AXGN · Axogen, Inc. · Holdings

$36.17 +0.93 (+2.64%) At close · Oct 20

Russell 2000 member Rebalance outlook locked

Market Cap

$1.96B

53.76M

Volume · Oct 20 974.62K Avg daily vol (3M) 1.21M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Use institutional holdings through the 13F API or MCP → See every holder of AXGN: 13F, 13D and 13G, insiders and funds →

Institutional holdings are being reconciled with SEC filings. Holder totals and changes may be incomplete until reconciliation finishes.

The Sep 30, 2026 filing window is still open — showing the latest known positions. Funds that have not filed yet carry their prior-quarter positions; a fund that filed without this stock counts as sold out. Each fund's Change is measured against its own previous reported quarter, shown in the Reported column. Pick a completed quarter in the Quarter selector below to put every fund on one quarter and see the stock-wide activity breakdown.

Quarter:

Holders: 254

Total Value: $2,395,874,749

Total Shares: 52,201,050

Showing 1–100 of 257 holdings by value · page 1 of 3

Institutional Ownership Trend

27 quarters

Holder Concentration

Across 254 13F filers

Top Holder

8.4%

of institutional value

Top 5

26.0%

of institutional value

Top 10

38.9%

of institutional value

HHI

253

Broadly held

Who Holds AXGN

Last 8 quarters through Sep 30, 2026 (filing window open)

Each kind of 13F filer's share of the common shares 13F filers report. Holder type comes from regulatory records, never the filer's name; style, turnover and horizon are measured from each filer's own 13F book. The quarter selector does not change this card.

Jun 30, 2026, the newest closed quarter: 253 filers, like-for-like change +5.5%; actively managed books +3.0%; index-like books +11.9%.

Holder type	Share	Filers	Change	New	Exited
Registered adviser	52.2%	132	-4.8%	27	10
Exempt adviser	1.5%	5	+5.4%	1	1
Adviser	3.3%	8	+47.2%	4	0
Bank	6.7%	16	+36.3%	1	0
Insurer	0.2%	2	+5.6%	0	0
Broker-dealer	0.0%	2	-16.3%	0	0
Benefit plan	0.2%	4	-29.3%	1	1
Non-US	4.7%	10	+104.5%	2	0
Holding company	21.1%	7	+12.5%	0	0
Unknown	10.0%	67	+2.6%	13	6

Style	Share	Filers	Change	New	Exited
Index-like	29.7%	30	+11.9%	3	1
Systematic-like	6.1%	18	-10.2%	5	3
Concentrated	6.8%	21	-24.3%	5	2
Diversified active	57.4%	184	+9.4%	36	12
Not measured	0.0%	0	—	0	0
Unknown	0.0%	0	—	0	0

Turnover	Share	Filers	Change	New	Exited
Low turnover	41.5%	122	+13.4%	16	7
Medium turnover	40.5%	78	+7.0%	18	3
High turnover	10.8%	38	-23.9%	11	7
Unknown	7.2%	15	+17.0%	4	1
Not measured	0.0%	0	—	0	0

Holding horizon	Share	Filers	Change	New	Exited
Short horizon	9.2%	32	+3.7%	13	7
Medium horizon	12.1%	35	+0.3%	5	2
Long horizon	59.8%	166	+11.2%	28	7
Unknown	18.9%	20	-5.7%	3	2
Not measured	0.0%	0	—	0	0

13F book size	Share	Filers	Change	New	Exited
Under $1B	3.2%	38	-16.6%	11	5
$1B to $10B	17.4%	66	-34.6%	12	11
$10B to $100B	28.2%	82	+28.5%	19	2
$100B or more	51.3%	67	+20.7%	7	0
Not measured	0.0%	0	—	0	0

Client base	Share	Filers	Change	New	Exited
Individuals and high net worth	3.0%	48	-12.1%	10	3
Pooled investment vehicles	20.1%	29	-19.6%	11	6
Registered funds and BDCs	19.3%	26	+40.0%	6	0
Pension and profit sharing plans	0.0%	1	+0.2%	0	0
Insurance companies	0.1%	3	+21.4%	0	0
Governments and sovereign funds	0.7%	5	+8.3%	0	0
Other investment advisers	0.0%	3	-4.3%	0	0
Diversified	8.9%	17	-24.3%	0	0
Other	0.0%	0	—	0	0
Unknown	47.8%	121	+19.7%	22	9

Private funds	Share	Filers	Change	New	Exited
Hedge funds	28.7%	49	-14.9%	13	7
Private equity funds	2.6%	8	+369.4%	2	0
Venture capital funds	0.0%	2	0.0%	0	0
Real estate funds	0.0%	3	+84.1%	1	1
Securitized asset funds	0.2%	1	+30.9%	0	0
Other private funds	7.5%	15	+9.9%	3	0
Mixed private funds	1.1%	3	-46.3%	0	1
Private funds, type not stated	0.1%	3	+7.1%	2	0
No private funds	13.6%	53	+4.8%	7	2
Unknown	46.2%	116	+20.2%	21	7

Location	Share	Filers	Change	New	Exited
United States	85.5%	205	+2.1%	40	15
Outside the US	14.5%	48	+31.6%	9	3

The Sep 30, 2026 filing window is still open, so its bar carries the Jun 30, 2026 positions of filers that have not filed yet.

How the mix is measured

Shares are 13F common shares restated to today's split basis; options and principal amounts are left out. The change compares only filers with a 13F in both quarters and keeps each filer in its bucket of the later quarter on both sides; a filer that continues under a new CIK is compared through its successor. Holder type comes from each filer's current regulatory profile; client base and private funds come from the Form ADV roster current at each quarter end (rosters from December 2019, so an earlier quarter reads Unknown). Style, turnover, horizon and book size are measured from each filer's 13F book in the newest measured quarter, and Not measured means none was. Actively managed books are those measured systematic-like, concentrated or diversified active.

Excluded 2 holders with no observed 2026-03-31 13F and 2 holders with no observed 2026-06-30 13F. Compared across CIK changes: JANUS HENDERSON GROUP PLC continues as Jupiter Topco LLC.

Conviction and Crowding

253 13F filers, quarter ended Jun 30, 2026

Measured across every 13F filer's common-share book once the quarter's filing window closed. Crowding counts filers with high turnover or that advise private funds on Form ADV.

Conviction holders

2.77% of filers hold it in their top 10

Mar 31, 2026: 6

Holders above 5%

median weight 0.003%

Mar 31, 2026: 3

Crowding

42.8%

of shares outstanding, 94th percentile

Mar 31, 2026: 48.8%

Private-fund advisers

39.8%

of shares outstanding

Institutional ownership

99.0%

101.3% of the estimated free float above 100%: filers can report lent or client shares more than once

Top 10 holders

38.9%

of 13F shares, Herfindahl 0.025

Active holders

+3.0%

like-for-like share change of active filers

5% stakes on record

0 13D, 3 13G (3 institutional, 0 passive, 0 exempt)

How long holders have held it value-weighted 12.0 quarters

Under 1 year 35.2% 1 to 3 years 26.3% 3 years or more 38.5% Since coverage start 0.000%

Managers in the top fifth by current three-year FundScore alpha were net sellers of $3,023,830 (0 buying, 1 selling).

Estimated Holder Cost Basis

Quarter ended Jun 30, 2026, on that quarter's share basis

Estimated, never filed: each 13F buy is costed at its quarter's average traded price and each sale keeps the average cost, so this is what holders paid on average if they traded evenly through each quarter.

Estimated avg cost

$27.63

estimated per share

Mar 31, 2026: $21.34

Quarter-end price

$46.19

implied by 13F values

Estimated gain or loss

+67.2%

price against the estimated cost

Estimated at a loss

0.000%

of costed 13F value

Estimated cost covers 60.4% of the 13F shares; positions already held when a filer's 13F history starts, or bought in a quarter without a trusted average price, have no estimate.

Shared Holders

Companies AXGN's active holders also own, quarter ended Jun 30, 2026

#	Company	Shared holders	Similarity
1	NPCE NeuroPace Inc	4	0.249
2	ATRC AtriCure, Inc.	6	0.226
3	VCYT Veracyte, Inc.	8	0.213
4	NEO Neogenomics Inc	6	0.210
5	TLSI TriSalus Life Sciences, Inc.	4	0.208
6	TCMD Tactile Systems Technology Inc	5	0.205
7	TNDM Tandem Diabetes Care Inc	5	0.205
8	XGN Exagen Inc.	3	0.186
9	IRTC iRhythm Holdings, Inc.	6	0.183
10	VCEL Vericel Corp	6	0.180

Shareholder Targets

13F filers that own its peers but not AXGN, quarter ended Jun 30, 2026

Found through its holder-overlap peers: NPCE, ATRC, VCYT, NEO, TLSI, TCMD, TNDM, XGN, IRTC, VCEL, UTI, KNSA, ALHC, HTFL, RIGL, IMVT, MIRM, BDSX, KRMD, KRYS. 818 filers in all.

#	Filer	In peers	Of book	Largest peers
1	Frazier Life Sciences Management, L.P. Registered adviser, Concentrated	$1,077,947,977 2 held	23.5%	MIRM, KRYS
2	Avoro Capital Advisors LLC Registered adviser, Concentrated	$1,073,713,002 1 held	9.64%	KRYS
3	WILLIAM BLAIR INVESTMENT MANAGEMENT, LLC Registered adviser, Diversified active	$733,900,549 5 held	2.19%	KRYS, ALHC, MIRM
4	Artisan Partners Limited Partnership Registered adviser, Diversified active	$522,763,983 2 held	0.850%	VCYT, IRTC
5	Capital World Investors Adviser, Diversified active	$517,239,029 3 held	0.060%	KRYS, MIRM, ALHC
6	BAKER BROS. ADVISORS LP Registered adviser, Concentrated	$442,904,738 3 held	2.23%	IMVT, KNSA, MIRM
7	T. Rowe Price Investment Management, Inc. Adviser, Diversified active	$415,740,866 3 held	0.270%	ALHC, IMVT, NEO
8	Deep Track Capital, LP Adviser, Diversified active	$359,914,548 1 held	5.48%	IMVT
9	TCG Crossover Management, LLC Unknown, Diversified active	$296,100,353 2 held	7.12%	IMVT, MIRM
10	TANG CAPITAL MANAGEMENT LLC Unknown, Concentrated	$289,221,581 2 held	13.2%	MIRM, KNSA

Fund Ownership of AXGN by Fund Type

Registered funds' Form N-PORT reports for the quarter ended Jun 30, 2026

Index funds and index ETFs hold 13.90% of shares outstanding.

Fund type	Of shares outstanding	Change	Shares	Funds
Index funds	3.17%	+0.62 pp	1,704,538	57
Index ETFs	10.73%	+0.86 pp	5,770,242	39
Active ETFs	0.99%	-0.06 pp	531,459	19
Active funds	15.18%	+5.66 pp	8,161,755	105
Unclassified	0.14%	+0.11 pp	77,194	8
All registered funds	30.22%	+7.19 pp	16,245,188	228

How fund types are measured

Each fund series counts once, with its latest Form N-PORT report dated inside the quarter; only long share positions count, restated to today's split basis. The fund type is the series' latest Form N-CEN answer on or before the quarter end (the index-fund box and the exchange-traded flag), never its name. Unclassified funds have no usable N-CEN; unit investment trusts are among them, so index ownership is a floor. Funds sit inside their advisers' 13F reports, so these shares overlap the 13F figures and are never added to them.

Holder type: Registered adviser 133 Exempt adviser 5 Adviser 8 Bank 16 Insurer 2 Broker-dealer 2 Benefit plan 4 Non-US 10 Holding company 7 Unknown 67

Showing 100 of 257

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

BlackRock, Inc.

$204,123,211

8.52%

4,419,208

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	40,787	$1,883,951	0.9%	Sole
BlackRock Investment Management (Australia) Ltd	193	$8,914	0.0%	Sole
BlackRock Asset Management Canada Ltd	10,546	$487,119	0.2%	Sole
BlackRock Investment Management, LLC	144,180	$6,659,674	3.3%	Sole
BLACKROCK ADVISORS LLC	115,177	$5,320,025	2.6%	Sole
BlackRock Fund Advisors	2,280,469	$105,334,863	51.6%	Sole
BlackRock Institutional Trust Company, N.A.	1,573,924	$72,699,549	35.6%	Sole
BlackRock Fund Managers Ltd.	9,810	$453,123	0.2%	Sole
BlackRock Investment Management (UK) Ltd.	13,854	$639,916	0.3%	Sole
BlackRock Asset Management Ireland Ltd	168,501	$7,783,061	3.8%	Sole
BlackRock Life Ltd	482	$22,263	0.0%	Sole
BlackRock Asset Management Schweiz AG	7,823	$361,344	0.2%	Sole
Aperio Group, LLC	53,462	$2,469,409	1.2%	Sole

First Light Asset Management, LLC

$126,946,794

5.30%

2,748,361

MORGAN STANLEY

$126,926,606

5.30%

2,747,924

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	2,166	$100,047	0.1%	Defined
MORGAN STANLEY & CO. LLC	19,793	$914,238	0.7%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	2,444,625	$112,917,228	89.0%	Defined
MORGAN STANLEY SMITH BARNEY LLC	132,774	$6,132,831	4.8%	Defined
Parametric Sas	1,208	$55,797	0.0%	Defined
Morgan Stanley Bank, N.A.	37,144	$1,715,681	1.4%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	110,214	$5,090,784	4.0%	Defined

VANGUARD CAPITAL MANAGEMENT LLC

$104,294,618

4.35%

2,257,948

Artemis Investment Management LLP

$74,659,184

3.12%

1,780,992

Polar Capital Holdings Plc

$69,322,968

2.89%

1,500,822

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Polar Capital LLP	1,500,822	$69,322,968	100.0%	Defined

WELLINGTON MANAGEMENT GROUP LLP

$64,280,496

2.68%

1,391,654

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wellington Management Company LLP	727,236	$33,591,030	52.3%	Defined
Wellington Management Singapore Pte. Ltd.	57,261	$2,644,885	4.1%	Defined
Wellington Management International Ltd	172,600	$7,972,394	12.4%	Defined
Wellington Trust Company, NA joint	434,557	$20,072,187	31.2%	Defined

GEODE CAPITAL MANAGEMENT, LLC

$63,522,334

2.65%

1,375,240

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	195,188	$9,015,733	14.2%	Defined
Held directly	1,180,052	$54,506,601	85.8%	Defined

CITADEL ADVISORS LLC

$62,856,783

2.62%

1,360,831

GOLDMAN SACHS GROUP INC

$58,269,562

2.43%

1,261,519

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	1,018,533	$47,046,039	80.7%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	242,986	$11,223,523	19.3%	Defined

STATE STREET CORP

$57,590,661

2.40%

1,246,821

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	1,246,821	$57,590,661	100.0%	Defined

FMR LLC

$56,484,087

2.36%

1,222,864

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	1,120,339	$51,748,458	91.6%	Defined
STRATEGIC ADVISERS LLC	397	$18,337	0.0%	Defined
Fidelity Institutional Asset Management Trust Co	91,900	$4,244,861	7.5%	Defined
FIAM LLC	10,228	$472,431	0.8%	Defined

Hood River Capital Management LLC

$49,288,055

2.06%

1,067,072

WASATCH ADVISORS LP

$47,024,095

1.96%

1,121,758

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 1	1,121,758	$47,024,095	100.0%	Sole

PRICE T ROWE ASSOCIATES INC /MD/

$47,023,359

1.96%

1,018,042

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	73,892	$3,413,071	7.3%	Defined
Held directly	944,150	$43,610,288	92.7%	Sole

RTW INVESTMENTS, LP

$46,724,695

1.95%

1,011,576

NEXT CENTURY GROWTH INVESTORS LLC

$40,302,899

1.68%

872,546

UBS Group AG

$38,624,492

1.61%

836,209

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	5,382	$248,594	0.6%	Defined
UBS AG	826,560	$38,178,806	98.8%	Defined
UBS Switzerland AG	3,479	$160,695	0.4%	Defined
UBS Securities LLC	788	$36,397	0.1%	Defined

ARMISTICE CAPITAL, LLC

$35,797,250

1.49%

775,000

BAMCO INC /NY/

$34,190,299

1.43%

740,210

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARON CAPITAL MANAGEMENT INC /NY/ joint	740,210	$34,190,299	100.0%	Defined

ArrowMark Colorado Holdings LLC

$32,877,811

1.37%

711,795

Driehaus Capital Management LLC

$32,667,138

1.36%

707,234

TimesSquare Capital Management, LLC

$31,863,386

1.33%

689,833

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GuideStone Capital Management, LLC joint	689,833	$31,863,386	100.0%	Sole

BESSEMER GROUP INC

$31,359,405

1.31%

678,970

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BESSEMER TRUST CO	12,158	$561,577	1.8%	Defined
BESSEMER TRUST Co NATIONAL ASSOCIATION	559,631	$25,849,355	82.4%	Defined
Bessemer Trust CO of Delaware, N.A.	49,155	$2,270,469	7.2%	Defined
BESSEMER TRUST Co OF FLORIDA	55,684	$2,572,043	8.2%	Defined
Bessemer Trust Co of Nevada, National Association	2,342	$105,961	0.3%	Defined

DEUTSCHE BANK AG\

$30,105,902

1.26%

651,784

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DWS INVESTMENT MANAGEMENT AMERICAS, INC	41,194	$1,902,750	6.3%	Defined
DWS Investments UK Ltd	610,590	$28,203,152	93.7%	Defined

VANGUARD PORTFOLIO MANAGEMENT LLC

$26,710,060

1.11%

578,265

Nuveen, LLC

$25,214,750

1.05%

545,892

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	30,463	$1,407,085	5.6%	Defined
TIAA CREF INVESTMENT MANAGEMENT LLC joint	84,050	$3,882,269	15.4%	Defined
TEACHERS ADVISORS, LLC joint	431,379	$19,925,396	79.0%	Defined

Holocene Advisors, LP

$24,793,175

1.03%

536,765

NORTHERN TRUST CORP

$22,151,474

0.92%

479,573

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	443,128	$20,468,081	92.4%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	28,283	$1,306,391	5.9%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	8,162	$377,002	1.7%	Other

Capricorn Fund Managers Ltd

$21,472,021

0.90%

464,863

NOMURA ASSET MANAGEMENT INTERNATIONAL INC.

$21,058,021

0.88%

455,900

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NOMURA HOLDINGS INC joint	455,900	$21,058,021	100.0%	Defined

ARROWSTREET CAPITAL, LIMITED PARTNERSHIP

$20,463,324

0.85%

443,025

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$20,393,808

0.85%

441,520

DIMENSIONAL FUND ADVISORS LP

$19,000,763

0.79%

411,361

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	4,033	$186,284	1.0%	Defined
Dimensional Ireland Ltd	19,686	$909,296	4.8%	Defined
Held directly	387,642	$17,905,183	94.2%	Sole

OBERWEIS ASSET MANAGEMENT INC/

$17,930,680

0.75%

388,194

JACOBS LEVY EQUITY MANAGEMENT, INC

$17,848,878

0.74%

386,423

VANGUARD FIDUCIARY TRUST CO

$17,492,106

0.73%

378,699

Assenagon Asset Management S.A.

$17,259,050

0.72%

411,714

FRED ALGER MANAGEMENT, LLC

$16,232,967

0.68%

351,439

HEALTHCARE OF ONTARIO PENSION PLAN TRUST FUND

$16,071,902

0.67%

347,952

HighMark Wealth Management LLC

$15,768,804

0.66%

341,390

First Eagle Investment Management, LLC

$15,754,346

0.66%

341,077

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FIRST EAGLE SEPARATE ACCOUNT MANAGEMENT, LLC	122	$5,635	0.0%	Sole
Held directly	340,955	$15,748,711	100.0%	Sole

WCM INVESTMENT MANAGEMENT, LLC

$15,570,602

0.65%

337,099

RAYMOND JAMES FINANCIAL INC

$15,550,415

0.65%

336,662

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	20,410	$942,737	6.1%	Defined
Raymond James Financial Services Advisors, Inc.	56,794	$2,623,314	16.9%	Defined
EAGLE ASSET MANAGEMENT INC joint	259,458	$11,984,364	77.1%	Defined

TWO SIGMA INVESTMENTS, LP

$14,743,986

0.62%

319,203

BANK OF AMERICA CORP /DE/

$14,475,899

0.60%

313,399

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	234,797	$10,845,273	74.9%	Defined
BOFA SECURITIES, INC.	54,379	$2,511,766	17.4%	Defined
MANAGED ACCOUNT ADVISORS LLC	24,223	$1,118,860	7.7%	Defined

MARSHALL WACE, LLP

$13,778,522

0.58%

298,301

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marshall Wace North America L.P. joint	293,774	$13,569,420	98.5%	Other
Marshall Wace Middle East Ltd	4,527	$209,102	1.5%	Other

NZS Capital, LLC

$12,848,672

0.54%

278,170

D. E. Shaw & Co., Inc.

$12,828,718

0.54%

277,738

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	277,738	$12,828,718	100.0%	Defined

FIRST TRUST ADVISORS LP

$12,525,065

0.52%

271,164

Stephens Investment Management Group LLC

$11,958,821

0.50%

258,905

Balyasny Asset Management L.P.

$11,709,072

0.49%

253,498

SILVERCREST ASSET MANAGEMENT GROUP LLC

$11,635,030

0.49%

251,895

PERCEPTIVE ADVISORS LLC

$11,423,341

0.48%

247,312

BROWN CAPITAL MANAGEMENT LLC

$10,402,311

0.43%

225,207

Point72 Asset Management, L.P.

$10,375,105

0.43%

224,618

Atika Capital Management LLC

$10,207,988

0.43%

221,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Atika Capital Partners LP	128,929	$5,955,230	58.3%	Defined
Optika Master Portfolio Ltd	55,856	$2,579,988	25.3%	Other
Held directly	36,215	$1,672,770	16.4%	Sole

DAFNA Capital Management LLC

$9,651,862

0.40%

208,960

Uniplan Investment Counsel, Inc.

$8,956,471

0.37%

193,905

JPMORGAN CHASE & CO

$8,268,054

0.35%

179,001

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	34,976	$1,615,540	19.5%	Other
J.P. Morgan Investment Management Inc.	144,022	$6,652,376	80.5%	Defined
J.P. Morgan Securities LLC	3	$138	0.0%	Defined

TOCQUEVILLE ASSET MANAGEMENT L.P.

$8,032,441

0.34%

173,900

F&V Capital Management, LLC

$7,906,531

0.33%

188,610

Divisadero Street Capital Management, LP

$7,871,515

0.33%

170,416

Invesco Ltd.

$7,151,734

0.30%

154,833

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	8,257	$381,390	5.3%	Defined
Invesco Trust Co joint	20,936	$967,033	13.5%	Defined
Invesco Capital Management LLC	125,640	$5,803,311	81.1%	Defined

Trexquant Investment LP

$7,142,913

0.30%

154,642

Legal & General Group Plc

$7,116,030

0.30%

154,060

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	5,111	$236,077	3.3%	Defined
Legal & General Investment Management Ltd joint	148,949	$6,879,953	96.7%	Defined

VICTORY CAPITAL MANAGEMENT INC

$7,092,566

0.30%

153,552

Bank of New York Mellon Corp

$7,041,709

0.29%

152,451

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Investments Corporation	55,742	$2,574,722	36.6%	Defined
The Bank of New York Mellon	94,944	$4,385,463	62.3%	Defined
BNY Mellon, NA joint	1,763	$81,432	1.2%	Defined
Held directly	2	$92	0.0%	Defined

PERKINS CAPITAL MANAGEMENT INC

$6,828,821

0.29%

147,842

PRIMORIS WEALTH ADVISORS, LLC

$6,513,205

0.27%

141,009

DISCIPLINED GROWTH INVESTORS INC /MN

$5,905,807

0.25%

127,859

Russell Investments Group, Ltd.

$5,865,479

0.24%

126,986

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC joint	47,165	$2,178,550	37.1%	Defined
Russell Investments Canada Limited	969	$44,758	0.8%	Defined
Russell Investments Capital, LLC joint	36,932	$1,705,888	29.1%	Other
Russell Investments France SAS	10,570	$488,228	8.3%	Defined
Russell Investments Limited	1,493	$68,961	1.2%	Defined
Russell Investments Trust Company joint	29,857	$1,379,094	23.5%	Other

Verition Fund Management LLC

$5,849,547

0.24%

126,641

Susquehanna Portfolio Strategies, LLC

$5,828,438

0.24%

126,184

SG Americas Securities, LLC

$5,547,511

0.23%

120,102

FRANKLIN RESOURCES INC

$4,902,050

0.20%

106,128

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FIDUCIARY TRUST INTERNATIONAL LLC	23	$1,062	0.0%	Defined
FRANKLIN ADVISERS INC	100,351	$4,635,212	94.6%	Defined
FRANKLIN ADVISORY SERVICES LLC	3,360	$155,198	3.2%	Defined
PUTNAM ADVISORY CO LLC	2,394	$110,578	2.3%	Defined

Swiss National Bank

$4,568,191

0.19%

98,900

Calamos Advisors LLC

$4,394,793

0.18%

95,146

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC	22,018	$1,017,011	23.1%	Other
Held directly	73,128	$3,377,782	76.9%	Sole

PRINCIPAL FINANCIAL GROUP INC

$4,348,556

0.18%

94,145

Mariner, LLC

$4,249,295

0.18%

91,996

Lisanti Capital Growth, LLC

$4,183,197

0.17%

90,565

CITIGROUP INC

$4,170,864

0.17%

90,298

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	90,298	$4,170,864	100.0%	Defined

NEW YORK STATE COMMON RETIREMENT FUND

$4,104,304

0.17%

88,857

WELLS FARGO & COMPANY/MN

$3,979,821

0.17%

86,162

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	48,027	$2,218,367	55.7%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	11,920	$550,584	13.8%	Defined
WELLS FARGO CLEARING SERVICES, LLC	26,148	$1,207,776	30.3%	Defined
WELLS FARGO INVESTMENT INSTITUTE, INC.	67	$3,094	0.1%	Defined

XTX Topco Ltd

$3,748,780

0.16%

81,160

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	81,160	$3,748,780	100.0%	Defined

B. Metzler seel. Sohn & Co. AG

$3,745,916

0.16%

81,098

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Metzler Asset Management GmbH	81,098	$3,745,916	100.0%	Defined

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$3,673,212

0.15%

79,524

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	586	$27,067	0.7%	Defined
UBS Asset Management Switzerland AG	74,061	$3,420,877	93.1%	Defined
UBS Asset Management (UK) Ltd.	4,877	$225,268	6.1%	Sole

ROYAL BANK OF CANADA

$3,583,558

0.15%

77,583

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC CMA LTD.	1,601	$73,950	2.1%	Defined
RBC Capital Markets, LLC	75,981	$3,509,562	97.9%	Defined
Held directly	1	$46	0.0%	Sole

RICE HALL JAMES & ASSOCIATES, LLC

$3,413,579

0.14%

73,903

ROYCE & ASSOCIATES LP

$3,349,514

0.14%

72,516

KENNEDY CAPITAL MANAGEMENT LLC

$3,261,891

0.14%

70,619

SummitTX Capital, L.P.

$3,245,078

0.14%

70,255

Connor, Clark & Lunn Investment Management Ltd.

$3,100,180

0.13%

67,118

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 1	67,118	$3,100,180	100.0%	Sole

RHUMBLINE ADVISERS

$3,005,490

0.13%

65,068

ALLIANCEBERNSTEIN L.P.

$2,984,797

0.12%

64,620

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	64,620	$2,984,797	100.0%	Defined

Janney Montgomery Scott LLC

$2,848,814

0.12%

61,676

GLOBEFLEX CAPITAL L P

$2,701,976

0.11%

58,497

BARCLAYS PLC

$2,698,142

0.11%

58,414

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	54,790	$2,530,750	93.8%	Defined
BARCLAYS CAPITAL INC.	3,624	$167,392	6.2%	Defined

CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM

$2,661,005

0.11%

57,610

AMERIPRISE FINANCIAL INC

$2,655,970

0.11%

57,501

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	44,494	$2,055,177	77.4%	Defined
Ameriprise Financial Services, LLC	6	$277	0.0%	Defined
Columbia Threadneedle Management Ltd	13,001	$600,516	22.6%	Defined

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Key facts CIK 805928 CUSIP 05463X106 13F (30d) 22 filings 22 filers Visit website Investor relations