ALEC · Alector, Inc. · Holdings

$2.18 +0.08 (+3.81%) Q2 2025 earnings call transcript

Russell 2000 member Rebalance outlook locked

Market Cap

$234.48M

111.66M

Volume · Oct 28 1.38M Avg daily vol (3M) 3.24M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Use institutional holdings through the 13F API or MCP → See every holder of ALEC: 13F, 13D and 13G, insiders and funds →

The Sep 30, 2026 filing window is still open — showing the latest known positions. Funds that have not filed yet carry their prior-quarter positions; a fund that filed without this stock counts as sold out. Each fund's Change is measured against its own previous reported quarter, shown in the Reported column. Pick a completed quarter in the Quarter selector below to put every fund on one quarter and see the stock-wide activity breakdown.

Quarter:

Holders: 139

Total Value: $185,008,496

Total Shares: 92,044,768

Showing 1–100 of 143 holdings by value · page 1 of 2

Institutional Ownership Trend

27 quarters

Holder Concentration

Across 141 13F filers

Top Holder

10.9%

of institutional value

Top 5

33.2%

of institutional value

Top 10

51.9%

of institutional value

HHI

402

Broadly held

Who Holds ALEC

Last 8 quarters through Sep 30, 2026 (filing window open)

Each kind of 13F filer's share of the common shares 13F filers report. Holder type comes from regulatory records, never the filer's name; style, turnover and horizon are measured from each filer's own 13F book. The quarter selector does not change this card.

Jun 30, 2026, the newest closed quarter: 140 filers, like-for-like change +3.2%; actively managed books +3.6%; index-like books +2.0%.

Holder type	Share	Filers	Change	New	Exited
Registered adviser	55.0%	71	+17.2%	6	5
Exempt adviser	0.9%	3	+31.4%	2	0
Adviser	0.6%	3	-8.9%	0	0
Bank	3.9%	12	+3.1%	0	0
Insurer	0.0%	1	-32.7%	0	0
Broker-dealer	0.0%	1	new	1	0
Benefit plan	0.2%	3	-2.1%	0	0
Non-US	1.4%	5	-3.9%	0	0
Holding company	10.8%	7	-6.5%	1	0
Unknown	27.1%	34	-13.9%	3	4

Style	Share	Filers	Change	New	Exited
Index-like	21.0%	20	+2.0%	0	0
Systematic-like	11.7%	11	-1.9%	3	1
Concentrated	16.3%	24	-10.5%	4	3
Diversified active	51.0%	85	+10.6%	6	5
Not measured	0.0%	0	—	0	0
Unknown	0.0%	0	—	0	0

Turnover	Share	Filers	Change	New	Exited
Low turnover	28.8%	76	-2.5%	7	3
Medium turnover	44.5%	35	+9.3%	1	2
High turnover	14.9%	19	-3.6%	4	3
Unknown	11.8%	10	+5.9%	1	1
Not measured	0.0%	0	—	0	0

Holding horizon	Share	Filers	Change	New	Exited
Short horizon	12.3%	13	+3.3%	1	2
Medium horizon	34.6%	25	+12.0%	3	1
Long horizon	37.2%	91	-6.9%	6	6
Unknown	15.9%	11	+12.8%	3	0
Not measured	0.0%	0	—	0	0

13F book size	Share	Filers	Change	New	Exited
Under $1B	13.9%	23	-24.2%	3	4
$1B to $10B	32.8%	29	+26.9%	4	3
$10B to $100B	14.8%	39	+5.7%	5	1
$100B or more	38.5%	49	-0.4%	1	1
Not measured	0.0%	0	—	0	0

Client base	Share	Filers	Change	New	Exited
Individuals and high net worth	0.3%	24	+40.9%	5	3
Pooled investment vehicles	38.5%	23	+21.9%	1	1
Registered funds and BDCs	11.4%	8	+5.5%	0	0
Pension and profit sharing plans	0.0%	2	+49.6%	0	0
Insurance companies	0.1%	3	+48.3%	0	0
Governments and sovereign funds	1.0%	2	+38.1%	0	0
Other investment advisers	0.1%	2	-2.3%	0	0
Other	0.1%	1	-55.0%	0	1
Diversified	3.5%	6	+7.5%	0	0
Unknown	45.0%	69	-9.8%	7	4

Private funds	Share	Filers	Change	New	Exited
Hedge funds	42.2%	27	+20.4%	2	1
Private equity funds	0.2%	5	+103.1%	2	0
Venture capital funds	0.0%	1	+183.3%	0	0
Real estate funds	0.0%	3	+395.4%	0	1
Other private funds	1.8%	12	+1.4%	0	0
Mixed private funds	0.1%	1	+17.2%	0	0
Private funds, type not stated	0.2%	1	new	1	0
No private funds	11.3%	24	+7.5%	3	3
Securitized asset funds	0.0%	0	—	0	0
Unknown	44.1%	66	-10.4%	5	4

Location	Share	Filers	Change	New	Exited
United States	95.4%	116	+3.0%	10	8
Outside the US	4.6%	24	+7.7%	3	1

The Sep 30, 2026 filing window is still open, so its bar carries the Jun 30, 2026 positions of filers that have not filed yet.

How the mix is measured

Shares are 13F common shares restated to today's split basis; options and principal amounts are left out. The change compares only filers with a 13F in both quarters and keeps each filer in its bucket of the later quarter on both sides; a filer that continues under a new CIK is compared through its successor. Holder type comes from each filer's current regulatory profile; client base and private funds come from the Form ADV roster current at each quarter end (rosters from December 2019, so an earlier quarter reads Unknown). Style, turnover, horizon and book size are measured from each filer's 13F book in the newest measured quarter, and Not measured means none was. Actively managed books are those measured systematic-like, concentrated or diversified active.

Excluded 2 holders with no observed 2026-06-30 13F.

Conviction and Crowding

140 13F filers, quarter ended Jun 30, 2026

Measured across every 13F filer's common-share book once the quarter's filing window closed. Crowding counts filers with high turnover or that advise private funds on Form ADV.

Conviction holders

2.14% of filers hold it in their top 10

Mar 31, 2026: 4

Holders above 5%

median weight 0.000%

Mar 31, 2026: 0

Crowding

39.7%

of shares outstanding, 92nd percentile

Mar 31, 2026: 34.7%

Private-fund advisers

36.6%

of shares outstanding

Institutional ownership

82.2%

89.2% of the estimated free float

Top 10 holders

51.9%

of 13F shares, Herfindahl 0.041

Active holders

+3.6%

like-for-like share change of active filers

5% stakes on record

0 13D, 4 13G (2 institutional, 1 passive, 1 exempt)

How long holders have held it value-weighted 10.2 quarters

Under 1 year 41.3% 1 to 3 years 23.5% 3 years or more 35.1% Since coverage start 0.000%

Managers in the top fifth by current three-year FundScore alpha were net buyers of $0 (0 buying, 0 selling).

Estimated Holder Cost Basis

Quarter ended Jun 30, 2026, on that quarter's share basis

Estimated, never filed: each 13F buy is costed at its quarter's average traded price and each sale keeps the average cost, so this is what holders paid on average if they traded evenly through each quarter.

Estimated avg cost

$2.43

estimated per share

Mar 31, 2026: $2.47

Quarter-end price

$2.01

implied by 13F values

Estimated gain or loss

-17.3%

price against the estimated cost

Estimated at a loss

47.0%

of costed 13F value

Estimated cost covers 57.3% of the 13F shares; positions already held when a filer's 13F history starts, or bought in a quarter without a trusted average price, have no estimate.

Shared Holders

Companies ALEC's active holders also own, quarter ended Jun 30, 2026

#	Company	Shared holders	Similarity
1	LXEO Lexeo Therapeutics, Inc.	7	0.412
2	STRO Sutro Biopharma, Inc.	5	0.327
3	PRQR ProQR Therapeutics N.V.	6	0.324
4	AMAN Amanat Acquisition Corp.	4	0.314
5	IFRX InflaRx N.V.	5	0.304
6	BCRX Biocryst Pharmaceuticals Inc	7	0.296
7	AMLX Amylyx Pharmaceuticals, Inc.	7	0.292
8	EQ Equillium, Inc.	4	0.284
9	JATT JATT II Acquisition Corp.	4	0.284
10	SVRA Savara Inc	7	0.283

Shareholder Targets

13F filers that own its peers but not ALEC, quarter ended Jun 30, 2026

Found through its holder-overlap peers: LXEO, STRO, PRQR, AMAN, IFRX, BCRX, AMLX, EQ, JATT, SVRA, TRAX, PLRX, KRRO, SRZN, RLAY, PHVS, JANX, CATX, RACC, AVR. 549 filers in all.

#	Filer	In peers	Of book	Largest peers
1	PERCEPTIVE ADVISORS LLC Registered adviser, Diversified active	$614,878,103 7 held	9.57%	RLAY, AMLX, BCRX
2	SB INVESTMENT ADVISERS (UK) LTD Registered adviser, Concentrated	$522,101,857 1 held	6.81%	RLAY
3	Commodore Capital LP Registered adviser, Concentrated	$418,580,926 6 held	24.9%	RLAY, AMLX, PHVS
4	RA CAPITAL MANAGEMENT, L.P. Registered adviser, Concentrated	$402,368,819 7 held	3.50%	JANX, BCRX, AVR
5	Casdin Capital, LLC Registered adviser, Concentrated	$300,019,340 1 held	15.0%	RLAY
6	Jupiter Topco LLC Unknown, Diversified active	$266,945,525 10 held	0.110%	BCRX, AMLX, TRAX
7	GENERAL ATLANTIC, L.P. Unknown, Concentrated	$226,525,177 1 held	7.79%	PHVS
8	Bain Capital Life Sciences Investors, LLC Unknown, Concentrated	$200,938,926 2 held	12.0%	SVRA, PHVS
9	Foresite Capital Management IV, LLC Unknown, Concentrated	$188,630,830 2 held	65.0%	PHVS, RLAY
10	Polar Capital Holdings Plc Registered adviser, Diversified active	$172,697,169 3 held	0.490%	PHVS, SVRA, RLAY

Fund Ownership of ALEC by Fund Type

Registered funds' Form N-PORT reports for the quarter ended Jun 30, 2026

Index funds and index ETFs hold 11.10% of shares outstanding.

Fund type	Of shares outstanding	Change	Shares	Funds
Index funds	2.80%	+0.45 pp	3,126,405	54
Index ETFs	8.30%	+0.98 pp	9,263,273	22
Active ETFs	0.06%	+0.06 pp	67,665	4
Active funds	4.57%	-0.62 pp	5,107,579	46
Unclassified	0.20%	+0.20 pp	228,555	1
All registered funds	15.94%	+1.08 pp	17,793,477	127

How fund types are measured

Each fund series counts once, with its latest Form N-PORT report dated inside the quarter; only long share positions count, restated to today's split basis. The fund type is the series' latest Form N-CEN answer on or before the quarter end (the index-fund box and the exchange-traded flag), never its name. Unclassified funds have no usable N-CEN; unit investment trusts are among them, so index ownership is a floor. Funds sit inside their advisers' 13F reports, so these shares overlap the 13F figures and are never added to them.

Holder type: Registered adviser 71 Exempt adviser 3 Adviser 3 Bank 12 Insurer 1 Broker-dealer 1 Benefit plan 3 Non-US 4 Holding company 7 Unknown 34

Showing 100 of 143

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

TCG Crossover Management, LLC

$20,100,000

10.86%

10,000,000

BlackRock, Inc.

$15,774,232

8.53%

7,847,879

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	469,149	$942,989	6.0%	Sole
BlackRock Asset Management Canada Ltd	846	$1,700	0.0%	Sole
BlackRock Investment Management, LLC	295,587	$594,129	3.8%	Sole
BLACKROCK ADVISORS LLC	242,278	$486,978	3.1%	Sole
BlackRock Fund Advisors	3,919,590	$7,878,375	49.9%	Sole
BlackRock Institutional Trust Company, N.A.	2,905,617	$5,840,290	37.0%	Sole
BlackRock Japan Co. Ltd	9,725	$19,547	0.1%	Sole
BlackRock (Netherlands) B.V.	4,325	$8,693	0.1%	Sole
Aperio Group, LLC	762	$1,531	0.0%	Sole

ACADIAN ASSET MANAGEMENT LLC

$8,872,833

4.80%

4,414,345

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Acadian Inc	4,414,345	$8,872,833	100.0%	Defined

Nantahala Capital Management, LLC

$8,679,399

4.69%

4,318,109

VANGUARD CAPITAL MANAGEMENT LLC

$8,058,773

4.36%

4,009,340

Kynam Capital Management, LP

$7,655,072

4.14%

3,808,494

Merck & Co., Inc.

$7,126,895

3.85%

3,545,719

Balyasny Asset Management L.P.

$6,527,326

3.53%

3,247,426

683 Capital Management, LLC

$6,432,000

3.48%

3,200,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
683 Capital Partners, LP	3,200,000	$6,432,000	100.0%	Defined

D. E. Shaw & Co., Inc.

$5,949,565

3.22%

2,959,983

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	2,454,675	$4,933,896	82.9%	Defined
D. E. Shaw Investment Management, L.L.C. joint	505,308	$1,015,669	17.1%	Defined

ADAR1 Capital Management, LLC

$5,796,001

3.13%

2,883,583

VANGUARD PORTFOLIO MANAGEMENT LLC

$5,669,190

3.06%

2,820,493

GEODE CAPITAL MANAGEMENT, LLC

$4,933,084

2.67%

2,454,271

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	353,618	$710,772	14.4%	Defined
Held directly	2,100,653	$4,222,312	85.6%	Defined

Spruce Street Capital LP

$4,223,146

2.28%

2,101,068

MILLENNIUM MANAGEMENT LLC

$3,997,558

2.16%

1,988,835

Ikarian Capital, LLC

$3,985,038

2.15%

1,982,606

STATE STREET CORP

$3,778,327

2.04%

1,879,765

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	1,879,765	$3,778,327	100.0%	Defined

JACOBS LEVY EQUITY MANAGEMENT, INC

$3,549,517

1.92%

1,765,929

MEDICI CAPITAL LLC

$3,321,565

1.80%

1,652,520

Affinity Asset Advisors, LLC

$3,035,100

1.64%

1,510,000

Eversept Partners, LP

$2,861,984

1.55%

1,423,873

FMR LLC

$2,806,759

1.52%

1,396,398

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	1,396,028	$2,806,016	100.0%	Defined
STRATEGIC ADVISERS LLC	370	$743	0.0%	Defined

AQR CAPITAL MANAGEMENT LLC

$2,632,247

1.42%

1,309,577

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RF&L WEALTH MANAGEMENT, LLC	47	$94	0.0%	Other
Compound Planning, Inc.	22	$44	0.0%	Other
Arax Advisory Partners	5,096	$10,242	0.4%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	4,687	$9,420	0.4%	Other
Stokes Family Office, LLC	3,420	$6,874	0.3%	Other
Held directly	1,296,305	$2,605,573	99.0%	Defined

Euclidean Capital LLC

$2,628,794

1.42%

1,307,858

MACKENZIE FINANCIAL CORP

$2,230,637

1.21%

1,109,770

DEUTSCHE BANK AG\

$2,207,759

1.19%

1,098,388

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DWS INVESTMENT MANAGEMENT AMERICAS, INC	1,730	$3,477	0.2%	Defined
DWS Investments UK Ltd	1,096,658	$2,204,282	99.8%	Defined

DEERFIELD MANAGEMENT COMPANY, L.P.

$2,120,935

1.15%

1,055,192

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Deerfield UGP, LLC	1,055,192	$2,120,935	100.0%	Defined

TWO SIGMA INVESTMENTS, LP

$2,101,205

1.14%

1,045,376

RENAISSANCE TECHNOLOGIES LLC

$1,879,683

1.02%

935,166

DAFNA Capital Management LLC

$1,825,785

0.99%

908,351

Elmind Capital, LP

$1,809,000

0.98%

900,000

PANAGORA ASSET MANAGEMENT INC

$1,700,457

0.92%

845,999

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$1,645,850

0.89%

818,831

NORTHERN TRUST CORP

$1,554,657

0.84%

773,462

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	758,132	$1,523,844	98.0%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	15,330	$30,813	2.0%	Other

VANGUARD FIDUCIARY TRUST CO

$1,333,819

0.72%

663,592

MARSHALL WACE, LLP

$1,203,102

0.65%

598,559

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marshall Wace North America L.P. joint	598,559	$1,203,102	100.0%	Other

BOOTHBAY FUND MANAGEMENT, LLC

$1,071,028

0.58%

532,850

Balyasny Asset Management L.P.

$1,012,839

0.55%

503,900

GOLDMAN SACHS GROUP INC

$872,088

0.47%

433,875

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	274,318	$551,379	63.2%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	159,557	$320,709	36.8%	Defined

XTX Topco Ltd

$803,358

0.43%

399,681

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	399,681	$803,358	100.0%	Defined

CITADEL ADVISORS LLC

$786,376

0.43%

391,232

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	71,306	$143,325	18.2%	Defined
Held directly	319,926	$643,051	81.8%	Defined

CANTOR FITZGERALD, L. P.

$593,938

0.32%

295,492

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Cantor Fitzgerald & Co.	295,492	$593,938	100.0%	Defined

UBS Group AG

$550,297

0.30%

273,780

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS AG	273,780	$550,297	100.0%	Defined

Invesco Ltd.

$516,847

0.28%

257,139

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	14,074	$28,288	5.5%	Defined
Invesco Trust Co joint	38,786	$77,959	15.1%	Defined
Invesco Capital Management LLC	204,279	$410,600	79.4%	Defined

Bank of New York Mellon Corp

$502,686

0.27%

250,093

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Investments Corporation	81,174	$163,159	32.5%	Defined
The Bank of New York Mellon	168,918	$339,525	67.5%	Defined
Held directly	1	$2	0.0%	Defined

AMERIPRISE FINANCIAL INC

$451,305

0.24%

224,530

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	224,530	$451,305	100.0%	Defined

LMR Partners LLP

$402,000

0.22%

200,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
LMR Partners LLC	200,000	$402,000	100.0%	Sole

NANO CAP NEW MILLENNIUM GROWTH FUND L P

$402,000

0.22%

200,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AZG TANGIBLE ASSETS FUND LLC	200,000	$402,000	100.0%	Sole

Nuveen, LLC

$354,230

0.19%

176,234

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	1,425	$2,864	0.8%	Defined
TEACHERS ADVISORS, LLC joint	174,809	$351,366	99.2%	Defined

Utah Retirement Systems

$310,122

0.17%

154,290

BANK OF AMERICA CORP /DE/

$286,781

0.16%

142,678

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	92,398	$185,719	64.8%	Defined
MERRILL LYNCH, PIERCE, FENNER & SMITH INC.	25,976	$52,211	18.2%	Defined
BOFA SECURITIES, INC.	24,304	$48,851	17.0%	Defined

BRIDGEWAY CAPITAL MANAGEMENT, LLC

$286,425

0.15%

142,500

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$273,159

0.15%

135,900

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	135,900	$273,159	100.0%	Other

CANADA PENSION PLAN INVESTMENT BOARD

$263,913

0.14%

131,300

MORGAN STANLEY

$242,163

0.13%

120,480

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	530	$1,065	0.4%	Defined
MORGAN STANLEY & CO. LLC	28,673	$57,632	23.8%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	69,404	$139,502	57.6%	Defined
MORGAN STANLEY SMITH BARNEY LLC	10,408	$20,920	8.6%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	11,465	$23,044	9.5%	Defined

JANE STREET GROUP, LLC

$241,401

0.13%

120,100

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	120,100	$241,401	100.0%	Defined

PUBLIC EMPLOYEES RETIREMENT SYSTEM OF OHIO

$240,735

0.13%

119,769

RHUMBLINE ADVISERS

$220,159

0.12%

109,532

ALLIANCEBERNSTEIN L.P.

$218,969

0.12%

108,940

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	108,940	$218,969	100.0%	Defined

CITADEL ADVISORS LLC

$218,889

0.12%

108,900

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	108,900	$218,889	100.0%	Defined

STRS OHIO

$204,216

0.11%

101,600

Russell Investments Group, Ltd.

$197,973

0.11%

98,496

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC joint	66,609	$133,882	67.6%	Defined
Russell Investments Capital, LLC joint	9,499	$19,092	9.6%	Other
Russell Investments France SAS	5,248	$10,548	5.3%	Defined
Russell Investments Trust Company joint	17,140	$34,451	17.4%	Other

WELLS FARGO & COMPANY/MN

$179,235

0.10%

89,172

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	88,715	$178,317	99.5%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	257	$516	0.3%	Defined
WELLS FARGO CLEARING SERVICES, LLC	200	$402	0.2%	Defined

SIMPLEX TRADING, LLC

$171,830

0.09%

85,488

PRICE T ROWE ASSOCIATES INC /MD/

$164,126

0.09%

81,655

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	81,655	$164,126	100.0%	Defined

Blackstone Inc.

$155,061

0.08%

77,145

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Blackstone Real Estate Advisors L.P.	77,145	$155,061	100.0%	Other

Jasper Ridge Partners, L.P.

$147,797

0.08%

73,531

SEI INVESTMENTS CO

$143,013

0.08%

71,151

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI TRUST CO	71,151	$143,013	100.0%	Sole

DZ BANK AG Deutsche Zentral Genossenschafts Bank, Frankfurt am Main

$140,028

0.08%

69,666

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UNION ASSET MANAGEMENT HOLDING AG joint	69,666	$140,028	100.0%	Defined

IEQ CAPITAL, LLC

$121,745

0.07%

60,570

Verdad Advisers, LP

$121,623

0.07%

60,509

MetLife Investment Management, LLC

$117,978

0.06%

58,696

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
METLIFE INC	58,696	$117,978	100.0%	Defined

Mariner, LLC

$105,945

0.06%

52,709

Vanguard Personalized Indexing Management, LLC

$99,384

0.05%

49,445

HSBC HOLDINGS PLC

$97,531

0.05%

48,523

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
HSBC BANK PLC	48,523	$97,531	100.0%	Defined

Graham Capital Management, L.P.

$93,456

0.05%

46,496

CITIGROUP INC

$93,012

0.05%

46,275

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	46,275	$93,012	100.0%	Defined

MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.

$90,831

0.05%

45,190

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Global X Management CO LLC joint	45,190	$90,831	100.0%	Defined

BARCLAYS PLC

$90,363

0.05%

44,957

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	41,317	$83,047	91.9%	Defined
BARCLAYS CAPITAL INC.	3,640	$7,316	8.1%	Defined

JPMORGAN CHASE & CO

$86,425

0.05%

42,998

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	42,000	$84,420	97.7%	Defined
J.P. Morgan Investment Management Inc.	493	$990	1.1%	Defined
J.P. Morgan Securities plc	505	$1,015	1.2%	Defined

Diversified Trust Co

$85,125

0.05%

42,351

DIMENSIONAL FUND ADVISORS LP

$67,792

0.04%

33,728

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	3,487	$7,008	10.3%	Defined
Dimensional Ireland Ltd	11,623	$23,362	34.5%	Defined
Held directly	18,618	$37,422	55.2%	Sole

Cerity Partners LLC

$65,366

0.04%

32,521

Vanguard Global Advisers, LLC

$62,917

0.03%

31,302

Cannell & Spears LLC

$60,987

0.03%

30,342

Point72 Asset Management, L.P.

$53,664

0.03%

26,699

ROYAL BANK OF CANADA

$52,504

0.03%

26,122

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC CMA LTD.	2,973	$5,975	11.4%	Defined
RBC Capital Markets, LLC	23,149	$46,529	88.6%	Defined

MERCER GLOBAL ADVISORS INC /ADV

$51,263

0.03%

25,504

GROUP ONE TRADING LLC

$50,668

0.03%

25,208

INTECH INVESTMENT MANAGEMENT LLC

$49,305

0.03%

24,530

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$48,487

0.03%

24,123

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	24,123	$48,487	100.0%	Other

Police & Firemen's Retirement System of New Jersey

$47,086

0.03%

23,426

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Capital	23,426	$47,086	100.0%	Defined

VOYA INVESTMENT MANAGEMENT LLC

$44,014

0.02%

21,898

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	21,898	$44,014	100.0%	Defined

BNP PARIBAS ARBITRAGE, SA

$42,612

0.02%

21,200

MANUFACTURERS LIFE INSURANCE COMPANY, THE

$38,987

0.02%

19,397

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MANULIFE FINANCIAL CORP joint	19,397	$38,987	100.0%	Defined

Qube Research & Technologies Ltd

$33,303

0.02%

16,569

CAPTRUST FINANCIAL ADVISORS

$32,234

0.02%

16,037

PRUDENTIAL FINANCIAL INC

$32,190

0.02%

16,015

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PGIM Quantitative Solutions LLC	16,015	$32,190	100.0%	Defined

Creative Planning

$30,180

0.02%

15,015

Tower Research Capital LLC (TRC)

$29,044

0.02%

14,450

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Latour Trading LLC (Latour)	14,450	$29,044	100.0%	Defined

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Key facts CIK 1653087 CUSIP 014442107 13F (30d) 17 filings 15 filers Visit website Investor relations