FDMT · 4D Molecular Therapeutics, Inc.

$12.19 +0.54 (+4.64%) At close · Aug 14

Market Cap

$879.87M

72.18M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Quarter:

Holders: 160

Total Value: $778,351,347

Total Shares: 58,552,645

Institutional Ownership Trend

23 quarters

Holder Concentration

Across 160 13F filers · Δ vs Mar 31, 2026

Top Holder

9.8%

−0.5 pts

Top 5

39.0%

−4.8 pts

Top 10

60.9%

−5.4 pts

HHI

466

Broadly held−71

Quarterly Activity

vs Mar 31, 2026

New

Increased

Decreased

Sold Out

Unchanged

Holder count: +8

Institutional value: +56.1%

Shares held: +9.4%

Top Buyers

vs Mar 31, 2026

Institution	Added	Now Holds
Small Trails Edge Capital Partners, LP	+925,000	2,227,917
Large ADAGE CAPITAL PARTNERS GP, L.L.C.	+644,047	2,249,203
Mega CITADEL ADVISORS LLC	+584,101	901,197
Small Parkman Healthcare Partners LLC	+425,607	948,184
Mega BlackRock, Inc.	+400,756	4,546,059

Top Sellers

vs Mar 31, 2026

Institution	Sold	Now Holds
Mega STATE STREET CORP	−2,504,641	2,911,109
Large FEDERATED HERMES, INC.	−1,209,844	222,416
Mid NOVO HOLDINGS A/S	−401,559	3,249,178
Mega Invesco Ltd.	−160,302	150,382
Small Propel Bio Management, LLC	−127,766	252,654

Top New Positions

vs Mar 31, 2026

Institution	Acquired	Value (USD)
Mega Jupiter Topco LLC	5,746,289	$76,425,643
Mega T. Rowe Price Investment Management, Inc.	2,511,887	$33,408,097
Small DIADEMA PARTNERS LP	571,107	$7,595,723
Small ACUTA CAPITAL PARTNERS, LLC	507,801	$6,753,753
Small Catalio Capital Management, LP	471,254	$6,267,678

Top Sold-Out Positions

vs Mar 31, 2026

Institution	Previously Held	Last Value (USD)
Mega JANUS HENDERSON GROUP PLC	5,247,151	$48,850,974
Small Peapod Lane Capital LLC	241,480	$2,248,178
Small Exome Asset Management LLC	130,564	$1,215,550
Mid ADAR1 Capital Management, LLC	116,712	$1,086,588
Small Simplify Asset Management Inc.	70,430	$655,703

Filter: New 33 Increased 61 Reduced 38 Sold out 25 Unchanged 28

Showing 100 of 164

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

Jupiter Topco LLC

$76,425,643

9.82%

5,746,289

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	17,262	$229,584	0.3%	Defined
JANUS HENDERSON INVESTORS US LLC	5,729,027	$76,196,059	99.7%	Defined

RA CAPITAL MANAGEMENT, L.P.

$67,162,060

8.63%

5,049,779

BlackRock, Inc.

$60,462,581

7.77%

4,546,059

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	80,761	$1,074,121	1.8%	Sole
BlackRock Asset Management Canada Ltd	328	$4,362	0.0%	Sole
BlackRock Investment Management, LLC	148,261	$1,971,871	3.3%	Sole
BLACKROCK ADVISORS LLC	474,983	$6,317,273	10.4%	Sole
BlackRock Fund Advisors	2,212,188	$29,422,100	48.7%	Sole
BlackRock Institutional Trust Company, N.A.	1,524,936	$20,281,648	33.5%	Sole
BlackRock Japan Co. Ltd	2,291	$30,470	0.1%	Sole
BlackRock Investment Management (UK) Ltd.	20,630	$274,379	0.5%	Sole
BlackRock Asset Management Ireland Ltd	41,354	$550,008	0.9%	Sole
Aperio Group, LLC	40,327	$536,349	0.9%	Sole

GOLDMAN SACHS GROUP INC

$56,640,284

7.28%

4,258,668

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	3,837,410	$51,037,553	90.1%	Defined
Goldman Sachs Trust Company, N.A.	421,258	$5,602,731	9.9%	Defined

NOVO HOLDINGS A/S

$43,214,067

5.55%

3,249,178

STATE STREET CORP

$38,717,749

4.97%

2,911,109

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	2,911,109	$38,717,749	100.0%	Defined

BVF INC/IL

$38,294,543

4.92%

2,879,289

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BVF PARTNERS L P/IL	2,879,289	$38,294,543	100.0%	Defined

T. Rowe Price Investment Management, Inc.

$33,408,097

4.29%

2,511,887

ADAGE CAPITAL PARTNERS GP, L.L.C.

$29,914,399

3.84%

2,249,203

Trails Edge Capital Partners, LP

$29,631,296

3.81%

2,227,917

VANGUARD CAPITAL MANAGEMENT LLC

$27,917,431

3.59%

2,099,055

ARMISTICE CAPITAL, LLC

$26,600,000

3.42%

2,000,000

GEODE CAPITAL MANAGEMENT, LLC

$15,008,597

1.93%

1,128,466

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	155,949	$2,074,121	13.8%	Defined
Held directly	972,517	$12,934,476	86.2%	Defined

BANK OF AMERICA CORP /DE/

$14,904,963

1.91%

1,120,674

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	604,436	$8,038,998	53.9%	Defined
MERRILL LYNCH, PIERCE, FENNER & SMITH INC.	482,511	$6,417,396	43.1%	Defined
BOFA SECURITIES, INC.	33,727	$448,569	3.0%	Defined

Evergreen Quality Fund GP, Ltd.

$12,654,604

1.63%

951,474

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
HHLR ADVISORS, LTD.	951,474	$12,654,604	100.0%	Other

Parkman Healthcare Partners LLC

$12,610,847

1.62%

948,184

DIMENSIONAL FUND ADVISORS LP

$9,720,396

1.25%

730,857

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	6,190	$82,327	0.8%	Defined
DFA Australia Ltd.	26,673	$354,750	3.6%	Defined
Dimensional Ireland Ltd	31,772	$422,567	4.3%	Defined
Held directly	666,222	$8,860,752	91.2%	Sole

DEUTSCHE BANK AG\

$9,455,528

1.21%

710,942

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DWS Investment GmbH	198,000	$2,633,400	27.9%	Defined
DWS Investments UK Ltd	512,942	$6,822,128	72.1%	Defined

CITADEL ADVISORS LLC

$9,145,039

1.17%

687,597

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	193,994	$2,580,120	28.2%	Defined
Held directly	493,603	$6,564,919	71.8%	Defined

RENAISSANCE TECHNOLOGIES LLC

$8,173,820

1.05%

614,573

VANGUARD PORTFOLIO MANAGEMENT LLC

$7,712,709

0.99%

579,903

DIADEMA PARTNERS LP

$7,595,723

0.98%

571,107

NORTHERN TRUST CORP

$7,486,955

0.96%

562,929

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	555,729	$7,391,195	98.7%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	7,200	$95,760	1.3%	Other

ACUTA CAPITAL PARTNERS, LLC

$6,753,753

0.87%

507,801

Nuveen, LLC

$6,690,843

0.86%

503,071

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	12,693	$168,816	2.5%	Defined
TEACHERS ADVISORS, LLC joint	490,378	$6,522,027	97.5%	Defined

MORGAN STANLEY

$6,690,163

0.86%

503,020

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	7	$93	0.0%	Defined
MORGAN STANLEY & CO. LLC	210,107	$2,794,423	41.8%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	115,573	$1,537,120	23.0%	Defined
MORGAN STANLEY SMITH BARNEY LLC	88,488	$1,176,890	17.6%	Defined
Morgan Stanley Bank, N.A.	678	$9,017	0.1%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	88,167	$1,172,620	17.5%	Defined

RTW INVESTMENTS, LP

$6,327,302

0.81%

475,737

Catalio Capital Management, LP

$6,267,678

0.81%

471,254

Schonfeld Strategic Advisors LLC

$5,419,351

0.70%

407,470

JPMORGAN CHASE & CO

$5,348,420

0.69%

402,137

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	29,714	$395,196	7.4%	Defined
J.P. Morgan Investment Management Inc.	126,662	$1,684,604	31.5%	Defined
J.P. Morgan Securities LLC	4,663	$62,017	1.2%	Defined
JPMorgan Asset Management (UK) Ltd	241,098	$3,206,603	60.0%	Defined

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$4,512,357

0.58%

339,275

Opaleye Management Inc.

$4,448,012

0.57%

334,437

VANGUARD FIDUCIARY TRUST CO

$3,673,965

0.47%

276,238

Burkehill Global Management, LP

$3,657,500

0.47%

275,000

UBS Group AG

$3,413,750

0.44%

256,673

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	11,456	$152,364	4.5%	Defined
UBS AG	245,217	$3,261,386	95.5%	Defined

Propel Bio Management, LLC

$3,360,298

0.43%

252,654

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Simplify Asset Management Inc.	252,654	$3,360,298	100.0%	Defined

RAYMOND JAMES FINANCIAL INC

$3,223,240

0.41%

242,349

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	221,063	$2,940,137	91.2%	Defined
Raymond James Financial Services Advisors, Inc.	21,286	$283,103	8.8%	Defined

FEDERATED HERMES, INC.

$2,958,132

0.38%

222,416

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FEDERATED EQUITY MANAGEMENT CO OF PENNSYLVANIA	478	$6,357	0.2%	Defined
FEDERATED MDTA LLC	221,938	$2,951,775	99.8%	Defined

ArrowMark Colorado Holdings LLC

$2,698,091

0.35%

202,864

CITADEL ADVISORS LLC

$2,679,950

0.34%

201,500

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	201,500	$2,679,950	100.0%	Defined

Woodline Partners LP

$2,660,013

0.34%

200,001

Brevan Howard Capital Management LP

$2,352,105

0.30%

176,850

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Brevan Howard Investment Products Ltd	176,850	$2,352,105	100.0%	Defined

ACADIAN ASSET MANAGEMENT LLC

$2,175,148

0.28%

163,545

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Acadian Inc	163,545	$2,175,148	100.0%	Defined

Bank of New York Mellon Corp

$2,026,080

0.26%

152,337

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Investments Corporation	63,677	$846,904	41.8%	Defined
The Bank of New York Mellon	83,659	$1,112,664	54.9%	Defined
BNY Mellon, NA joint	5,000	$66,499	3.3%	Defined
Held directly	1	$13	0.0%	Defined

Invesco Ltd.

$2,000,079

0.26%

150,382

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	6,531	$86,862	4.3%	Defined
Invesco Trust Co joint	18,186	$241,873	12.1%	Defined
Invesco Capital Management LLC	125,665	$1,671,344	83.6%	Defined

Perigon Wealth Management, LLC

$1,944,260

0.25%

146,185

Persistent Asset Partners Ltd

$1,936,280

0.25%

145,585

VOYA INVESTMENT MANAGEMENT LLC

$1,864,659

0.24%

140,200

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	140,200	$1,864,659	100.0%	Defined

WELLINGTON MANAGEMENT GROUP LLP

$1,708,570

0.22%

128,464

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wellington Management Company LLP	40,804	$542,693	31.8%	Defined
Wellington Trust Company, NA joint	81,849	$1,088,591	63.7%	Defined
Wellington Management Australia Pty Ltd	5,811	$77,286	4.5%	Defined

FMR LLC

$1,706,602

0.22%

128,316

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	128,281	$1,706,137	100.0%	Defined
STRATEGIC ADVISERS LLC	35	$465	0.0%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$1,627,920

0.21%

122,400

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	122,400	$1,627,920	100.0%	Other

AMERICAN CENTURY COMPANIES INC

$1,516,200

0.19%

114,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AMERICAN CENTURY INVESTMENT MANAGEMENT INC	114,000	$1,516,200	100.0%	Defined

Susquehanna Portfolio Strategies, LLC

$1,422,235

0.18%

106,935

Greenland Capital Management LP

$1,330,000

0.17%

100,000

Privium Fund Management B.V.

$1,198,463

0.15%

90,110

Assenagon Asset Management S.A.

$1,108,328

0.14%

83,333

Greenland Capital Management LP

$961,018

0.12%

72,257

Caption Management, LLC

$931,000

0.12%

70,000

TWO SIGMA INVESTMENTS, LP

$924,097

0.12%

69,481

CITIGROUP INC

$920,519

0.12%

69,212

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	69,212	$920,519	100.0%	Defined

Man Group plc

$909,041

0.12%

68,349

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Numeric Investors LLC	68,349	$909,041	100.0%	Defined

Engineers Gate Manager LP

$907,139

0.12%

68,206

XTX Topco Ltd

$892,722

0.11%

67,122

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	67,122	$892,722	100.0%	Defined

SG Americas Securities, LLC

$883,652

0.11%

66,440

Fisher Asset Management, LLC

$874,488

0.11%

65,751

RHUMBLINE ADVISERS

$793,185

0.10%

59,638

Quantinno Capital Management LP

$772,490

0.10%

58,082

STATE OF WISCONSIN INVESTMENT BOARD

$748,151

0.10%

56,252

ALLIANCEBERNSTEIN L.P.

$741,342

0.10%

55,740

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	55,740	$741,342	100.0%	Defined

PRICE T ROWE ASSOCIATES INC /MD/

$729,757

0.09%

54,869

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	54,869	$729,757	100.0%	Defined

ALPS ADVISORS INC

$705,644

0.09%

53,056

TD Asset Management Inc

$684,710

0.09%

51,482

AQR CAPITAL MANAGEMENT LLC

$668,735

0.09%

50,281

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RF&L WEALTH MANAGEMENT, LLC	27	$359	0.1%	Other
Arax Advisory Partners	2,337	$31,082	4.6%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	256	$3,404	0.5%	Other
Stokes Family Office, LLC	468	$6,224	0.9%	Other
Held directly	47,193	$627,666	93.9%	Defined

Birchview Capital, LP

$645,116

0.08%

48,505

PANAGORA ASSET MANAGEMENT INC

$624,860

0.08%

46,982

Rafferty Asset Management, LLC

$594,696

0.08%

44,714

D. E. Shaw & Co., Inc.

$589,908

0.08%

44,354

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	22,000	$292,600	49.6%	Defined
D. E. Shaw Investment Management, L.L.C. joint	22,354	$297,308	50.4%	Defined

WELLS FARGO & COMPANY/MN

$555,035

0.07%

41,732

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	38,382	$510,480	92.0%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	1,600	$21,280	3.8%	Defined
WELLS FARGO CLEARING SERVICES, LLC	1,750	$23,275	4.2%	Defined

Allspring Global Investments Holdings, LLC

$494,759

0.06%

37,200

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Allspring Funds Management, LLC reporting on behalf of Morgan Stanley Smith Barney, LLC	139	$1,848	0.4%	Defined
Allspring Global Investments, LLC reporting on behalf of UBS ASSET MANAGEMENT AMERICAS INC	37,061	$492,911	99.6%	Defined

Virtus Investment Advisers, LLC

$492,831

0.06%

37,055

COMMONWEALTH EQUITY SERVICES, LLC

$380,034

0.05%

28,574

Creative Planning

$376,669

0.05%

28,321

OMERS ADMINISTRATION Corp

$369,740

0.05%

27,800

Mitsubishi UFJ Asset Management Co., Ltd.

$354,365

0.05%

26,644

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MITSUBISHI UFJ FINANCIAL GROUP INC	26,644	$354,365	100.0%	Defined

MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.

$351,704

0.05%

26,444

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Global X Management CO LLC joint	21,109	$280,749	79.8%	Defined
Global X Management (AUS) Ltd joint	5,335	$70,955	20.2%	Defined

MetLife Investment Management, LLC

$346,558

0.04%

26,057

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
METLIFE INC	26,057	$346,558	100.0%	Defined

Squarepoint Ops LLC

$340,160

0.04%

25,576

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Squarepoint Capital LLP	25,576	$340,160	100.0%	Defined

TRUIST FINANCIAL CORP

$332,500

0.04%

25,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Truist Advisory Services, Inc.	25,000	$332,500	100.0%	Defined

BRIDGEWAY CAPITAL MANAGEMENT, LLC

$332,500

0.04%

25,000

Peloton Wealth Strategists

$332,500

0.04%

25,000

CANADA PENSION PLAN INVESTMENT BOARD

$328,510

0.04%

24,700

Kera Capital Partners, Inc.

$320,104

0.04%

24,068

BARCLAYS PLC

$316,871

0.04%

23,825

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	22,496	$299,196	94.4%	Defined
BARCLAYS CAPITAL INC.	1,329	$17,675	5.6%	Defined

Rockefeller Capital Management L.P.

$315,594

0.04%

23,729

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ROCKEFELLER & CO. LLC	150	$1,995	0.6%	Defined
ROCKEFELLER TRUST COMPANY, N.A.	203	$2,699	0.9%	Defined
ROCKEFELLER FINANCIAL LLC	23,376	$310,900	98.5%	Defined

Jain Global LLC

$313,042

0.04%

23,537

Russell Investments Group, Ltd.

$312,004

0.04%

23,459

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC	23,459	$312,004	100.0%	Defined

PROFUND ADVISORS LLC

$301,896

0.04%

22,699

MARSHALL WACE, LLP

$283,848

0.04%

21,342

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marshall Wace North America L.P. joint	21,342	$283,848	100.0%	Other

Rangeley Capital, LLC

$278,834

0.04%

20,965

Headlands Technologies LLC

$255,865

0.03%

19,238

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI INVESTMENTS MANAGEMENT CORP	48	$638	0.3%	Sole
SEI TRUST CO	16,616	$220,992	99.7%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Aperio Group, LLC	16,359	$217,574	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jane Street Capital, LLC	15,600	$207,480	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Manager 1	14,899	$198,156	100.0%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Capital	13,831	$183,952	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MANULIFE FINANCIAL CORP joint	12,489	$166,103	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	12,161	$161,741	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	12,100	$160,930	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Latour Trading LLC (Latour)	6,110	$81,263	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	4,448	$59,158	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
EATON VANCE MANAGEMENT	4,159	$55,314	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RBC CMA LTD.	1,391	$18,500	72.8%	Defined
RBC Capital Markets, LLC	520	$6,916	27.2%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Optiver VOF	1,782	$23,700	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PNC Bank, National Association	1,482	$19,710	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	691	$9,190	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
US BANK NATIONAL ASSOCIATION	58	$771	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BMSS Wesson Wealth Solutions, LLC	6	$79	14.9%	Sole
Held directly	34	$451	85.1%	Sole

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
OSAIC WEALTH, INC.	35	$465	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Deka Vermoegensmanagement GmbH	30,000	$256	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	17,187	$160,010	0.3%	Defined
JANUS HENDERSON INVESTORS US LLC	5,229,964	$48,690,964	99.7%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Quadrature US Inc.	32,524	$302,798	100.0%	Other

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Virtu Americas LLC	26,368	$245,486	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fifth Third Bank, National Association	35	$325	100.0%	Defined

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Institutional Asset Management Trust Co	8	$74	100.0%	Sole

Key facts CIK 1650648 CUSIP 35104E100 13F (30d) 156 filings 155 filers Visit website Investor relations