GRAL · GRAIL, Inc. · Holdings

$128.22 -4.38 (-3.30%)

Russell 2000 member Rebalance outlook locked

Market Cap

$5.92B

44.67M

Volume · Oct 22 99.63K Avg daily vol (3M) 992.95K

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Use institutional holdings through the 13F API or MCP → See every holder of GRAL: 13F, 13D and 13G, insiders and funds →

Institutional holdings are being reconciled with SEC filings. Holder totals and changes may be incomplete until reconciliation finishes.

The Sep 30, 2026 filing window is still open — showing the latest known positions. Funds that have not filed yet carry their prior-quarter positions; a fund that filed without this stock counts as sold out. Each fund's Change is measured against its own previous reported quarter, shown in the Reported column. Pick a completed quarter in the Quarter selector below to put every fund on one quarter and see the stock-wide activity breakdown.

Quarter:

Holders: 392

Total Value: $2,757,329,916

Total Shares: 39,961,304

Showing 1–100 of 406 holdings by value · page 1 of 5

Institutional Ownership Trend

11 quarters

Superinvestors Holding GRAL

All superinvestors →

Mario Gabelli $4,554,496 Joel Greenblatt $1,634,383

Holder Concentration

Across 380 13F filers

Top Holder

10.4%

of institutional value

Top 5

40.9%

of institutional value

Top 10

61.8%

of institutional value

HHI

485

Broadly held

Who Holds GRAL

Last 8 quarters through Sep 30, 2026 (filing window open)

Each kind of 13F filer's share of the common shares 13F filers report. Holder type comes from regulatory records, never the filer's name; style, turnover and horizon are measured from each filer's own 13F book. The quarter selector does not change this card.

Jun 30, 2026, the newest closed quarter: 375 filers, like-for-like change +8.4%; actively managed books +6.1%; index-like books +13.3%.

Holder type	Share	Filers	Change	New	Exited
Registered adviser	55.5%	198	+7.8%	29	29
Exempt adviser	0.1%	3	-55.7%	1	0
Adviser	6.5%	9	+22.4%	2	0
Bank	12.5%	21	+15.8%	1	1
Insurer	0.2%	2	+7.2%	0	0
Broker-dealer	0.1%	2	-37.6%	1	0
Benefit plan	0.3%	7	+5.6%	2	0
Non-US	0.8%	11	-35.0%	1	1
Holding company	18.7%	11	+12.0%	2	0
Unknown	5.5%	111	-10.3%	19	17

Style	Share	Filers	Change	New	Exited
Index-like	32.7%	38	+13.3%	5	2
Systematic-like	0.9%	14	-69.8%	8	10
Concentrated	20.9%	73	+2.0%	10	8
Diversified active	45.5%	250	+13.9%	35	28
Not measured	0.0%	0	—	0	0
Unknown	0.0%	0	—	0	0

Turnover	Share	Filers	Change	New	Exited
Low turnover	60.4%	240	+8.5%	28	23
Medium turnover	28.2%	70	+14.5%	10	12
High turnover	4.6%	28	-26.2%	13	10
Unknown	6.8%	37	+17.6%	7	3
Not measured	0.0%	0	—	0	0

Holding horizon	Share	Filers	Change	New	Exited
Short horizon	7.2%	44	+18.7%	12	8
Medium horizon	19.8%	63	+18.3%	13	11
Long horizon	63.1%	253	+6.8%	28	26
Unknown	9.8%	15	-4.8%	5	3
Not measured	0.0%	0	—	0	0

13F book size	Share	Filers	Change	New	Exited
Under $1B	6.0%	114	+20.3%	20	17
$1B to $10B	14.2%	93	+12.1%	19	17
$10B to $100B	28.1%	100	-1.3%	14	9
$100B or more	51.7%	68	+12.0%	5	5
Not measured	0.0%	0	—	0	0

Client base	Share	Filers	Change	New	Exited
Individuals and high net worth	2.3%	129	-12.3%	15	14
Pooled investment vehicles	30.3%	20	+4.8%	7	10
Registered funds and BDCs	17.1%	19	+15.9%	4	4
Pension and profit sharing plans	0.0%	3	+20.1%	0	0
Insurance companies	0.1%	3	+25.1%	0	0
Governments and sovereign funds	0.2%	2	-4.9%	0	0
Other investment advisers	0.7%	6	-1.5%	2	0
Diversified	4.8%	16	+14.3%	1	0
Charitable organizations	0.0%	0	-100.0%	0	1
Corporations	0.0%	0	—	0	0
Other	0.0%	0	—	0	0
Unknown	44.5%	177	+9.1%	29	19

Private funds	Share	Filers	Change	New	Exited
Hedge funds	33.0%	34	+4.7%	11	8
Private equity funds	0.3%	9	-5.5%	2	0
Venture capital funds	0.0%	2	-99.9%	0	1
Real estate funds	0.0%	5	-22.9%	1	1
Securitized asset funds	0.2%	2	+22.4%	1	0
Other private funds	2.5%	12	+48.5%	2	1
Mixed private funds	1.4%	4	+96.9%	0	0
Private funds, type not stated	0.1%	3	-48.4%	1	4
No private funds	18.1%	130	+13.2%	12	13
Liquidity funds	0.0%	0	-100.0%	0	1
Unknown	44.4%	174	+9.4%	28	19

Location	Share	Filers	Change	New	Exited
United States	96.1%	322	+9.5%	51	39
Outside the US	3.9%	53	-14.0%	7	9

The Sep 30, 2026 filing window is still open, so its bar carries the Jun 30, 2026 positions of filers that have not filed yet.

How the mix is measured

Shares are 13F common shares restated to today's split basis; options and principal amounts are left out. The change compares only filers with a 13F in both quarters and keeps each filer in its bucket of the later quarter on both sides; a filer that continues under a new CIK is compared through its successor. Holder type comes from each filer's current regulatory profile; client base and private funds come from the Form ADV roster current at each quarter end (rosters from December 2019, so an earlier quarter reads Unknown). Style, turnover, horizon and book size are measured from each filer's 13F book in the newest measured quarter, and Not measured means none was. Actively managed books are those measured systematic-like, concentrated or diversified active.

Excluded 4 holders with no observed 2026-03-31 13F and 5 holders with no observed 2026-06-30 13F. Compared across CIK changes: 4 filers continue as Evelyn Partners Group Ltd; JANUS HENDERSON GROUP PLC continues as Jupiter Topco LLC.

Conviction and Crowding

375 13F filers, quarter ended Jun 30, 2026

Measured across every 13F filer's common-share book once the quarter's filing window closed. Crowding counts filers with high turnover or that advise private funds on Form ADV.

Conviction holders

0.800% of filers hold it in their top 10

Mar 31, 2026: 1

Holders above 5%

median weight 0.001%

Mar 31, 2026: 1

Crowding

36.0%

of shares outstanding, 88th percentile

Mar 31, 2026: 36.7%

Private-fund advisers

33.3%

of shares outstanding

Institutional ownership

88.8%

110.1% of the estimated free float above 100%: filers can report lent or client shares more than once

Top 10 holders

63.9%

of 13F shares, Herfindahl 0.052

Active holders

+6.1%

like-for-like share change of active filers

5% stakes on record

0 13D, 7 13G (5 institutional, 2 passive, 0 exempt)

How long holders have held it value-weighted 6.7 quarters

Under 1 year 19.5% 1 to 3 years 80.5% 3 years or more 0.000% Since coverage start 0.000%

Managers in the top fifth by current three-year FundScore alpha were net buyers of $0 (0 buying, 0 selling).

Estimated Holder Cost Basis

Quarter ended Jun 30, 2026, on that quarter's share basis

Estimated, never filed: each 13F buy is costed at its quarter's average traded price and each sale keeps the average cost, so this is what holders paid on average if they traded evenly through each quarter.

Estimated avg cost

$61.69

estimated per share

Mar 31, 2026: $63.30

Quarter-end price

$68.27

implied by 13F values

Estimated gain or loss

+10.7%

price against the estimated cost

Estimated at a loss

20.4%

of costed 13F value

Estimated cost covers 41.1% of the 13F shares; positions already held when a filer's 13F history starts, or bought in a quarter without a trusted average price, have no estimate.

Shared Holders

Companies GRAL's active holders also own, quarter ended Jun 30, 2026

#	Company	Shared holders	Similarity
1	WGS GeneDx Holdings Corp.	5	0.148
2	AD Array Digital Infrastructure, Inc.	3	0.147
3	QS QuantumScape Corp	5	0.137
4	PSNL Personalis, Inc.	4	0.135
5	ILMN Illumina, Inc.	11	0.135
6	PRME Prime Medicine, Inc.	3	0.130
7	IMVT Immunovant, Inc.	4	0.127
8	PACB Pacific Biosciences Of California, Inc.	3	0.122
9	BEAM Beam Therapeutics Inc.	4	0.113
10	QDEL QuidelOrtho Corp	3	0.112

Shareholder Targets

13F filers that own its peers but not GRAL, quarter ended Jun 30, 2026

Found through its holder-overlap peers: WGS, AD, QS, PSNL, ILMN, PRME, IMVT, PACB, BEAM, QDEL, TSHA, DRVN, GH, CRSP, PBLS, RNA, TNGX, MAZE, NTLA, ATEX. 1,387 filers in all.

#	Filer	In peers	Of book	Largest peers
1	Capital World Investors Adviser, Diversified active	$2,971,269,240 2 held	0.350%	ILMN, CRSP
2	ARK Investment Management LLC Registered adviser, Diversified active	$1,714,733,415 9 held	11.1%	CRSP, BEAM, NTLA
3	WCM INVESTMENT MANAGEMENT, LLC Registered adviser, Diversified active	$1,116,380,746 2 held	2.01%	ILMN, GH
4	RA CAPITAL MANAGEMENT, L.P. Registered adviser, Concentrated	$1,028,915,913 3 held	9.06%	PBLS, TSHA, NTLA
5	RTW INVESTMENTS, LP Registered adviser, Diversified active	$897,422,640 6 held	7.66%	TNGX, GH, IMVT
6	Corvex Management LP Registered adviser, Concentrated	$826,156,688 2 held	25.7%	ILMN, WGS
7	BAILLIE GIFFORD & CO Adviser, Diversified active	$588,358,043 4 held	0.530%	GH, ILMN, BEAM
8	Capital International Investors Adviser, Diversified active	$579,221,505 2 held	0.120%	GH, CRSP
9	Sumitomo Mitsui Trust Group, Inc. Bank, Index-like	$559,592,979 9 held	0.320%	ILMN, BEAM, CRSP
10	Owl Creek Asset Management, L.P. Registered adviser, Concentrated	$557,088,221 1 held	49.4%	ATEX

Fund Ownership of GRAL by Fund Type

Registered funds' Form N-PORT reports for the quarter ended Jun 30, 2026

Index funds and index ETFs hold 15.34% of shares outstanding.

Fund type	Of shares outstanding	Change	Shares	Funds
Index funds	2.78%	+0.43 pp	1,242,662	62
Index ETFs	12.56%	+1.62 pp	5,610,873	39
Active ETFs	0.81%	+0.21 pp	362,965	20
Active funds	7.17%	+3.81 pp	3,202,149	97
Unclassified	0.23%	+0.22 pp	101,461	10
All registered funds	23.55%	+6.29 pp	10,520,110	228

How fund types are measured

Each fund series counts once, with its latest Form N-PORT report dated inside the quarter; only long share positions count, restated to today's split basis. The fund type is the series' latest Form N-CEN answer on or before the quarter end (the index-fund box and the exchange-traded flag), never its name. Unclassified funds have no usable N-CEN; unit investment trusts are among them, so index ownership is a floor. Funds sit inside their advisers' 13F reports, so these shares overlap the 13F figures and are never added to them.

Holder type: Registered adviser 208 Exempt adviser 3 Adviser 9 Bank 21 Insurer 2 Broker-dealer 2 Benefit plan 7 Non-US 11 Holding company 11 Unknown 118

Showing 100 of 406

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

FARALLON CAPITAL MANAGEMENT LLC

$284,111,339

10.30%

4,161,584

BAKER BROS. ADVISORS LP

$275,033,477

9.97%

4,028,614

BlackRock, Inc.

$244,206,904

8.86%

3,577,075

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	45,875	$3,131,886	1.3%	Sole
BlackRock Investment Management (Australia) Ltd	78	$5,325	0.0%	Sole
BlackRock Asset Management Canada Ltd	7,637	$521,377	0.2%	Sole
BlackRock Investment Management, LLC	114,819	$7,838,693	3.2%	Sole
BLACKROCK ADVISORS LLC	127,139	$8,679,779	3.6%	Sole
BlackRock Fund Advisors	1,889,073	$128,967,013	52.8%	Sole
BlackRock Institutional Trust Company, N.A.	1,151,308	$78,599,797	32.2%	Sole
BlackRock Japan Co. Ltd	479	$32,701	0.0%	Sole
BlackRock Fund Managers Ltd.	7,219	$492,841	0.2%	Sole
BlackRock Investment Management (UK) Ltd.	7,383	$504,037	0.2%	Sole
BlackRock (Netherlands) B.V.	846	$57,756	0.0%	Sole
BlackRock Asset Management Ireland Ltd	129,340	$8,830,041	3.6%	Sole
BlackRock Asset Management Schweiz AG	6,612	$451,401	0.2%	Sole
Aperio Group, LLC	89,265	$6,094,121	2.5%	Sole
SpiderRock Advisors, LLC	2	$136	0.0%	Sole

STATE STREET CORP

$162,859,040

5.91%

2,385,514

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	2,385,514	$162,859,040	100.0%	Defined

PRIMECAP MANAGEMENT CO/CA/

$148,835,836

5.40%

2,180,106

Sessa Capital IM, L.P.

$139,101,353

5.04%

2,037,518

Deep Track Capital, LP

$125,271,626

4.54%

1,834,944

VANGUARD CAPITAL MANAGEMENT LLC

$112,039,057

4.06%

1,641,117

MORGAN STANLEY

$99,821,863

3.62%

1,462,163

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	1,159	$79,124	0.1%	Defined
MORGAN STANLEY & CO. LLC	294,852	$20,129,546	20.2%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	286,765	$19,577,446	19.6%	Defined
MORGAN STANLEY AIP GP LP	38	$2,593	0.0%	Defined
MORGAN STANLEY INVESTMENT MANAGEMENT INC	614,576	$41,957,103	42.0%	Defined
MORGAN STANLEY SMITH BARNEY LLC	194,090	$13,250,524	13.3%	Defined
Morgan Stanley Bank, N.A.	2,152	$146,917	0.1%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	55,668	$3,800,454	3.8%	Defined
EATON VANCE MANAGEMENT	8,295	$566,299	0.6%	Defined
BOSTON MANAGEMENT & RESEARCH	4,568	$311,857	0.3%	Defined

Arbiter Partners Capital Management LLC

$70,611,456

2.56%

1,034,297

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Broken Clock Management LLC joint	1,034,297	$70,611,456	100.0%	Defined

GEODE CAPITAL MANAGEMENT, LLC

$67,681,512

2.45%

991,380

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	135,198	$9,229,967	13.6%	Defined
Held directly	856,182	$58,451,545	86.4%	Defined

GOLDMAN SACHS GROUP INC

$51,066,983

1.85%

748,015

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	623,687	$42,579,111	83.4%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	124,328	$8,487,872	16.6%	Defined

Jefferies Financial Group Inc.

$49,427,480

1.79%

724,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JEFFERIES LLC joint	724,000	$49,427,480	100.0%	Defined

BANK OF AMERICA CORP /DE/

$49,389,043

1.79%

723,437

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	617,434	$42,152,219	85.3%	Defined
MERRILL LYNCH, PIERCE, FENNER & SMITH INC.	4,087	$279,019	0.6%	Defined
BOFA SECURITIES, INC.	101,916	$6,957,805	14.1%	Defined

DIMENSIONAL FUND ADVISORS LP

$46,395,402

1.68%

679,587

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	4,599	$313,973	0.7%	Defined
DFA Australia Ltd.	7,016	$478,982	1.0%	Defined
Dimensional Fund Advisors Pte. Ltd.	232	$15,838	0.0%	Defined
Dimensional Ireland Ltd	27,521	$1,878,858	4.0%	Defined
Held directly	640,219	$43,707,751	94.2%	Sole

Jefferies Financial Group Inc.

$44,375,500

1.61%

650,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JEFFERIES LLC	650,000	$44,375,500	100.0%	Defined

Casdin Capital, LLC

$32,769,600

1.19%

480,000

VANGUARD PORTFOLIO MANAGEMENT LLC

$32,631,831

1.18%

477,982

NORTHERN TRUST CORP

$25,436,580

0.92%

372,588

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	344,757	$23,536,559	92.5%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	21,424	$1,462,616	5.8%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	6,407	$437,405	1.7%	Other

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$25,396,440

0.92%

372,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	372,000	$25,396,440	100.0%	Other

CITIGROUP INC

$25,345,441

0.92%

371,253

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citibank, N.A. joint	2	$136	0.0%	Defined
Citigroup Financial Products Inc. joint	371,251	$25,345,305	100.0%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$24,563,546

0.89%

359,800

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	359,800	$24,563,546	100.0%	Other

KINGDON CAPITAL MANAGEMENT, L.L.C.

$23,211,800

0.84%

340,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 1 joint	340,000	$23,211,800	100.0%	Defined

Douglas Lane & Associates, LLC

$22,640,589

0.82%

161,580

Granahan Investment Management, LLC

$21,926,685

0.80%

321,176

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$20,523,873

0.74%

300,628

RAYMOND JAMES FINANCIAL INC

$19,991,435

0.73%

292,829

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	153,731	$10,495,215	52.5%	Defined
Raymond James Financial Services Advisors, Inc.	139,098	$9,496,220	47.5%	Defined

T. Rowe Price Investment Management, Inc.

$17,093,442

0.62%

250,380

UBS Group AG

$17,053,571

0.62%

249,796

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	26,882	$1,835,234	10.8%	Defined
UBS AG	204,939	$13,991,185	82.0%	Defined
UBS Switzerland AG	17,866	$1,219,711	7.2%	Defined
UBS Securities LLC	109	$7,441	0.0%	Defined

Aberdeen Group plc

$15,668,169

0.57%

229,503

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
abrdn Inc.	229,503	$15,668,169	100.0%	Defined

VANGUARD FIDUCIARY TRUST CO

$15,357,336

0.56%

224,950

Slow Capital, Inc.

$14,832,476

0.54%

217,262

CITADEL ADVISORS LLC

$14,616,607

0.53%

214,100

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	214,100	$14,616,607	100.0%	Defined

AQR CAPITAL MANAGEMENT LLC

$14,428,247

0.52%

211,341

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Petros Family Wealth, LLC	22	$1,501	0.0%	Other
RF&L WEALTH MANAGEMENT, LLC	8	$546	0.0%	Other
Arax Advisory Partners	1,717	$117,219	0.8%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	1,776	$121,247	0.8%	Other
Stokes Family Office, LLC	671	$45,809	0.3%	Other
Held directly	207,147	$14,141,925	98.0%	Defined

WELLS FARGO & COMPANY/MN

$13,259,875

0.48%

194,227

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	168,719	$11,518,446	86.9%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	2,411	$164,598	1.2%	Defined
WELLS FARGO CLEARING SERVICES, LLC	22,790	$1,555,873	11.7%	Defined
Wells Fargo Securities, LLC	204	$13,927	0.1%	Defined
WELLS FARGO INVESTMENT INSTITUTE, INC.	103	$7,031	0.1%	Defined

TCW GROUP INC

$13,101,900

0.48%

191,913

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Sepulveda Management LLC	191,913	$13,101,900	100.0%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$10,601,989

0.38%

155,295

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	155,295	$10,601,989	100.0%	Other

Bank of New York Mellon Corp

$10,522,245

0.38%

154,127

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNY Mellon Trust of Delaware joint	56	$3,822	0.0%	Defined
Mellon Investments Corporation	51,106	$3,489,006	33.2%	Defined
BNY Mellon Securities Corporation	364	$24,850	0.2%	Defined
The Bank of New York Mellon	73,875	$5,043,446	47.9%	Defined
BNY Mellon, NA joint	14,887	$1,016,334	9.7%	Defined
Newton Investment Management North America, LLC	13,836	$944,583	9.0%	Defined
Held directly	3	$204	0.0%	Defined

FRANKLIN RESOURCES INC

$10,369,322

0.38%

151,887

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FIDUCIARY TRUST CO INTERNATIONAL	22	$1,501	0.0%	Defined
Fiduciary Trust Co International of Pennsylvania	1,299	$88,682	0.9%	Defined
Fiduciary Trust International of California	5	$341	0.0%	Defined
FIDUCIARY TRUST INTERNATIONAL LLC	185	$12,629	0.1%	Defined
FRANKLIN ADVISERS INC	150,376	$10,266,169	99.0%	Defined

Newtyn Management, LLC

$10,240,500

0.37%

150,000

180 WEALTH ADVISORS, LLC

$8,723,130

0.32%

127,774

FMR LLC

$8,651,583

0.31%

126,726

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	119,226	$8,139,559	94.1%	Defined
FIDELITY MANAGEMENT TRUST CO	1,201	$81,992	0.9%	Defined
STRATEGIC ADVISERS LLC	6,299	$430,032	5.0%	Defined

Voleon Capital Management LP

$8,264,902

0.30%

121,062

D. E. Shaw & Co., Inc.

$7,926,282

0.29%

116,102

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	65,106	$4,444,786	56.1%	Defined
D. E. Shaw Investment Management, L.L.C. joint	50,996	$3,481,496	43.9%	Defined

JPMORGAN CHASE & CO

$7,837,802

0.28%

114,806

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	20,559	$1,403,562	17.9%	Defined
J.P. Morgan Investment Management Inc.	66,387	$4,532,240	57.8%	Defined
J.P. Morgan Securities LLC	24,117	$1,646,467	21.0%	Defined
J.P. Morgan Securities plc	3,710	$253,281	3.2%	Defined
55I, LLC	33	$2,252	0.0%	Other

Invesco Ltd.

$7,740,039

0.28%

113,374

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	5,511	$376,234	4.9%	Defined
Invesco Trust Co joint	16,374	$1,117,852	14.4%	Defined
Invesco Taiwan Ltd	88	$6,007	0.1%	Defined
Invesco Capital Management LLC	91,401	$6,239,946	80.6%	Defined

WIM INVESTMENT MANAGEMENT Ltd

$7,263,928

0.26%

106,400

Alyeska Investment Group, L.P.

$7,168,350

0.26%

105,000

Legal & General Group Plc

$6,870,554

0.25%

100,638

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	4,011	$273,830	4.0%	Defined
Legal & General Investment Management Ltd joint	96,627	$6,596,724	96.0%	Defined

Legal Advantage Investments, Inc.

$6,813,335

0.25%

48,625

AMERICAN CENTURY COMPANIES INC

$6,774,227

0.25%

99,227

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AMERICAN CENTURY INVESTMENT MANAGEMENT INC	99,227	$6,774,227	100.0%	Defined

Nuveen, LLC

$6,264,523

0.23%

91,761

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	4,753	$324,487	5.2%	Defined
TEACHERS ADVISORS, LLC joint	87,008	$5,940,036	94.8%	Defined

NOMURA ASSET MANAGEMENT INTERNATIONAL INC.

$5,809,026

0.21%

85,089

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NOMURA HOLDINGS INC joint	85,089	$5,809,026	100.0%	Defined

Swiss National Bank

$5,202,174

0.19%

76,200

Reynders McVeigh Capital Management, LLC

$4,867,924

0.18%

71,304

Susquehanna Portfolio Strategies, LLC

$4,853,177

0.18%

71,088

NATIXIS

$4,794,329

0.17%

70,226

BARCLAYS PLC

$4,778,148

0.17%

69,989

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	66,751	$4,557,090	95.4%	Defined
BARCLAYS CAPITAL INC.	3,238	$221,058	4.6%	Defined

CITADEL ADVISORS LLC

$4,601,398

0.17%

67,400

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	67,400	$4,601,398	100.0%	Defined

GAMCO INVESTORS, INC. ET AL

$4,554,496

0.17%

66,713

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$4,511,072

0.16%

66,077

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	585	$39,937	0.9%	Defined
UBS Asset Management Switzerland AG	58,354	$3,983,827	88.3%	Defined
UBS Asset Management (UK) Ltd.	3,646	$248,911	5.5%	Defined
Held directly	3,492	$238,397	5.3%	Defined

ROYAL BANK OF CANADA

$4,081,655

0.15%

59,787

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC CMA LTD.	1,257	$85,815	2.1%	Defined
RBC Capital Markets, LLC	50,196	$3,426,880	84.0%	Defined
RBC Dominion Securities Inc.	364	$24,850	0.6%	Defined
RBC Trust Co (Delaware) Ltd	6,046	$412,760	10.1%	Defined
RBC Rochdale, LLC	5	$341	0.0%	Defined
RBC Europe Ltd	60	$4,096	0.1%	Defined
Held directly	1,859	$126,913	3.1%	Sole

Rockefeller Capital Management L.P.

$3,944,571

0.14%

57,779

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ROCKEFELLER & CO. LLC	649	$44,307	1.1%	Defined
ROCKEFELLER TRUST COMPANY, N.A.	190	$12,971	0.3%	Defined
Rockefeller Trust Co (Delaware)	853	$58,234	1.5%	Defined
ROCKEFELLER FINANCIAL LLC	56,087	$3,829,059	97.1%	Defined

PRICE T ROWE ASSOCIATES INC /MD/

$3,926,208

0.14%

57,521

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	57,510	$3,926,207	100.0%	Defined
Held directly	11	$1	0.0%	Sole

JACOBS LEVY EQUITY MANAGEMENT, INC

$3,810,967

0.14%

55,822

Allianz Asset Management GmbH

$3,810,694

0.14%

55,818

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Pacific Investment Management Company LLC	55,818	$3,810,694	100.0%	Defined

JANE STREET GROUP, LLC

$3,775,331

0.14%

55,300

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	55,300	$3,775,331	100.0%	Defined

RHUMBLINE ADVISERS

$3,773,214

0.14%

55,269

WEALTH ENHANCEMENT ADVISORY SERVICES, LLC

$3,636,114

0.13%

25,950

Public Sector Pension Investment Board

$3,633,534

0.13%

53,223

PVG ASSET MANAGEMENT CORP

$3,595,339

0.13%

25,659

Pictet Asset Management Holding SA

$3,416,162

0.12%

50,039

Centiva Capital, LP

$3,413,500

0.12%

50,000

Verition Fund Management LLC

$3,334,921

0.12%

48,849

ALLIANCEBERNSTEIN L.P.

$3,292,389

0.12%

48,226

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	48,226	$3,292,389	100.0%	Defined

Creative Planning

$3,254,020

0.12%

47,664

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CREATIVE PLANNING TRUST Co LLC	1,576	$107,593	3.3%	Defined
Held directly	46,088	$3,146,427	96.7%	Sole

D. E. Shaw & Co., Inc.

$3,072,150

0.11%

45,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	45,000	$3,072,150	100.0%	Defined

CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM

$2,966,058

0.11%

43,446

PRINCIPAL FINANCIAL GROUP INC

$2,903,864

0.11%

42,535

DEUTSCHE BANK AG\

$2,842,690

0.10%

41,639

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Deutsche Bank Trust Co Americas	77	$5,256	0.2%	Other
DWS INVESTMENT MANAGEMENT AMERICAS, INC	766	$52,294	1.8%	Defined
DEUTSCHE BANK NATIONAL TRUST Co	21	$1,433	0.1%	Defined
Deutsche Bank Trust Co Delaware	31	$2,116	0.1%	Defined
DEUTSCHE BANK TRUST COMPANY, NATIONAL ASSOCIATION	59	$4,027	0.1%	Other
DWS Investments UK Ltd	40,685	$2,777,564	97.7%	Defined

HighTower Advisors, LLC

$2,769,372

0.10%

40,565

Rafferty Asset Management, LLC

$2,697,962

0.10%

39,519

Timelo Investment Management Inc.

$2,662,530

0.10%

39,000

NORDEA INVESTMENT MANAGEMENT AB

$2,395,320

0.09%

35,086

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nordea Funds Ltd	6,650	$453,995	19.0%	Defined
Nordea Investment Funds S.A.	28,436	$1,941,325	81.0%	Defined

Lido Advisors, LLC

$2,270,250

0.08%

33,254

Bouvel Investment Partners, LLC

$2,269,803

0.08%

16,199

STIFEL FINANCIAL CORP

$2,151,322

0.08%

31,512

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STIFEL NICOLAUS & CO INC \MO\	31,424	$2,145,316	99.7%	Defined
1919 Investment Counsel, LLC	74	$5,051	0.2%	Defined
Held directly	14	$955	0.0%	Defined

BNP PARIBAS ARBITRAGE, SA

$2,133,983

0.08%

31,258

Summit Financial, LLC

$2,103,876

0.08%

30,817

GABELLI FUNDS LLC

$2,067,420

0.07%

30,283

JONES FINANCIAL COMPANIES LLLP

$2,059,637

0.07%

30,169

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Edward D. Jones & Co., L.P.	30,169	$2,059,637	100.0%	Sole

Russell Investments Group, Ltd.

$1,939,070

0.07%

28,403

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC joint	22,881	$1,562,085	80.6%	Defined
Russell Investments Capital, LLC joint	2,060	$140,635	7.3%	Other
Russell Investments France SAS	400	$27,308	1.4%	Defined
Russell Investments Trust Company joint	3,062	$209,042	10.8%	Other

OSAIC HOLDINGS, INC.

$1,916,884

0.07%

28,078

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
OSAIC WEALTH, INC.	27,934	$1,907,054	99.5%	Defined
OSAIC INSTITUTIONS, INC.	72	$4,915	0.3%	Defined
Osaic Advisory Services, LLC	72	$4,915	0.3%	Defined

WILLIAM JOHN SEMPOLINSKI

$1,895,123

0.07%

13,525

MOODY NATIONAL BANK TRUST DIVISION

$1,888,537

0.07%

13,478

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MOODY BANCSHARES INC joint	13,478	$1,888,537	100.0%	Defined

TWO SIGMA INVESTMENTS, LP

$1,872,304

0.07%

27,425

COMMERCE BANK

$1,841,992

0.07%

26,981

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
COMMERCE BANCSHARES INC /MO/	26,981	$1,841,992	100.0%	Sole

Gem Investment Advisors, LLC

$1,828,338

0.07%

26,781

VANGUARD ASSET MANAGEMENT, Ltd

$1,823,764

0.07%

26,714

Legato Capital Management LLC

$1,804,649

0.07%

26,434

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Key facts CIK 1699031 CUSIP 384747101 13F (30d) 65 filings 65 filers Visit website Investor relations