ALNY · Alnylam Pharmaceuticals, Inc.

$233.48 -3.08 (-1.30%)

Nasdaq-100 member Russell 1000 member Rebalance outlook locked

Market Cap

$31.98B

133.81M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Quarter:

Holders: 786

Total Value: $40,708,325,511

Total Shares: 135,230,139

Showing 1–100 of 817 holdings by value · page 1 of 9

Institutional Ownership Trend

26 quarters

Superinvestors Holding ALNY

All superinvestors →

Ray Dalio $29,977,771 Joel Greenblatt $12,658,612

Holder Concentration

Across 786 13F filers · Δ vs Mar 31, 2026

Top Holder

11.1%

−0.1 pts

Top 5

39.5%

−0.2 pts

Top 10

55.5%

−2.2 pts

HHI

421

Broadly held−13

Quarterly Activity

vs Mar 31, 2026

New

Increased

321

Decreased

281

Sold Out

Unchanged

Holder count: -11

Institutional value: -5.7%

Shares held: +3.6%

Top Buyers

vs Mar 31, 2026

Institution	Added	Now Holds
Mega Invesco Ltd.	+2,462,592	3,754,318
Large Orbis Allan Gray Ltd	+652,582	1,987,407
Large LOS ANGELES CAPITAL MANAGEMENT LLC	+641,357	729,635
Mega BlackRock, Inc.	+462,767	10,907,675
Mega T. Rowe Price Investment Management, Inc.	+448,849	5,642,173

Top Sellers

vs Mar 31, 2026

Institution	Sold	Now Holds
Mega JPMORGAN CHASE & CO	−3,648,235	2,675,517
Large MARSHALL WACE, LLP	−839,519	259,799
Mega PRICE T ROWE ASSOCIATES INC /MD/	−283,547	1,149,201
Large Allspring Global Investments Holdings, LLC	−279,247	2,546
Large Squarepoint Ops LLC	−250,324	257,899

Top New Positions

vs Mar 31, 2026

Institution	Acquired	Value (USD)
Mega NORGES BANK	2,271,813	$683,883,867
Mid Deep Track Capital, LP	825,000	$248,349,750
Mid Paradigm Biocapital Advisors LP	480,000	$144,494,400
Mega Jupiter Topco LLC	135,545	$40,803,110
Small Davern Capital Partners, LP	113,085	$34,041,977

Top Sold-Out Positions

vs Mar 31, 2026

Institution	Previously Held	Last Value (USD)
Large CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM	196,863	$65,136,060
Mega JANUS HENDERSON GROUP PLC	147,089	$48,667,337
Large HRT FINANCIAL LP	128,633	$42,560,800
Mid Andra AP-fonden	86,697	$28,685,436
Large PARNASSUS INVESTMENTS, LLC	48,991	$16,209,652

Filter: New 88 Increased 321 Reduced 281 Sold out 99 Unchanged 96

Showing 100 of 817

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

FMR LLC

$4,522,663,276

11.11%

15,023,962

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC joint	10,325,698	$3,108,344,868	68.7%	Defined
FIDELITY MANAGEMENT TRUST CO	2,380,552	$716,617,568	15.8%	Defined
STRATEGIC ADVISERS LLC	756,165	$227,628,349	5.0%	Defined
Fidelity Institutional Asset Management Trust Co	102,664	$30,904,943	0.7%	Defined
FIAM LLC	1,457,197	$438,660,012	9.7%	Defined
Fidelity Diversifying Solutions LLC	1,686	$507,536	0.0%	Defined

Capital World Investors

$3,354,932,991

8.24%

11,144,846

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CAPITAL GROUP COMPANIES INC joint	11,144,846	$3,354,932,991	100.0%	Defined

BlackRock, Inc.

$3,283,537,392

8.07%

10,907,675

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	341,519	$102,807,464	3.1%	Sole
BlackRock Investment Management (Australia) Ltd	91,954	$27,680,912	0.8%	Sole
BlackRock Asset Management Canada Ltd	117,758	$35,448,690	1.1%	Sole
BLACKROCK (SINGAPORE) LTD.	9,715	$2,924,506	0.1%	Sole
BlackRock Asset Management North Asia Ltd	17,497	$5,267,121	0.2%	Sole
BlackRock Investment Management, LLC	167,576	$50,445,403	1.5%	Sole
BLACKROCK ADVISORS LLC	303,580	$91,386,687	2.8%	Sole
BlackRock Fund Advisors	4,171,597	$1,255,775,844	38.2%	Sole
BlackRock Institutional Trust Company, N.A.	3,428,111	$1,031,964,254	31.4%	Sole
BlackRock Japan Co. Ltd	138,832	$41,792,596	1.3%	Sole
BlackRock Fund Managers Ltd.	149,228	$44,922,104	1.4%	Sole
BlackRock Investment Management (UK) Ltd.	426,523	$128,396,218	3.9%	Sole
BlackRock (Netherlands) B.V.	230,932	$69,517,459	2.1%	Sole
BlackRock International, Ltd.	146	$43,950	0.0%	Sole
BlackRock Asset Management Ireland Ltd	999,406	$300,851,188	9.2%	Sole
BlackRock Advisors (UK) Ltd	6,941	$2,089,449	0.1%	Sole
BlackRock Asset Management Deutschland AG	36,024	$10,844,304	0.3%	Sole
BlackRock (Luxembourg) S.A.	108,675	$32,714,435	1.0%	Sole
BlackRock Life Ltd	34,397	$10,354,528	0.3%	Sole
BlackRock Asset Management Schweiz AG	5,011	$1,508,461	0.0%	Sole
Aperio Group, LLC	116,696	$35,128,996	1.1%	Sole
SpiderRock Advisors, LLC	5,557	$1,672,823	0.1%	Sole

Capital Research Global Investors

$2,935,252,919

7.21%

9,750,699

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CAPITAL GROUP COMPANIES INC joint	9,750,699	$2,935,252,919	100.0%	Defined

VANGUARD PORTFOLIO MANAGEMENT LLC

$1,968,251,541

4.84%

6,538,390

VANGUARD CAPITAL MANAGEMENT LLC

$1,815,053,762

4.46%

6,029,478

T. Rowe Price Investment Management, Inc.

$1,698,463,338

4.17%

5,642,173

Invesco Ltd.

$1,130,162,341

2.78%

3,754,318

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	44,188	$13,301,911	1.2%	Defined
Invesco Trust Co joint	54,395	$16,374,526	1.4%	Defined
Invesco Management S.A.	4,244	$1,277,571	0.1%	Defined
Invesco Taiwan Ltd	220	$66,226	0.0%	Defined
Invesco Asset Management (Japan) Limited	2,988	$899,477	0.1%	Defined
Invesco Investment Advisers LLC	865	$260,390	0.0%	Defined
Invesco Capital Management LLC	3,647,418	$1,097,982,240	97.2%	Defined

STATE STREET CORP

$1,070,369,059

2.63%

3,555,689

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS (JAPAN) CO., LTD. joint	3,555,689	$1,070,369,059	100.0%	Defined

JPMORGAN CHASE & CO

$805,410,876

1.98%

2,675,517

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
J.P. Morgan Trust Co of Delaware	199	$59,904	0.0%	Other
JPMorgan Chase Bank, N.A.	99,676	$30,005,466	3.7%	Defined
J.P. Morgan Investment Management Inc.	2,237,814	$673,649,148	83.6%	Other
J.P. Morgan Securities LLC	74,040	$22,288,260	2.8%	Defined
JPMorgan Asset Management (UK) Ltd	152,972	$46,049,161	5.7%	Defined
JPMorgan Asset Management (Canada) Inc.	913	$274,840	0.0%	Defined
JPMorgan Asset Management (Asia Pacific) Ltd	3,785	$1,139,398	0.1%	Defined
JPMorgan Asset Management (Japan) Ltd	33,023	$9,940,913	1.2%	Defined
JPMorgan Asset Management (Taiwan) Ltd	3,500	$1,053,605	0.1%	Defined
55I, LLC	59	$17,760	0.0%	Other
J.P. Morgan SE	9,516	$2,864,601	0.4%	Defined
J.P. MORGAN MANSART MANAGEMENT Ltd	60,020	$18,067,820	2.2%	Defined

GEODE CAPITAL MANAGEMENT, LLC

$793,466,914

1.95%

2,635,840

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	347,115	$104,492,028	13.2%	Defined
Held directly	2,288,725	$688,974,886	86.8%	Defined

BAILLIE GIFFORD & CO

$734,479,783

1.80%

2,439,889

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BAILLIE GIFFORD & CO LTD	550,940	$165,849,468	22.6%	Defined
BAILLIE GIFFORD OVERSEAS LTD	852,262	$256,556,429	34.9%	Defined
Baillie Gifford Investment Management (Europe) Ltd	247,199	$74,414,314	10.1%	Defined
Held directly	789,488	$237,659,572	32.4%	Sole

NORGES BANK

$683,883,867

1.68%

2,271,813

WELLINGTON MANAGEMENT GROUP LLP

$635,295,816

1.56%

2,110,407

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wellington Management Company LLP	2,042,733	$614,923,914	96.8%	Defined
Wellington Managment Canada LLC	1,344	$404,584	0.1%	Defined
Wellington Management Japan Pte Ltd	719	$216,440	0.0%	Defined
Wellington Management Singapore Pte. Ltd.	14,339	$4,316,469	0.7%	Defined
Wellington Management International Ltd	19,225	$5,787,301	0.9%	Defined
Wellington Trust Company, NA joint	26,634	$8,017,633	1.3%	Defined
Wellington Management Australia Pty Ltd	5,413	$1,629,475	0.3%	Defined

Orbis Allan Gray Ltd

$598,269,128

1.47%

1,987,407

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ORBIS INVESTMENT MANAGEMENT LTD	1,984,586	$597,419,923	99.9%	Defined
Held directly	2,821	$849,205	0.1%	Defined

Dodge & Cox

$588,631,954

1.45%

1,955,393

TWO SIGMA INVESTMENTS, LP

$564,713,315

1.39%

1,875,937

AMERICAN CENTURY COMPANIES INC

$558,096,374

1.37%

1,853,956

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AMERICAN CENTURY INVESTMENT MANAGEMENT INC	1,853,956	$558,096,374	100.0%	Defined

Clearbridge Investments, LLC

$379,142,167

0.93%

1,259,483

PRICE T ROWE ASSOCIATES INC /MD/

$345,943,976

0.85%

1,149,201

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	139,541	$42,006,027	12.1%	Defined
Held directly	1,009,660	$303,937,949	87.9%	Sole

D. E. Shaw & Co., Inc.

$332,369,932

0.82%

1,104,109

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	794,407	$239,140,339	72.0%	Defined
D. E. Shaw Investment Management, L.L.C. joint	309,702	$93,229,593	28.0%	Defined

MORGAN STANLEY

$310,708,106

0.76%

1,032,150

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	994	$299,223	0.1%	Defined
MORGAN STANLEY & CO. LLC	137,251	$41,316,668	13.3%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	308,083	$92,742,225	29.8%	Defined
MORGAN STANLEY AIP GP LP	138	$41,541	0.0%	Defined
MORGAN STANLEY INVESTMENT MANAGEMENT INC	8,412	$2,532,263	0.8%	Defined
MORGAN STANLEY SMITH BARNEY LLC	182,492	$54,935,566	17.7%	Defined
Morgan Stanley Strategic Investments, Inc	1	$301	0.0%	Defined
Parametric Sas	110	$33,113	0.0%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	340,402	$102,471,213	33.0%	Defined
EATON VANCE MANAGEMENT	11,509	$3,464,554	1.1%	Defined
Calvert Research & Management	26,003	$7,827,682	2.5%	Defined
BOSTON MANAGEMENT & RESEARCH	16,755	$5,043,757	1.6%	Defined

BANK OF AMERICA CORP /DE/

$299,367,107

0.74%

994,476

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	586,016	$176,408,396	58.9%	Defined
MERRILL LYNCH, PIERCE, FENNER & SMITH INC.	20,420	$6,147,032	2.1%	Defined
BOFA SECURITIES, INC.	140,417	$42,269,729	14.1%	Defined
MERRILL LYNCH INTERNATIONAL	16,614	$5,001,312	1.7%	Defined
MANAGED ACCOUNT ADVISORS LLC joint	223,952	$67,416,270	22.5%	Defined
BofA Securities Europe SA	7,057	$2,124,368	0.7%	Defined

Legal & General Group Plc

$278,320,596

0.68%

924,561

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	323,780	$97,467,493	35.0%	Defined
Legal & General Investment Management Ltd joint	600,781	$180,853,103	65.0%	Defined

MANUFACTURERS LIFE INSURANCE COMPANY, THE

$269,059,108

0.66%

893,795

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MANULIFE FINANCIAL CORP joint	893,795	$269,059,108	100.0%	Defined

NORTHERN TRUST CORP

$266,983,504

0.66%

886,900

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	687,322	$206,904,540	77.5%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	177,360	$53,390,680	20.0%	Defined
Northern Trust Asset Management Australia Pty Ltd	7,881	$2,372,417	0.9%	Defined
Northern Trust Co of Hong Kong Ltd	14,337	$4,315,867	1.6%	Defined

ROYAL BANK OF CANADA

$263,659,830

0.65%

875,859

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC Global Asset Management Inc.	514,893	$154,998,239	58.8%	Defined
RBC CMA LTD.	2,226	$670,092	0.3%	Defined
RBC Capital Markets, LLC	61,059	$18,380,590	7.0%	Defined
RBC Dominion Securities Inc.	31,703	$9,543,554	3.6%	Defined
RBC Phillips, Hager & North Investment Counsel Inc.	93	$27,995	0.0%	Defined
RBC Private Counsel (USA) Inc.	559	$168,275	0.1%	Defined
RBC Trust Co (Delaware) Ltd	138	$41,542	0.0%	Defined
CITY NATIONAL BANK	580	$174,597	0.1%	Defined
RBC Rochdale, LLC	1,803	$542,757	0.2%	Defined
Held directly	262,805	$79,112,189	30.0%	Sole

VANGUARD FIDUCIARY TRUST CO

$259,723,265

0.64%

862,782

Deep Track Capital, LP

$248,349,750

0.61%

825,000

DEUTSCHE BANK AG\

$245,459,858

0.60%

815,400

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Deutsche Bank Trust Co Americas	3	$903	0.0%	Defined
DWS Investment GmbH	58,287	$17,546,135	7.1%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC	18,782	$5,653,945	2.3%	Defined
DWS Investments UK Ltd	360,796	$108,610,419	44.2%	Defined
DBX Advisors LLC	5,660	$1,703,829	0.7%	Defined
DWS Investments Hong Kong Ltd	6,507	$1,958,802	0.8%	Defined
DWS International GmbH	66,893	$20,136,799	8.2%	Defined
Held directly	298,472	$89,849,026	36.6%	Sole

Qube Research & Technologies Ltd

$240,437,176

0.59%

798,715

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$227,391,137

0.56%

755,377

LOS ANGELES CAPITAL MANAGEMENT LLC

$219,642,024

0.54%

729,635

AQR CAPITAL MANAGEMENT LLC

$219,161,577

0.54%

728,039

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
KEB ASSET MANAGEMENT, LLC	12	$3,612	0.0%	Other
RF&L WEALTH MANAGEMENT, LLC	1	$301	0.0%	Other
Prism Planning Partners LLC	96	$28,898	0.0%	Other
Compound Planning, Inc.	6	$1,806	0.0%	Other
Arax Advisory Partners	575	$173,092	0.1%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	1,761	$530,113	0.2%	Other
Stokes Family Office, LLC	737	$221,859	0.1%	Other
NATIONAL MUTUAL FUNDS MANAGEMENT LTD	4,544	$1,367,880	0.6%	Other
Held directly	720,307	$216,834,016	98.9%	Defined

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$219,104,078

0.54%

727,848

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	80,464	$24,222,077	11.1%	Defined
UBS Asset Management Switzerland AG	285,400	$85,913,961	39.2%	Defined
UBS Asset Management (UK) Ltd.	275,156	$82,830,209	37.8%	Defined
UBS Asset Management Life Ltd.	5,715	$1,720,386	0.8%	Defined
Held directly	81,113	$24,417,445	11.1%	Defined

FEDERATED HERMES, INC.

$216,713,000

0.53%

719,905

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FEDERATED EQUITY MANAGEMENT CO OF PENNSYLVANIA	3,664	$1,102,973	0.5%	Defined
FEDERATED MDTA LLC	708,065	$213,148,806	98.4%	Defined
HERMES INVESTMENT MANAGEMENT LTD	8,176	$2,461,221	1.1%	Defined

UBS Group AG

$188,747,012

0.46%

627,004

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	46,272	$13,929,260	7.4%	Defined
UBS AG	223,582	$67,304,889	35.7%	Defined
UBS Switzerland AG	355,617	$107,051,385	56.7%	Defined
UBS Securities LLC	1,533	$461,478	0.2%	Defined

Amundi

$185,877,294

0.46%

617,471

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Amundi Asset Management joint	611,030	$183,938,360	99.0%	Defined
CPR Asset Management	6,441	$1,938,934	1.0%	Defined

LOOMIS SAYLES & CO L P

$174,653,692

0.43%

580,187

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
LOOMIS SAYLES TRUST COMPANY, LLC	13,943	$4,197,261	2.4%	Defined
Held directly	566,244	$170,456,431	97.6%	Defined

GOLDMAN SACHS GROUP INC

$173,019,098

0.43%

574,757

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	123,010	$37,029,700	21.4%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	445,738	$134,180,510	77.6%	Defined
GOLDMAN SACHS INTERNATIONAL	3,508	$1,056,013	0.6%	Defined
GOLDMAN SACHS BANK AG	865	$260,390	0.2%	Defined
Goldman Sachs Trust Company, N.A.	1,636	$492,485	0.3%	Defined

Bank of New York Mellon Corp

$148,128,128

0.36%

492,071

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNY Mellon Trust of Delaware	75	$22,577	0.0%	Defined
Mellon Investments Corporation joint	345,788	$104,092,561	70.3%	Defined
BNY Mellon Securities Corporation	31	$9,331	0.0%	Defined
The Bank of New York Mellon	114,641	$34,510,380	23.3%	Defined
BNY Mellon, NA joint	10,920	$3,287,246	2.2%	Defined
BNY Mellon Advisors, Inc. joint	15,359	$4,623,519	3.1%	Defined
Newton Investment Management North America, LLC joint	5,256	$1,582,213	1.1%	Defined
Held directly	1	$301	0.0%	Defined

Polar Capital Holdings Plc

$145,322,533

0.36%

482,751

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Polar Capital LLP	482,751	$145,322,533	100.0%	Defined

Paradigm Biocapital Advisors LP

$144,494,400

0.35%

480,000

FRANKLIN RESOURCES INC

$134,770,826

0.33%

447,699

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Brandywine Global Investment Management, LLC	136	$40,940	0.0%	Defined
FIDUCIARY TRUST CO INTERNATIONAL	368	$110,779	0.1%	Defined
Fiduciary Trust Co International of Pennsylvania	800	$240,824	0.2%	Defined
Fiduciary Trust International of the South	127	$38,230	0.0%	Defined
FIDUCIARY TRUST INTERNATIONAL LLC	2,125	$639,688	0.5%	Defined
FRANKLIN ADVISERS INC	369,655	$111,277,244	82.6%	Defined
FRANKLIN ADVISORY SERVICES LLC	54,860	$16,514,505	12.3%	Defined
PUTNAM INVESTMENT MANAGEMENT LLC	19,628	$5,908,616	4.4%	Defined

D. E. Shaw & Co., Inc.

$129,533,209

0.32%

430,300

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	430,300	$129,533,209	100.0%	Defined

FIRST TRUST ADVISORS LP

$126,894,380

0.31%

421,534

RAYMOND JAMES FINANCIAL INC

$118,244,281

0.29%

392,799

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	81,925	$24,661,882	20.9%	Defined
Raymond James Financial Services Advisors, Inc.	4,911	$1,478,358	1.3%	Defined
ClariVest Asset Management LLC joint	6	$1,806	0.0%	Defined
EAGLE ASSET MANAGEMENT INC joint	304,904	$91,785,251	77.6%	Defined
RAYMOND JAMES TRUST N.A.	1,053	$316,984	0.3%	Defined

DIMENSIONAL FUND ADVISORS LP

$118,010,380

0.29%

392,022

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	6,308	$1,898,897	1.6%	Defined
DFA Australia Ltd.	33,295	$10,022,793	8.5%	Defined
Dimensional Fund Advisors Pte. Ltd.	1,763	$530,715	0.4%	Defined
Dimensional Ireland Ltd	36,110	$10,870,193	9.2%	Defined
Held directly	314,546	$94,687,782	80.2%	Sole

Swiss National Bank

$112,609,302

0.28%

374,080

WELLS FARGO & COMPANY/MN

$111,381,100

0.27%

370,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	370,000	$111,381,100	100.0%	Defined

MASSACHUSETTS FINANCIAL SERVICES CO /MA/

$105,529,075

0.26%

350,560

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MFS Institutional Advisors, Inc.	36,368	$10,947,859	10.4%	Defined
MFS Investment Management CO (LUX) S.A.R.L.	2,073	$624,035	0.6%	Defined
MFS Investment Management K.K.	2,012	$605,672	0.6%	Defined
MFS Heritage Trust CO	55,689	$16,764,059	15.9%	Defined
MFS INVESTMENT MANAGEMENT CANADA LTD	581	$174,898	0.2%	Defined
Held directly	253,837	$76,412,552	72.4%	Sole

CANADA PENSION PLAN INVESTMENT BOARD

$100,618,073

0.25%

334,246

Sumitomo Mitsui Trust Group, Inc.

$97,948,840

0.24%

325,379

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUMITOMO MITSUI TRUST BANK Ltd	325,379	$97,948,840	100.0%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$97,804,647

0.24%

324,900

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	324,900	$97,804,647	100.0%	Other

MILLENNIUM MANAGEMENT LLC

$95,125,781

0.23%

316,001

Nuveen, LLC

$95,014,096

0.23%

315,630

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	29,148	$8,774,421	9.2%	Defined
TIAA CREF INVESTMENT MANAGEMENT LLC joint	15,565	$4,685,531	4.9%	Defined
TEACHERS ADVISORS, LLC joint	270,917	$81,554,144	85.8%	Defined

Candriam S.C.A.

$94,677,547

0.23%

314,512

WELLS FARGO & COMPANY/MN

$93,333,446

0.23%

310,047

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	126,661	$38,128,760	40.9%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	51,357	$15,459,997	16.6%	Defined
WELLS FARGO CLEARING SERVICES, LLC	117,443	$35,353,866	37.9%	Defined
Wells Fargo Securities, LLC	9,945	$2,993,743	3.2%	Defined
WELLS FARGO INVESTMENT INSTITUTE, INC.	4,641	$1,397,080	1.5%	Defined

NOMURA ASSET MANAGEMENT INTERNATIONAL INC.

$90,367,098

0.22%

300,193

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NOMURA HOLDINGS INC joint	300,193	$90,367,098	100.0%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$88,641,293

0.22%

294,460

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	294,460	$88,641,293	100.0%	Other

Pictet Asset Management Holding SA

$83,870,871

0.21%

278,613

BNP PARIBAS ARBITRAGE, SA

$81,586,053

0.20%

271,023

GILDER GAGNON HOWE & CO LLC

$80,992,422

0.20%

269,051

CONGRESS ASSET MANAGEMENT CO

$79,983,972

0.20%

265,701

MARSHALL WACE, LLP

$78,207,292

0.19%

259,799

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marshall Wace North America L.P. joint	259,799	$78,207,292	100.0%	Other

VANGUARD ASSET MANAGEMENT, Ltd

$77,578,441

0.19%

257,710

California Public Employees Retirement System

$76,477,574

0.19%

254,053

WESTFIELD CAPITAL MANAGEMENT CO LP

$75,974,854

0.19%

252,383

Korea Investment CORP

$74,553,089

0.18%

247,660

Woodline Partners LP

$74,138,270

0.18%

246,282

Squarepoint Ops LLC

$70,440,718

0.17%

233,999

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Squarepoint Capital LLP	233,999	$70,440,718	100.0%	Defined

TD Asset Management Inc

$70,349,506

0.17%

233,696

CITIGROUP INC

$68,570,419

0.17%

227,786

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citibank, N.A. joint	22,902	$6,894,189	10.1%	Defined
Citigroup Financial Products Inc. joint	204,884	$61,676,230	89.9%	Defined

ROYAL LONDON ASSET MANAGEMENT LTD

$64,296,696

0.16%

213,589

AMERIPRISE FINANCIAL INC

$61,591,036

0.15%

204,601

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	93,649	$28,191,158	45.8%	Defined
Threadneedle Asset Management Holdings LTD	590	$177,607	0.3%	Defined
Ameriprise Financial Services, LLC	14,230	$4,283,656	7.0%	Defined
Ameriprise Bank, FSB	167	$50,272	0.1%	Defined
Columbia Threadneedle Management Ltd	95,965	$28,888,343	46.9%	Defined

National Pension Service

$59,692,743

0.15%

198,295

PINEBRIDGE INVESTMENTS LLC

$58,914,280

0.14%

195,709

CITADEL ADVISORS LLC

$57,505,458

0.14%

191,029

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	177,597	$53,462,024	93.0%	Defined
Held directly	13,432	$4,043,434	7.0%	Defined

Robeco Institutional Asset Management B.V.

$56,618,926

0.14%

188,084

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$53,824,164

0.13%

178,800

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	178,800	$53,824,164	100.0%	Other

VICTORY CAPITAL MANAGEMENT INC

$52,030,627

0.13%

172,842

Man Group plc

$51,448,133

0.13%

170,907

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AHL Partners LLP	51,988	$15,649,947	30.4%	Defined
Numeric Investors LLC	118,919	$35,798,186	69.6%	Defined

Mitsubishi UFJ Asset Management Co., Ltd.

$50,772,923

0.12%

168,664

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MITSUBISHI UFJ FINANCIAL GROUP INC	168,664	$50,772,923	100.0%	Defined

RHUMBLINE ADVISERS

$49,844,246

0.12%

165,579

Life Cycle Investment Partners Ltd

$49,797,586

0.12%

165,424

CIBC Bancorp USA Inc.

$49,288,543

0.12%

163,733

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CIBC Private Wealth Advisors, Inc.	92,541	$27,857,617	56.5%	Defined
CIBC National Trust Co	64,062	$19,284,583	39.1%	Defined
CIBC WORLD MARKETS CORP	7,130	$2,146,343	4.4%	Defined

ANTIPODES PARTNERS Ltd

$49,228,338

0.12%

163,533

BNP PARIBAS ASSET MANAGEMENT Holding S.A.

$48,698,525

0.12%

161,773

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNP PARIBAS ASSET MANAGEMENT Europe	96,408	$29,021,700	59.6%	Defined
AXA INVESTMENT MANAGERS UK LIMITED	65,365	$19,676,825	40.4%	Defined

Russell Investments Group, Ltd.

$47,631,369

0.12%

158,228

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management Limited	8,837	$2,660,202	5.6%	Defined
Russell Investment Management, LLC joint	60,058	$18,079,258	38.0%	Defined
Russell Investments Canada Limited	38,017	$11,444,257	24.0%	Defined
Russell Investments Capital, LLC	1,104	$332,337	0.7%	Defined
Russell Investments Implementation Services, LLC	3,473	$1,045,477	2.2%	Defined
Russell Investments Limited joint	44,437	$13,376,868	28.1%	Other
Russell Investments Trust Company joint	2,302	$692,970	1.5%	Other

Universal- Beteiligungs- und Servicegesellschaft mbH

$47,328,839

0.12%

157,223

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Universal-Investment-Gesellschaft mbH joint	157,223	$47,328,839	100.0%	Defined

CAPITAL FUND MANAGEMENT S.A.

$46,931,781

0.12%

155,904

Assenagon Asset Management S.A.

$46,868,865

0.12%

155,695

Vinva Investment Management Ltd

$46,651,221

0.11%

154,972

Hudson Bay Capital Management LP

$44,769,181

0.11%

148,720

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Hudson Bay Capital UK LLP	8,520	$2,564,775	5.7%	Defined
Held directly	140,200	$42,204,406	94.3%	Sole

ENVESTNET ASSET MANAGEMENT INC

$41,058,986

0.10%

136,395

Jupiter Topco LLC

$40,803,110

0.10%

135,545

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	9,392	$2,827,273	6.9%	Defined
JANUS HENDERSON INVESTORS US LLC	120,453	$36,259,966	88.9%	Defined
Held directly	5,700	$1,715,871	4.2%	Defined

BARCLAYS PLC

$40,794,982

0.10%

135,518

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	92,945	$27,979,233	68.6%	Defined
BARCLAYS CAPITAL INC.	3,818	$1,149,332	2.8%	Defined
BARCLAYS CAPITAL SECURITIES LTD	38,755	$11,666,417	28.6%	Defined

LMR Partners LLP

$40,402,439

0.10%

134,214

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wabon Partners, L.P.	119,196	$35,881,571	88.8%	Sole
Held directly joint	15,018	$4,520,868	11.2%	Other

ProShare Advisors LLC

$38,686,569

0.10%

128,514

SEI INVESTMENTS CO

$36,713,618

0.09%

121,960

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI INVESTMENTS MANAGEMENT CORP	19,279	$5,803,557	15.8%	Sole
SEI TRUST CO	102,681	$30,910,061	84.2%	Sole

Key facts CIK 1178670 CUSIP 02043Q107 13F (30d) 607 filings 601 filers Visit website Investor relations