STATE RETIREMENT SYSTEM

$189,110

0.03%

39,398

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$175,680

0.03%

36,600

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	36,600	$175,680	100.0%	Other

STATE BOARD OF ADMINISTRATION OF FLORIDA RETIREMENT SYSTEM

$171,936

0.03%

35,820

NEW YORK STATE COMMON RETIREMENT FUND

$169,286

0.03%

35,268

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RF&L WEALTH MANAGEMENT, LLC	90	$432	0.3%	Other
Arax Advisory Partners	112	$537	0.4%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	76	$364	0.2%	Other
Stokes Family Office, LLC	22	$105	0.1%	Other
Held directly	31,441	$150,916	99.1%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
First Sentier Investors RQI Pty Ltd	31,688	$152,102	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	31,473	$151,070	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	27,300	$131,040	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PGIM Quantitative Solutions LLC	26,380	$126,624	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF NOVA SCOTIA	14,740	$70,752	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DNB FUND	14,643	$70,286	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
EATON VANCE MANAGEMENT	14,597	$70,065	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Amundi Asset Management	11,878	$57,014	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	9,467	$45,441	90.4%	Defined
Ameriprise Financial Services, LLC	1,000	$4,800	9.6%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Latour Trading LLC (Latour)	7,719	$37,051	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC CMA LTD.	4,890	$23,472	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FEDERATED MDTA LLC	4,041	$19,396	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNP PARIBAS ASSET MANAGEMENT Europe	2,293	$11,006	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PNC Bank, National Association	1,690	$8,112	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
US BANK NATIONAL ASSOCIATION	1,494	$7,171	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Edward D. Jones & Co., L.P.	830	$3,984	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
National Bank Financial Inc.	700	$3,360	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	2,754	$12,393	1.4%	Defined
Raymond James Financial Services Advisors, Inc.	188,601	$848,704	98.6%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Schroder Investment Management Limited	121,313	$545,908	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DRW Investments, LLC joint	69,089	$310,900	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	60,900	$274,050	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	33,000	$148,500	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	25,500	$114,750	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Optiver VOF	669	$3,010	100.0%	Defined

Key facts CIK 1447028 CUSIP 03879J100 13F (30d) 174 filings 170 filers Visit website