DNA · Ginkgo Bioworks Holdings, Inc. · Holdings

$9.80 -0.06 (-0.61%)

Russell 2000 member Rebalance outlook locked

Market Cap

$663.98M

67.34M

Volume · Oct 20 2.49M Avg daily vol (3M) 2M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Use institutional holdings through the 13F API or MCP → See every holder of DNA: 13F, 13D and 13G, insiders and funds →

Institutional holdings are being reconciled with SEC filings. Holder totals and changes may be incomplete until reconciliation finishes.

The Sep 30, 2026 filing window is still open — showing the latest known positions. Funds that have not filed yet carry their prior-quarter positions; a fund that filed without this stock counts as sold out. Each fund's Change is measured against its own previous reported quarter, shown in the Reported column. Pick a completed quarter in the Quarter selector below to put every fund on one quarter and see the stock-wide activity breakdown.

Quarter:

Holders: 183

Total Value: $370,735,110

Total Shares: 37,093,477

Showing 1–100 of 189 holdings by value · page 1 of 2

Institutional Ownership Trend

22 quarters

Superinvestors Holding DNA

All superinvestors →

Andreas Halvorsen $35,449,338

Holder Concentration

Across 186 13F filers

Top Holder

11.5%

of institutional value

Top 5

40.1%

of institutional value

Top 10

59.3%

of institutional value

HHI

468

Broadly held

Who Holds DNA

Last 8 quarters through Sep 30, 2026 (filing window open)

Each kind of 13F filer's share of the common shares 13F filers report. Holder type comes from regulatory records, never the filer's name; style, turnover and horizon are measured from each filer's own 13F book. The quarter selector does not change this card.

Jun 30, 2026, the newest closed quarter: 184 filers, like-for-like change +12.0%; actively managed books +17.6%; index-like books +5.4%.

Holder type	Share	Filers	Change	New	Exited
Registered adviser	47.8%	100	+14.3%	17	6
Exempt adviser	0.2%	1	new	1	0
Adviser	7.2%	3	-1.8%	1	1
Bank	9.8%	13	+21.5%	0	0
Insurer	0.2%	1	+53.3%	0	0
Broker-dealer	0.2%	2	-51.5%	0	1
Benefit plan	0.2%	1	+6.9%	0	1
Non-US	7.7%	6	+29.0%	1	0
Holding company	13.9%	5	+0.1%	0	0
Unknown	12.7%	52	+11.2%	8	10

Style	Share	Filers	Change	New	Exited
Index-like	36.2%	26	+5.4%	2	1
Systematic-like	4.7%	19	+14.0%	6	3
Concentrated	3.2%	32	+23.0%	6	5
Diversified active	51.1%	106	+17.6%	14	10
Unknown	4.8%	1	0.0%	0	0
Not measured	0.0%	0	—	0	0

Turnover	Share	Filers	Change	New	Exited
Low turnover	55.6%	114	+6.1%	16	9
Medium turnover	18.9%	31	+35.2%	2	2
High turnover	15.6%	29	+3.2%	9	5
Unknown	9.9%	10	+26.6%	1	3
Not measured	0.0%	0	—	0	0

Holding horizon	Share	Filers	Change	New	Exited
Short horizon	11.9%	22	+23.4%	5	4
Medium horizon	18.0%	30	+7.3%	6	4
Long horizon	58.0%	121	+14.0%	14	9
Unknown	12.1%	11	+0.9%	3	2
Not measured	0.0%	0	—	0	0

13F book size	Share	Filers	Change	New	Exited
Under $1B	6.0%	37	+7.9%	9	5
$1B to $10B	9.5%	44	-0.1%	4	9
$10B to $100B	24.4%	48	+14.8%	11	3
$100B or more	60.1%	55	+13.4%	4	2
Not measured	0.0%	0	—	0	0

Client base	Share	Filers	Change	New	Exited
Individuals and high net worth	4.3%	50	+4.1%	9	4
Pooled investment vehicles	24.1%	21	+6.4%	3	2
Registered funds and BDCs	15.4%	11	+23.7%	1	0
Pension and profit sharing plans	0.0%	1	new	1	0
Insurance companies	0.1%	2	+26.2%	0	0
Governments and sovereign funds	0.9%	4	+153.9%	2	0
Other investment advisers	0.0%	1	+24.3%	0	0
Diversified	2.5%	9	+12.3%	0	0
Charitable organizations	0.0%	0	—	0	0
Unknown	52.7%	85	+11.1%	12	13

Private funds	Share	Filers	Change	New	Exited
Hedge funds	24.4%	22	+10.7%	7	2
Private equity funds	2.1%	8	+4.1%	1	0
Venture capital funds	0.3%	2	+0.2%	0	0
Real estate funds	0.0%	3	+1363.8%	1	1
Securitized asset funds	0.2%	1	+1.1%	0	0
Other private funds	2.0%	8	+82.0%	1	0
Mixed private funds	0.3%	3	+40.5%	0	0
Private funds, type not stated	0.8%	4	+65.9%	1	0
No private funds	18.0%	50	+15.6%	7	3
Unknown	51.9%	83	+9.5%	10	13

Location	Share	Filers	Change	New	Exited
United States	73.8%	155	+12.3%	25	17
Outside the US	26.2%	29	+11.0%	3	2

The Sep 30, 2026 filing window is still open, so its bar carries the Jun 30, 2026 positions of filers that have not filed yet.

How the mix is measured

Shares are 13F common shares restated to today's split basis; options and principal amounts are left out. The change compares only filers with a 13F in both quarters and keeps each filer in its bucket of the later quarter on both sides; a filer that continues under a new CIK is compared through its successor. Holder type comes from each filer's current regulatory profile; client base and private funds come from the Form ADV roster current at each quarter end (rosters from December 2019, so an earlier quarter reads Unknown). Style, turnover, horizon and book size are measured from each filer's 13F book in the newest measured quarter, and Not measured means none was. Actively managed books are those measured systematic-like, concentrated or diversified active.

Excluded 1 holder with no observed 2026-03-31 13F and 2 holders with no observed 2026-06-30 13F. Compared across CIK changes: JANUS HENDERSON GROUP PLC continues as Jupiter Topco LLC.

Conviction and Crowding

183 13F filers, quarter ended Jun 30, 2026

Measured across every 13F filer's common-share book once the quarter's filing window closed. Crowding counts filers with high turnover or that advise private funds on Form ADV.

Conviction holders

1.64% of filers hold it in their top 10

Mar 31, 2026: 2

Holders above 5%

median weight 0.000%

Mar 31, 2026: 0

Crowding

—

the issuer has more than one class of common stock, so its count is the total across classes

Private-fund advisers

—

of shares outstanding

Institutional ownership

—

the issuer has more than one class of common stock, so its count is the total across classes

Top 10 holders

60.8%

of 13F shares, Herfindahl 0.049

Active holders

+17.6%

like-for-like share change of active filers

5% stakes on record

2 13D, 4 13G (1 institutional, 3 passive, 0 exempt)

How long holders have held it value-weighted 12.8 quarters

Under 1 year 17.6% 1 to 3 years 25.1% 3 years or more 57.4% Since coverage start 0.000%

Managers in the top fifth by current three-year FundScore alpha were net buyers of $0 (0 buying, 0 selling).

Estimated Holder Cost Basis

Quarter ended Jun 30, 2026, on that quarter's share basis

Estimated, never filed: each 13F buy is costed at its quarter's average traded price and each sale keeps the average cost, so this is what holders paid on average if they traded evenly through each quarter.

Estimated avg cost

$113.17

estimated per share

Mar 31, 2026: $126.00

Quarter-end price

$9.91

implied by 13F values

Estimated gain or loss

-91.2%

price against the estimated cost

Estimated at a loss

78.9%

of costed 13F value

Estimated cost covers 49.0% of the 13F shares; positions already held when a filer's 13F history starts, or bought in a quarter without a trusted average price, have no estimate.

Shared Holders

Companies DNA's active holders also own, quarter ended Jun 30, 2026

#	Company	Shared holders	Similarity
1	RXRX Recursion Pharmaceuticals, Inc.	3	0.146
2	AUR Aurora Innovation, Inc.	3	0.102
3	CPNG Coupang, Inc.	3	0.065
4	RDDT Reddit, Inc.	3	0.050
5	IBKR Interactive Brokers Group, Inc.	3	0.045
6	SPCX Space Exploration Technologies Corp	5	0.044
7	BA Boeing Co	5	0.033
8	ICE Intercontinental Exchange, Inc.	3	0.032
9	APH Amphenol Corp /De/	5	0.031
10	META Meta Platforms, Inc.	8	0.030

Shareholder Targets

13F filers that own its peers but not DNA, quarter ended Jun 30, 2026

Found through its holder-overlap peers: RXRX, AUR, CPNG, RDDT, IBKR, SPCX, BA, ICE, APH, META, SCHW, AMZN, MSFT, PWR, MMM, BX, NVDA, DIS, NOW, KO. 7,009 filers in all.

#	Filer	In peers	Of book	Largest peers
1	NORGES BANK Non-US, Index-like	$178,683,796,970 19 held	17.8%	NVDA, MSFT, AMZN
2	Capital Research Global Investors Adviser, Diversified active	$145,039,894,600 10 held	20.1%	NVDA, MSFT, AMZN
3	Capital World Investors Adviser, Diversified active	$97,106,241,962 15 held	11.5%	META, NVDA, MSFT
4	Alphabet Inc. Unknown, Concentrated	$94,176,237,970 1 held	95.1%	SPCX
5	Valor Management LLC Adviser, Concentrated	$86,013,406,595 1 held	99.8%	SPCX
6	Capital International Investors Adviser, Diversified active	$78,777,422,316 13 held	16.1%	MSFT, NVDA, APH
7	WELLINGTON MANAGEMENT GROUP LLP Holding company, Diversified active	$75,710,710,762 20 held	13.1%	NVDA, MSFT, AMZN
8	Amundi Holding company, Index-like	$73,595,337,092 19 held	17.7%	NVDA, MSFT, AMZN
9	AMERIPRISE FINANCIAL INC Adviser, Diversified active	$50,143,460,558 19 held	11.0%	NVDA, MSFT, AMZN
10	VY Capital Management Co. Ltd. Exempt adviser, Concentrated	$46,442,146,319 2 held	98.9%	SPCX, RDDT

Fund Ownership of DNA by Fund Type

Registered funds' Form N-PORT reports for the quarter ended Jun 30, 2026

Index funds and index ETFs hold 11.34% of shares outstanding.

Fund type	Of shares outstanding	Change	Shares	Funds
Index funds	2.43%	+0.47 pp	1,636,950	53
Index ETFs	8.90%	+0.84 pp	5,996,115	28
Active ETFs	0.06%	-0.12 pp	37,021	8
Active funds	1.97%	+0.55 pp	1,325,741	38
Unclassified	0.14%	+0.14 pp	96,542	5
All registered funds	13.50%	+1.88 pp	9,092,369	132

How fund types are measured

Each fund series counts once, with its latest Form N-PORT report dated inside the quarter; only long share positions count, restated to today's split basis. The fund type is the series' latest Form N-CEN answer on or before the quarter end (the index-fund box and the exchange-traded flag), never its name. Unclassified funds have no usable N-CEN; unit investment trusts are among them, so index ownership is a floor. Funds sit inside their advisers' 13F reports, so these shares overlap the 13F figures and are never added to them.

Holder type: Registered adviser 95 Exempt adviser 2 Adviser 3 Bank 13 Insurer 1 Broker-dealer 3 Benefit plan 1 Non-US 6 Holding company 5 Unknown 54

Showing 100 of 189

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

BlackRock, Inc.

$42,443,561

11.45%

4,282,903

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	53,168	$526,894	1.2%	Sole
BlackRock Investment Management (Australia) Ltd	62	$614	0.0%	Sole
BlackRock Asset Management Canada Ltd	3,108	$30,800	0.1%	Sole
BlackRock Investment Management, LLC	145,981	$1,446,671	3.4%	Sole
BLACKROCK ADVISORS LLC	106,388	$1,054,305	2.5%	Sole
BlackRock Fund Advisors	2,405,779	$23,841,269	56.2%	Sole
BlackRock Institutional Trust Company, N.A.	1,469,247	$14,560,237	34.3%	Sole
BlackRock Fund Managers Ltd.	5,603	$55,525	0.1%	Sole
BlackRock Investment Management (UK) Ltd.	6,785	$67,239	0.2%	Sole
BlackRock (Netherlands) B.V.	6,319	$62,621	0.1%	Sole
BlackRock Asset Management Ireland Ltd	36,697	$363,667	0.9%	Sole
BlackRock Life Ltd	190	$1,882	0.0%	Sole
BlackRock Asset Management Schweiz AG	6,531	$64,722	0.2%	Sole
Aperio Group, LLC	37,045	$367,115	0.9%	Sole

VIKING GLOBAL INVESTORS LP

$35,449,338

9.56%

3,577,128

Erste Asset Management GmbH

$25,141,065

6.78%

2,536,939

BAILLIE GIFFORD & CO

$23,701,241

6.39%

2,391,649

VANGUARD CAPITAL MANAGEMENT LLC

$20,685,073

5.58%

2,087,293

Anchorage Capital Group, L.L.C.

$17,306,457

4.67%

1,746,363

Legal & General Group Plc

$16,543,941

4.46%

1,669,419

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	4,618	$45,764	0.3%	Defined
Legal & General Investment Management Ltd joint	1,664,801	$16,498,177	99.7%	Defined

STATE STREET CORP

$14,591,186

3.94%

1,472,370

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	1,472,370	$14,591,186	100.0%	Defined

FORBES J M & CO LLP

$12,539,669

3.38%

977,371

GEODE CAPITAL MANAGEMENT, LLC

$12,110,177

3.27%

1,222,016

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	183,647	$1,819,941	15.0%	Defined
Held directly	1,038,369	$10,290,236	85.0%	Defined

VANGUARD PORTFOLIO MANAGEMENT LLC

$9,293,627

2.51%

937,803

Allianz Asset Management GmbH

$7,787,258

2.10%

785,798

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Pacific Investment Management Company LLC	785,798	$7,787,258	100.0%	Defined

UBS Group AG

$6,854,339

1.85%

691,659

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	4,491	$44,505	0.6%	Defined
UBS AG	521,868	$5,171,711	75.5%	Defined
UBS Switzerland AG	165,300	$1,638,123	23.9%	Defined

MILLENNIUM MANAGEMENT LLC

$6,329,279

1.71%

638,676

AQR CAPITAL MANAGEMENT LLC

$6,160,301

1.66%

621,625

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RF&L WEALTH MANAGEMENT, LLC	26	$257	0.0%	Other
Prism Planning Partners LLC	172	$1,704	0.0%	Other
Compound Planning, Inc.	41	$406	0.0%	Other
Arax Advisory Partners	6,152	$60,966	1.0%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	3,013	$29,858	0.5%	Other
Stokes Family Office, LLC	1,700	$16,847	0.3%	Other
Held directly	610,521	$6,050,263	98.2%	Defined

SB INVESTMENT ADVISERS (UK) LTD

$6,052,324

1.63%

610,729

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SB INVESTMENT ADVISERS (US) INC.	610,729	$6,052,324	100.0%	Sole

GOLDMAN SACHS GROUP INC

$6,019,095

1.62%

607,376

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	541,287	$5,364,154	89.1%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	66,089	$654,941	10.9%	Defined

BANK OF AMERICA CORP /DE/

$5,409,393

1.46%

545,852

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	480,399	$4,760,754	88.0%	Defined
BOFA SECURITIES, INC.	65,453	$648,639	12.0%	Defined

Prescott Group Capital Management, L.L.C.

$5,369,683

1.45%

541,845

RENAISSANCE TECHNOLOGIES LLC

$5,349,972

1.44%

539,856

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$4,109,677

1.11%

414,700

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	414,700	$4,109,677	100.0%	Other

NORTHERN TRUST CORP

$4,014,945

1.08%

405,141

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	388,962	$3,854,612	96.0%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	8,710	$86,316	2.1%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	7,469	$74,017	1.8%	Other

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$3,910,555

1.05%

394,607

EXCHANGE TRADED CONCEPTS, LLC

$3,555,539

0.96%

358,783

DIMENSIONAL FUND ADVISORS LP

$3,480,392

0.94%

351,200

TUDOR INVESTMENT CORP ET AL

$3,326,886

0.90%

335,710

VANGUARD FIDUCIARY TRUST CO

$3,097,598

0.84%

312,573

ACADIAN ASSET MANAGEMENT LLC

$2,866,695

0.77%

289,273

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Acadian Inc	289,273	$2,866,695	100.0%	Defined

Coastal Bridge Advisors, LLC

$2,743,256

0.74%

276,817

Invesco Ltd.

$2,098,172

0.57%

211,723

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	7,315	$72,490	3.5%	Defined
Invesco Trust Co joint	18,790	$186,208	8.9%	Defined
Invesco Capital Management LLC	185,618	$1,839,474	87.7%	Defined

BALANCE CAPITAL PARTNERS L.P.

$1,972,179

0.53%

199,009

ALGERT GLOBAL LLC

$1,784,394

0.48%

180,060

Susquehanna Portfolio Strategies, LLC

$1,746,449

0.47%

176,231

Bank of New York Mellon Corp

$1,689,465

0.46%

170,481

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Investments Corporation	82,628	$818,843	48.5%	Defined
The Bank of New York Mellon	87,608	$868,195	51.4%	Defined
BNY Mellon, NA joint	240	$2,378	0.1%	Defined
Held directly	5	$49	0.0%	Defined

BNP PARIBAS ARBITRAGE, SA

$1,637,934

0.44%

165,281

PEAK6 LLC

$1,619,294

0.44%

163,400

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PEAK6 Investments LLC joint	163,400	$1,619,294	100.0%	Defined

CAXTON ASSOCIATES LLP

$1,532,551

0.41%

154,647

D. E. Shaw & Co., Inc.

$1,499,937

0.40%

151,356

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	151,356	$1,499,937	100.0%	Defined

Nuveen, LLC

$1,455,233

0.39%

146,845

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	5,823	$57,705	4.0%	Defined
TEACHERS ADVISORS, LLC joint	141,022	$1,397,528	96.0%	Defined

MORGAN STANLEY

$1,430,149

0.39%

144,314

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	9	$89	0.0%	Defined
MORGAN STANLEY & CO. LLC	32,429	$321,371	22.5%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	45,149	$447,426	31.3%	Defined
MORGAN STANLEY SMITH BARNEY LLC	30,290	$300,173	21.0%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	36,437	$361,090	25.2%	Defined

Universal- Beteiligungs- und Servicegesellschaft mbH

$1,387,400

0.37%

140,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Universal-Investment-Gesellschaft mbH	140,000	$1,387,400	100.0%	Defined

JPMORGAN CHASE & CO

$1,328,335

0.36%

134,040

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	19,744	$195,663	14.7%	Defined
J.P. Morgan Investment Management Inc.	1,137	$11,267	0.8%	Defined
J.P. Morgan Securities LLC	113,159	$1,121,405	84.4%	Defined

CITADEL ADVISORS LLC

$1,297,565

0.35%

130,935

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	6,417	$63,592	4.9%	Defined
Held directly	124,518	$1,233,973	95.1%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$1,297,219

0.35%

130,900

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	130,900	$1,297,219	100.0%	Other

a16z Capital Management, L.L.C.

$1,075,908

0.29%

108,568

CITADEL ADVISORS LLC

$1,073,253

0.29%

108,300

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	108,300	$1,073,253	100.0%	Defined

Numerai GP LLC

$1,036,605

0.28%

104,602

AMERICAN CENTURY COMPANIES INC

$974,717

0.26%

98,357

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AMERICAN CENTURY INVESTMENT MANAGEMENT INC	98,357	$974,717	100.0%	Defined

Ensign Peak Advisors, Inc

$931,331

0.25%

93,979

PRICE T ROWE ASSOCIATES INC /MD/

$926,862

0.25%

93,528

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	93,528	$926,862	100.0%	Defined

PANAGORA ASSET MANAGEMENT INC

$803,591

0.22%

81,089

MARSHALL WACE, LLP

$772,355

0.21%

77,937

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marshall Wace North America L.P. joint	77,937	$772,355	100.0%	Other

Swiss National Bank

$744,865

0.20%

75,163

ALLSTATE CORP

$698,209

0.19%

70,455

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ALLSTATE INVESTMENTS LLC	70,455	$698,209	100.0%	Sole

Inspire Investing, LLC

$696,137

0.19%

70,246

Mariner, LLC

$667,527

0.18%

67,359

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$646,875

0.17%

65,275

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	900	$8,919	1.4%	Defined
UBS Asset Management Switzerland AG	64,375	$637,956	98.6%	Sole

DEUTSCHE BANK AG\

$645,368

0.17%

65,123

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Deutsche Bank Trust Co Americas	16,525	$163,762	25.4%	Defined
DWS Investments UK Ltd	48,598	$481,606	74.6%	Defined

IEQ CAPITAL, LLC

$624,766

0.17%

63,044

ALLEN OPERATIONS LLC

$613,934

0.17%

61,951

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Allen & CO LLC	61,951	$613,934	100.0%	Defined

RHUMBLINE ADVISERS

$610,624

0.16%

61,617

JACOBS LEVY EQUITY MANAGEMENT, INC

$609,078

0.16%

61,461

TWO SIGMA INVESTMENTS, LP

$599,099

0.16%

60,454

BROWN ADVISORY INC

$586,731

0.16%

59,206

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Brown Advisory LLC	59,206	$586,731	100.0%	Defined

ALLIANCEBERNSTEIN L.P.

$552,898

0.15%

55,792

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	55,792	$552,898	100.0%	Defined

CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM

$543,979

0.15%

54,892

Q Fund Management (Hong Kong) Ltd

$504,603

0.14%

39,330

EDGEWOOD MANAGEMENT LLC

$495,500

0.13%

50,000

MASSACHUSETTS FINANCIAL SERVICES CO /MA/

$466,660

0.13%

47,090

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MFS Heritage Trust CO	7,967	$78,952	16.9%	Defined
Held directly	39,123	$387,708	83.1%	Sole

Quadrature Capital Ltd

$456,732

0.12%

46,088

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Quadrature US Inc.	46,088	$456,732	100.0%	Other

CITIGROUP INC

$454,135

0.12%

45,826

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	45,826	$454,135	100.0%	Defined

Shay Capital LLC

$444,384

0.12%

44,842

LPL Financial LLC

$427,259

0.12%

43,114

WELLS FARGO & COMPANY/MN

$415,307

0.11%

41,908

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	39,887	$395,280	95.2%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	142	$1,407	0.3%	Defined
WELLS FARGO CLEARING SERVICES, LLC	1,879	$18,620	4.5%	Defined

JANE STREET GROUP, LLC

$414,238

0.11%

41,800

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	41,800	$414,238	100.0%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$390,175

0.11%

39,372

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	21,968	$217,702	55.8%	Other
G1 EXECUTION SERVICES, LLC	17,404	$172,473	44.2%	Other

Russell Investments Group, Ltd.

$373,595

0.10%

37,699

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC joint	32,684	$323,897	86.7%	Defined
Russell Investments Trust Company joint	5,015	$49,698	13.3%	Other

Casdin Capital, LLC

$361,645

0.10%

36,493

CITADEL ADVISORS LLC

$347,841

0.09%

35,100

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	35,100	$347,841	100.0%	Defined

Dynamic Technology Lab Private Ltd

$346,810

0.09%

34,996

GROUP ONE TRADING LLC

$326,138

0.09%

32,910

Mitsubishi UFJ Asset Management Co., Ltd.

$301,531

0.08%

30,427

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MITSUBISHI UFJ FINANCIAL GROUP INC	30,427	$301,531	100.0%	Defined

Welch & Forbes LLC

$297,300

0.08%

30,000

BARCLAYS PLC

$296,962

0.08%

29,966

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	26,127	$258,918	87.2%	Defined
BARCLAYS CAPITAL INC.	3,839	$38,044	12.8%	Defined

MetLife Investment Management, LLC

$285,110

0.08%

28,770

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
METLIFE INC	28,770	$285,110	100.0%	Defined

AXQ CAPITAL, LP

$262,387

0.07%

26,477

IMC-Chicago, LLC

$240,922

0.06%

24,311

Man Group plc

$239,901

0.06%

24,208

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Numeric Investors LLC	24,208	$239,901	100.0%	Defined

VANGUARD ASSET MANAGEMENT, Ltd

$238,424

0.06%

24,059

CANADA PENSION PLAN INVESTMENT BOARD

$222,975

0.06%

22,500

MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.

$217,356

0.06%

21,933

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Global X Management CO LLC joint	21,933	$217,356	100.0%	Defined

Modera Wealth Management, LLC

$216,315

0.06%

21,828

MARTINGALE ASSET MANAGEMENT L P

$209,061

0.06%

21,096

Boston Standard Wealth Management, LLC

$194,566

0.05%

15,165

Walleye Capital LLC

$188,944

0.05%

19,066

TriaGen Wealth Management LLC

$184,508

0.05%

14,381

FNY Investment Advisers, LLC

$181,929

0.05%

14,180

Trexquant Investment LP

$178,419

0.05%

18,004

SIMPLEX TRADING, LLC

$178,330

0.05%

17,995

Comprehensive Financial Management LLC

$174,970

0.05%

17,656

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Key facts CIK 1830214 CUSIP 37611X209 13F (30d) 16 filings 16 filers Visit website Investor relations