NTLA · Intellia Therapeutics, Inc. · Holdings

$11.84 +0.26 (+2.25%) At close · Oct 20

Russell 2000 member Rebalance outlook locked

Market Cap

$1.62B

140.13M

Volume · Oct 20 3.42M Avg daily vol (3M) 4.33M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Use institutional holdings through the 13F API or MCP → See every holder of NTLA: 13F, 13D and 13G, insiders and funds →

Institutional holdings are being reconciled with SEC filings. Holder totals and changes may be incomplete until reconciliation finishes.

The Sep 30, 2026 filing window is still open — showing the latest known positions. Funds that have not filed yet carry their prior-quarter positions; a fund that filed without this stock counts as sold out. Each fund's Change is measured against its own previous reported quarter, shown in the Reported column. Pick a completed quarter in the Quarter selector below to put every fund on one quarter and see the stock-wide activity breakdown.

Quarter:

Holders: 328

Total Value: $2,500,498,347

Total Shares: 148,280,102

Showing 1–100 of 347 holdings by value · page 1 of 4

Institutional Ownership Trend

27 quarters

Holder Concentration

Across 333 13F filers

Top Holder

11.4%

of institutional value

Top 5

37.4%

of institutional value

Top 10

53.8%

of institutional value

HHI

423

Broadly held

Who Holds NTLA

Last 8 quarters through Sep 30, 2026 (filing window open)

Each kind of 13F filer's share of the common shares 13F filers report. Holder type comes from regulatory records, never the filer's name; style, turnover and horizon are measured from each filer's own 13F book. The quarter selector does not change this card.

Jun 30, 2026, the newest closed quarter: 328 filers, like-for-like change +30.3%; actively managed books +34.2%; index-like books +25.9%.

Holder type	Share	Filers	Change	New	Exited
Registered adviser	51.0%	171	+41.1%	29	15
Exempt adviser	0.7%	6	-1.2%	2	0
Adviser	1.0%	7	+43.0%	2	0
Bank	13.2%	18	+19.7%	0	0
Insurer	0.0%	1	+26.6%	0	0
Broker-dealer	0.2%	2	+43.4%	0	0
Benefit plan	0.3%	6	+76.5%	2	0
Non-US	0.6%	6	+3.2%	0	1
Holding company	16.9%	8	+34.9%	0	0
Unknown	16.0%	103	+9.4%	16	12

Style	Share	Filers	Change	New	Exited
Index-like	32.2%	39	+25.9%	5	1
Systematic-like	12.3%	31	+16.2%	11	5
Concentrated	6.4%	45	+146.6%	11	7
Diversified active	46.5%	211	+31.4%	23	15
Unknown	2.6%	2	0.0%	1	0
Not measured	0.0%	0	—	0	0

Turnover	Share	Filers	Change	New	Exited
Low turnover	48.7%	179	+37.3%	17	13
Medium turnover	24.8%	74	+28.4%	12	5
High turnover	16.2%	48	+23.8%	17	8
Unknown	10.3%	27	+16.4%	5	2
Not measured	0.0%	0	—	0	0

Holding horizon	Share	Filers	Change	New	Exited
Short horizon	15.8%	48	+19.9%	15	6
Medium horizon	13.2%	48	+25.8%	7	7
Long horizon	60.5%	205	+34.6%	21	14
Unknown	10.5%	27	+29.2%	8	1
Not measured	0.0%	0	—	0	0

13F book size	Share	Filers	Change	New	Exited
Under $1B	3.9%	70	+8.6%	18	8
$1B to $10B	8.7%	89	+40.3%	16	10
$10B to $100B	26.0%	101	+50.0%	15	9
$100B or more	61.3%	68	+23.8%	2	1
Not measured	0.0%	0	—	0	0

Client base	Share	Filers	Change	New	Exited
Individuals and high net worth	1.6%	83	+12.8%	12	7
Pooled investment vehicles	19.7%	34	+95.0%	12	5
Registered funds and BDCs	26.8%	18	+18.5%	1	1
Pension and profit sharing plans	0.0%	3	+1.6%	0	0
Insurance companies	0.1%	3	+26.3%	0	0
Charitable organizations	0.0%	1	0.0%	0	0
Governments and sovereign funds	0.4%	4	+236.7%	2	0
Corporations	0.0%	1	+19.6%	0	0
Other investment advisers	0.0%	5	-2.4%	0	0
Other	0.0%	2	+45.4%	1	0
Diversified	2.3%	17	+35.1%	1	1
Unknown	49.0%	157	+20.7%	22	14

Private funds	Share	Filers	Change	New	Exited
Hedge funds	28.2%	43	+42.2%	12	5
Private equity funds	0.3%	15	+39.1%	2	1
Venture capital funds	3.1%	2	+1163718.8%	1	0
Real estate funds	0.0%	3	+119.3%	0	1
Securitized asset funds	0.2%	1	+10.2%	0	0
Other private funds	5.2%	14	+24.4%	0	0
Mixed private funds	0.3%	2	+0.3%	0	1
Private funds, type not stated	0.1%	7	+18.0%	3	2
No private funds	14.2%	90	+18.3%	13	4
Unknown	48.3%	151	+21.1%	20	14

Location	Share	Filers	Change	New	Exited
United States	89.5%	266	+34.4%	44	23
Outside the US	10.5%	62	+3.3%	7	5

The Sep 30, 2026 filing window is still open, so its bar carries the Jun 30, 2026 positions of filers that have not filed yet.

How the mix is measured

Shares are 13F common shares restated to today's split basis; options and principal amounts are left out. The change compares only filers with a 13F in both quarters and keeps each filer in its bucket of the later quarter on both sides; a filer that continues under a new CIK is compared through its successor. Holder type comes from each filer's current regulatory profile; client base and private funds come from the Form ADV roster current at each quarter end (rosters from December 2019, so an earlier quarter reads Unknown). Style, turnover, horizon and book size are measured from each filer's 13F book in the newest measured quarter, and Not measured means none was. Actively managed books are those measured systematic-like, concentrated or diversified active.

Excluded 2 holders with no observed 2026-03-31 13F and 1 holder with no observed 2026-06-30 13F. Compared across CIK changes: JANUS HENDERSON GROUP PLC continues as Jupiter Topco LLC.

Conviction and Crowding

328 13F filers, quarter ended Jun 30, 2026

Measured across every 13F filer's common-share book once the quarter's filing window closed. Crowding counts filers with high turnover or that advise private funds on Form ADV.

Conviction holders

0.610% of filers hold it in their top 10

Mar 31, 2026: 2

Holders above 5%

median weight 0.002%

Mar 31, 2026: 1

Crowding

—

holders exceed shares outstanding: 13F filers report more shares than the cover-page count, which predates an issuance, is misstated, or misses shares lent and reported twice

Mar 31, 2026: 37.3%

Private-fund advisers

—

of shares outstanding

Institutional ownership

—

holders exceed shares outstanding: 13F filers report more shares than the cover-page count, which predates an issuance, is misstated, or misses shares lent and reported twice

Top 10 holders

55.3%

of 13F shares, Herfindahl 0.044

Active holders

+34.2%

like-for-like share change of active filers

5% stakes on record

0 13D, 7 13G (7 institutional, 0 passive, 0 exempt)

How long holders have held it value-weighted 14.9 quarters

Under 1 year 23.6% 1 to 3 years 28.9% 3 years or more 47.6% Since coverage start 0.000%

Managers in the top fifth by current three-year FundScore alpha were net sellers of $96,574 (0 buying, 1 selling).

Estimated Holder Cost Basis

Quarter ended Jun 30, 2026, on that quarter's share basis

Estimated, never filed: each 13F buy is costed at its quarter's average traded price and each sale keeps the average cost, so this is what holders paid on average if they traded evenly through each quarter.

Estimated avg cost

$14.63

estimated per share

Mar 31, 2026: $15.27

Quarter-end price

$16.91

implied by 13F values

Estimated gain or loss

+15.6%

price against the estimated cost

Estimated at a loss

8.76%

of costed 13F value

Estimated cost covers 30.7% of the 13F shares; positions already held when a filer's 13F history starts, or bought in a quarter without a trusted average price, have no estimate.

Shared Holders

Companies NTLA's active holders also own, quarter ended Jun 30, 2026

#	Company	Shared holders	Similarity
1	BEAM Beam Therapeutics Inc.	9	0.296
2	TEM Tempus AI, Inc.	12	0.254
3	PRME Prime Medicine, Inc.	5	0.253
4	PACB Pacific Biosciences Of California, Inc.	5	0.236
5	CRSP CRISPR Therapeutics AG	10	0.233
6	ABSI Absci Corp	6	0.197
7	IRON Disc Medicine, Inc.	6	0.192
8	TWST Twist Bioscience Corp	7	0.187
9	BBOT BridgeBio Oncology Therapeutics, Inc.	4	0.183
10	EDIT Editas Medicine, Inc.	4	0.183

Shareholder Targets

13F filers that own its peers but not NTLA, quarter ended Jun 30, 2026

Found through its holder-overlap peers: BEAM, TEM, PRME, PACB, CRSP, ABSI, IRON, TWST, BBOT, EDIT, VYGR, RXRX, FDMT, CERS, PSNL, JBIO, AKTS, STOK, PONY, SANA. 952 filers in all.

#	Filer	In peers	Of book	Largest peers
1	BAILLIE GIFFORD & CO Adviser, Diversified active	$786,498,814 6 held	0.710%	TEM, TWST, PONY
2	FARALLON CAPITAL MANAGEMENT LLC Registered adviser, Diversified active	$575,708,508 4 held	3.40%	BEAM, TWST, PRME
3	Artisan Partners Limited Partnership Registered adviser, Diversified active	$505,652,114 2 held	0.830%	TWST, STOK
4	SOFTBANK GROUP CORP. Unknown, Concentrated	$397,539,816 3 held	2.19%	TEM, RXRX, PACB
5	T. Rowe Price Investment Management, Inc. Adviser, Diversified active	$371,362,435 6 held	0.240%	CRSP, STOK, AKTS
6	Orbis Allan Gray Ltd Unknown, Diversified active	$297,197,240 1 held	1.02%	CRSP
7	EcoR1 Capital, LLC Registered adviser, Concentrated	$276,825,168 4 held	13.7%	AKTS, CRSP, STOK
8	RTW INVESTMENTS, LP Registered adviser, Diversified active	$263,410,707 5 held	2.25%	STOK, AKTS, BEAM
9	EdgePoint Investment Group Inc. Registered adviser, Concentrated	$259,425,808 1 held	1.96%	TWST
10	Capital World Investors Adviser, Diversified active	$259,039,093 1 held	0.030%	CRSP

Fund Ownership of NTLA by Fund Type

Registered funds' Form N-PORT reports for the quarter ended Jun 30, 2026

Index funds and index ETFs hold 21.53% of shares outstanding.

Fund type	Of shares outstanding	Change	Shares	Funds
Index funds	3.10%	+0.75 pp	4,342,515	59
Index ETFs	18.43%	+3.85 pp	25,822,193	47
Active ETFs	10.53%	+1.26 pp	14,759,825	18
Active funds	7.84%	+3.03 pp	10,981,081	91
Unclassified	0.59%	+0.59 pp	830,059	10
All registered funds	40.49%	+9.48 pp	56,735,673	225

How fund types are measured

Each fund series counts once, with its latest Form N-PORT report dated inside the quarter; only long share positions count, restated to today's split basis. The fund type is the series' latest Form N-CEN answer on or before the quarter end (the index-fund box and the exchange-traded flag), never its name. Unclassified funds have no usable N-CEN; unit investment trusts are among them, so index ownership is a floor. Funds sit inside their advisers' 13F reports, so these shares overlap the 13F figures and are never added to them.

Holder type: Registered adviser 167 Exempt adviser 6 Adviser 7 Bank 18 Insurer 1 Broker-dealer 2 Benefit plan 6 Non-US 6 Holding company 8 Unknown 107

Showing 100 of 347

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

ARK Investment Management LLC

$285,536,573

11.42%

16,875,684

BlackRock, Inc.

$260,720,659

10.43%

15,409,023

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	540,106	$9,138,593	3.5%	Sole
BlackRock Investment Management (Australia) Ltd	246	$4,162	0.0%	Sole
BlackRock Asset Management Canada Ltd	3,514	$59,456	0.0%	Sole
BlackRock Investment Management, LLC	507,927	$8,594,124	3.3%	Sole
BLACKROCK ADVISORS LLC	1,863,137	$31,524,278	12.1%	Sole
BlackRock Fund Advisors	6,227,314	$105,366,152	40.4%	Sole
BlackRock Institutional Trust Company, N.A.	4,858,201	$82,200,760	31.5%	Sole
BlackRock Japan Co. Ltd	9,269	$156,831	0.1%	Sole
BlackRock Fund Managers Ltd.	24,289	$410,969	0.2%	Sole
BlackRock Investment Management (UK) Ltd.	206,690	$3,497,194	1.3%	Sole
BlackRock (Netherlands) B.V.	9,773	$165,359	0.1%	Sole
BlackRock Asset Management Ireland Ltd	237,167	$4,012,865	1.5%	Sole
BlackRock (Luxembourg) S.A.	752,366	$12,730,032	4.9%	Sole
BlackRock Life Ltd	718	$12,148	0.0%	Sole
BlackRock Asset Management Schweiz AG	18,382	$311,023	0.1%	Sole
Aperio Group, LLC	149,686	$2,532,687	1.0%	Sole
SpiderRock Advisors, LLC	238	$4,026	0.0%	Sole

STATE STREET CORP

$156,255,980

6.25%

9,234,987

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	9,234,987	$156,255,980	100.0%	Defined

VANGUARD PORTFOLIO MANAGEMENT LLC

$124,064,224

4.96%

7,332,401

D. E. Shaw & Co., Inc.

$109,021,025

4.36%

6,443,323

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	6,443,323	$109,021,025	100.0%	Defined

VANGUARD CAPITAL MANAGEMENT LLC

$102,003,675

4.08%

6,028,586

TWO SIGMA INVESTMENTS, LP

$88,582,088

3.54%

5,235,348

FMR LLC

$80,759,733

3.23%

4,773,034

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	4,098,904	$69,353,455	85.9%	Defined
STRATEGIC ADVISERS LLC	8,642	$146,222	0.2%	Defined
Fidelity Institutional Asset Management Trust Co	4,388	$74,244	0.1%	Defined
FIAM LLC	661,100	$11,185,812	13.9%	Defined

RA CAPITAL MANAGEMENT, L.P.

$76,595,587

3.06%

4,526,926

REGENERON PHARMACEUTICALS, INC.

$62,654,675

2.51%

3,702,995

GEODE CAPITAL MANAGEMENT, LLC

$61,437,551

2.46%

3,631,061

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	518,070	$8,765,744	14.3%	Defined
Held directly	3,112,991	$52,671,807	85.7%	Defined

ARROWSTREET CAPITAL, LIMITED PARTNERSHIP

$55,510,459

2.22%

3,280,760

Sumitomo Mitsui Trust Group, Inc.

$49,026,190

1.96%

2,897,529

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Amova Asset Management Co., Ltd.	2,897,529	$49,026,190	100.0%	Defined

Amova Asset Management Americas, Inc.

$49,026,190

1.96%

2,897,529

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Sumitomo Mitsui Trust Group, Inc. joint	2,897,529	$49,026,190	100.0%	Sole

DIMENSIONAL FUND ADVISORS LP

$44,041,810

1.76%

2,602,944

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	19,783	$334,728	0.8%	Defined
DFA Australia Ltd.	45,517	$770,147	1.7%	Defined
Dimensional Fund Advisors Pte. Ltd.	1,713	$28,983	0.1%	Defined
Dimensional Ireland Ltd	102,498	$1,734,266	3.9%	Defined
Held directly	2,433,433	$41,173,686	93.5%	Sole

GOLDMAN SACHS GROUP INC

$38,314,849

1.53%

2,264,471

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	1,338,510	$22,647,589	59.1%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	925,961	$15,667,260	40.9%	Defined

UBS Group AG

$36,427,269

1.46%

2,152,912

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	170,319	$2,881,797	7.9%	Defined
UBS AG	1,923,624	$32,547,718	89.3%	Defined
UBS Switzerland AG	52,002	$879,873	2.4%	Defined
UBS Securities LLC	6,967	$117,881	0.3%	Defined

VOLORIDGE INVESTMENT MANAGEMENT, LLC

$34,962,743

1.40%

2,066,356

Contrarius Group Holdings Ltd

$33,082,457

1.32%

1,955,228

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Contrarius Investment Management Ltd joint	1,955,228	$33,082,457	100.0%	Defined

JUPITER ASSET MANAGEMENT LTD

$31,293,657

1.25%

1,849,507

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jupiter Investment Management Limited	19,113	$323,391	1.0%	Defined
Held directly	1,830,394	$30,970,266	99.0%	Sole

MORGAN STANLEY

$29,303,828

1.17%

1,731,905

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	24,858	$420,597	1.4%	Defined
MORGAN STANLEY & CO. LLC	670,492	$11,344,724	38.7%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	173,900	$2,942,388	10.0%	Defined
MORGAN STANLEY SMITH BARNEY LLC	265,584	$4,493,680	15.3%	Defined
Morgan Stanley Wealth Management Australia Pty Ltd	10,688	$180,840	0.6%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	489,383	$8,280,359	28.3%	Defined
EATON VANCE MANAGEMENT	97,000	$1,641,240	5.6%	Defined

ARMISTICE CAPITAL, LLC

$25,380,000

1.01%

1,500,000

RENAISSANCE TECHNOLOGIES LLC

$23,688,067

0.95%

1,400,004

NORTHERN TRUST CORP

$21,524,539

0.86%

1,272,136

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	1,165,210	$19,715,353	91.6%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	86,000	$1,455,119	6.8%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	20,926	$354,067	1.6%	Other

HRT FINANCIAL LP

$21,036,923

0.84%

1,243,317

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$20,438,361

0.82%

1,207,941

Ghisallo Capital Management LLC

$20,304,000

0.81%

1,200,000

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$18,227,916

0.73%

1,077,300

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	1,077,300	$18,227,916	100.0%	Other

Woodline Partners LP

$18,194,905

0.73%

1,075,349

STIFEL FINANCIAL CORP

$17,163,427

0.69%

1,014,387

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STIFEL NICOLAUS & CO INC \MO\	1,014,315	$17,162,209	100.0%	Defined
Held directly	72	$1,218	0.0%	Defined

BANK OF AMERICA CORP /DE/

$16,517,571

0.66%

976,216

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	454,042	$7,682,390	46.5%	Defined
MERRILL LYNCH, PIERCE, FENNER & SMITH INC.	96,078	$1,625,639	9.8%	Defined
BOFA SECURITIES, INC.	287,092	$4,857,596	29.4%	Defined
MANAGED ACCOUNT ADVISORS LLC joint	139,004	$2,351,946	14.2%	Defined

BAKER BROS. ADVISORS LP

$15,739,525

0.63%

930,232

VANGUARD FIDUCIARY TRUST CO

$15,585,367

0.62%

921,121

Connor, Clark & Lunn Investment Management Ltd.

$15,210,453

0.61%

898,963

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 1	898,963	$15,210,453	100.0%	Sole

IQ EQ FUND MANAGEMENT (IRELAND) Ltd

$13,162,880

0.53%

777,948

JACOBS LEVY EQUITY MANAGEMENT, INC

$12,284,884

0.49%

726,057

FEDERATED HERMES, INC.

$12,283,411

0.49%

725,970

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FEDERATED EQUITY MANAGEMENT CO OF PENNSYLVANIA	57,188	$967,620	7.9%	Defined
FEDERATED MDTA LLC	668,782	$11,315,791	92.1%	Defined

RAYMOND JAMES FINANCIAL INC

$11,833,677

0.47%

699,390

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	557,211	$9,428,010	79.7%	Defined
Raymond James Financial Services Advisors, Inc.	142,128	$2,404,805	20.3%	Defined
ClariVest Asset Management LLC joint	51	$862	0.0%	Defined

ROYAL BANK OF CANADA

$11,392,910

0.46%

673,340

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC CMA LTD.	4,113	$69,591	0.6%	Defined
RBC Capital Markets, LLC	662,505	$11,209,584	98.4%	Defined
RBC Trust Co (Delaware) Ltd	6,720	$113,702	1.0%	Defined
Held directly	2	$33	0.0%	Sole

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$10,958,728

0.44%

647,679

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	647,679	$10,958,728	100.0%	Other

Marex Group plc

$10,551,312

0.42%

623,600

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marex Financial	623,600	$10,551,312	100.0%	Defined

BNP PARIBAS ARBITRAGE, SA

$9,776,799

0.39%

577,825

Vestmark Advisory Solutions, Inc.

$9,015,263

0.36%

532,817

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ELEVATION POINT WEALTH PARTNERS, LLC	176,787	$2,991,236	33.2%	Other
Held directly	356,030	$6,024,027	66.8%	Sole

JPMORGAN CHASE & CO

$8,928,038

0.36%

527,662

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	76,212	$1,289,507	14.4%	Defined
J.P. Morgan Investment Management Inc.	215,685	$3,649,390	40.9%	Defined
J.P. Morgan Securities LLC	115	$1,945	0.0%	Defined
J.P. Morgan Securities plc	10,427	$176,424	2.0%	Defined
JPMorgan Asset Management (UK) Ltd	218,240	$3,692,620	41.4%	Defined
J.P. Morgan SE	6,983	$118,152	1.3%	Defined

AQR CAPITAL MANAGEMENT LLC

$8,340,777

0.33%

492,954

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RF&L WEALTH MANAGEMENT, LLC	29	$490	0.0%	Other
Prism Planning Partners LLC	331	$5,600	0.1%	Other
Compound Planning, Inc.	23	$389	0.0%	Other
Arax Advisory Partners	3,642	$61,622	0.7%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	1,439	$24,347	0.3%	Other
Stokes Family Office, LLC	1,013	$17,139	0.2%	Other
Held directly	486,477	$8,231,190	98.7%	Defined

MILLENNIUM MANAGEMENT LLC

$8,311,543

0.33%

491,226

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$7,267,140

0.29%

429,500

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	429,500	$7,267,140	100.0%	Other

Invesco Ltd.

$6,971,207

0.28%

412,010

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	18,578	$314,338	4.5%	Defined
Invesco Trust Co joint	53,525	$905,643	13.0%	Defined
Invesco Capital Management LLC	339,907	$5,751,226	82.5%	Defined

Bank of New York Mellon Corp

$6,948,263

0.28%

410,654

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Investments Corporation	150,047	$2,538,795	36.5%	Defined
The Bank of New York Mellon	242,742	$4,107,194	59.1%	Defined
BNY Mellon, NA joint	15,888	$268,824	3.9%	Defined
BNY Mellon Advisors, Inc.	1,975	$33,417	0.5%	Defined
Held directly	2	$33	0.0%	Defined

Caption Management, LLC

$6,757,848

0.27%

399,400

Point72 Asset Management, L.P.

$6,741,993

0.27%

398,463

Birchview Capital, LP

$6,529,428

0.26%

385,900

Nuveen, LLC

$6,389,770

0.26%

377,646

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	16,459	$278,486	4.4%	Defined
TEACHERS ADVISORS, LLC joint	361,187	$6,111,284	95.6%	Defined

WELLS FARGO & COMPANY/MN

$6,378,499

0.26%

376,980

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	117,688	$1,991,280	31.2%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	70,280	$1,189,137	18.6%	Defined
WELLS FARGO CLEARING SERVICES, LLC	188,964	$3,197,270	50.1%	Defined
WELLS FARGO INVESTMENT INSTITUTE, INC.	48	$812	0.0%	Defined

CITADEL ADVISORS LLC

$6,321,312

0.25%

373,600

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	373,600	$6,321,312	100.0%	Defined

Casdin Capital, LLC

$6,006,600

0.24%

355,000

FIRST TRUST ADVISORS LP

$5,749,534

0.23%

339,807

HIMENSION CAPITAL (SINGAPORE) PTE. LTD.

$5,531,029

0.22%

326,893

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$5,292,723

0.21%

312,809

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	3,670	$62,096	1.2%	Defined
UBS Asset Management Switzerland AG	232,988	$3,942,156	74.5%	Defined
UBS Asset Management (UK) Ltd.	52,221	$883,577	16.7%	Defined
Held directly	23,930	$404,894	7.7%	Defined

CAPITAL FUND MANAGEMENT S.A.

$5,048,589

0.20%

298,380

Trexquant Investment LP

$5,045,104

0.20%

298,174

JANE STREET GROUP, LLC

$5,030,316

0.20%

297,300

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	297,300	$5,030,316	100.0%	Defined

JANE STREET GROUP, LLC

$4,999,860

0.20%

295,500

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	295,500	$4,999,860	100.0%	Defined

UBS Group AG

$4,967,508

0.20%

293,588

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS AG	293,588	$4,967,508	100.0%	Defined

PINNACLE ASSOCIATES LTD

$4,399,427

0.18%

364,795

Swiss National Bank

$4,350,132

0.17%

257,100

CITADEL ADVISORS LLC

$4,238,460

0.17%

250,500

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	250,500	$4,238,460	100.0%	Defined

Opaleye Management Inc.

$4,230,000

0.17%

250,000

AMERIPRISE FINANCIAL INC

$4,068,430

0.16%

240,451

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	111,821	$1,892,011	46.5%	Defined
Ameriprise Financial Services, LLC	128,524	$2,174,626	53.5%	Defined
Ameriprise Bank, FSB	106	$1,793	0.0%	Defined

AMERICAN CENTURY COMPANIES INC

$4,044,150

0.16%

239,016

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AMERICAN CENTURY INVESTMENT MANAGEMENT INC	239,016	$4,044,150	100.0%	Defined

CITIGROUP INC

$3,946,708

0.16%

233,257

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	233,257	$3,946,708	100.0%	Defined

Assenagon Asset Management S.A.

$3,874,564

0.15%

321,274

DEUTSCHE BANK AG\

$3,562,506

0.14%

210,550

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DWS Investments UK Ltd	210,550	$3,562,506	100.0%	Defined

Susquehanna Portfolio Strategies, LLC

$3,544,080

0.14%

209,461

MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.

$3,414,505

0.14%

201,803

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Global X Management CO LLC joint	183,265	$3,100,843	90.8%	Defined
Global X Management (AUS) Ltd joint	18,538	$313,662	9.2%	Defined

XTX Topco Ltd

$3,408,297

0.14%

201,436

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	201,436	$3,408,297	100.0%	Defined

DIADEMA PARTNERS LP

$3,384,000

0.14%

200,000

LPL Financial LLC

$3,239,841

0.13%

191,480

Universal- Beteiligungs- und Servicegesellschaft mbH

$3,226,187

0.13%

190,673

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Universal-Investment-Gesellschaft mbH joint	190,673	$3,226,187	100.0%	Defined

PRICE T ROWE ASSOCIATES INC /MD/

$3,225,138

0.13%

190,611

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	190,611	$3,225,138	100.0%	Defined

Hennion & Walsh Asset Management, Inc.

$3,209,998

0.13%

266,169

ELEVATION POINT WEALTH PARTNERS, LLC

$3,184,462

0.13%

188,207

SECTOR GAMMA AS

$3,133,194

0.13%

185,177

RHUMBLINE ADVISERS

$2,990,187

0.12%

176,725

CITADEL ADVISORS LLC

$2,786,927

0.11%

164,712

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	164,712	$2,786,927	100.0%	Defined

MARSHALL WACE, LLP

$2,776,198

0.11%

164,078

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marshall Wace North America L.P. joint	164,078	$2,776,198	100.0%	Other

Canal Insurance CO

$2,772,594

0.11%

229,900

Rafferty Asset Management, LLC

$2,700,110

0.11%

159,581

BLAIR WILLIAM & CO/IL

$2,582,195

0.10%

152,612

CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM

$2,580,587

0.10%

152,517

BARCLAYS PLC

$2,564,546

0.10%

151,569

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	143,617	$2,429,999	94.8%	Defined
BARCLAYS CAPITAL INC.	7,952	$134,547	5.2%	Defined

Point72 Asset Management, L.P.

$2,431,404

0.10%

143,700

ALLIANCEBERNSTEIN L.P.

$2,425,482

0.10%

143,350

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	143,350	$2,425,482	100.0%	Defined

Rockefeller Capital Management L.P.

$2,400,777

0.10%

141,890

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ROCKEFELLER & CO. LLC	443	$7,495	0.3%	Defined
ROCKEFELLER TRUST COMPANY, N.A.	492	$8,324	0.3%	Defined
ROCKEFELLER FINANCIAL LLC	140,955	$2,384,958	99.3%	Defined

Jefferies Financial Group Inc.

$2,233,440

0.09%

132,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JEFFERIES LLC	132,000	$2,233,440	100.0%	Defined

Russell Investments Group, Ltd.

$2,170,036

0.09%

128,253

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC joint	84,830	$1,435,322	66.1%	Defined
Russell Investments Capital, LLC joint	15,133	$256,049	11.8%	Other
Russell Investments France SAS	3,802	$64,329	3.0%	Defined
Russell Investments Trust Company joint	24,488	$414,336	19.1%	Other

TUDOR INVESTMENT CORP ET AL

$1,978,083

0.08%

116,908

Legal & General Group Plc

$1,928,980

0.08%

114,006

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	13,117	$221,939	11.5%	Defined
Legal & General Investment Management Ltd joint	100,889	$1,707,041	88.5%	Defined

GROUP ONE TRADING LLC

$1,925,969

0.08%

113,828

OPPENHEIMER & CO INC

$1,818,104

0.07%

107,453

Showing the top 100 of 347 holders. Create a free account to see every holder, past quarters, search, filters and CSV export.

Key facts CIK 1652130 CUSIP 45826J105 13F (30d) 38 filings 34 filers Visit website Investor relations