VRDN · Viridian Therapeutics, Inc.\DE · Holdings

$22.48 +1.68 (+8.08%)

Russell 2000 member Rebalance outlook locked

Market Cap

$2.46B

113.29M

Volume · Oct 20 1.97M Avg daily vol (3M) 1.79M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Use institutional holdings through the 13F API or MCP → See every holder of VRDN: 13F, 13D and 13G, insiders and funds →

Institutional holdings are being reconciled with SEC filings. Holder totals and changes may be incomplete until reconciliation finishes.

The Sep 30, 2026 filing window is still open — showing the latest known positions. Funds that have not filed yet carry their prior-quarter positions; a fund that filed without this stock counts as sold out. Each fund's Change is measured against its own previous reported quarter, shown in the Reported column. Pick a completed quarter in the Quarter selector below to put every fund on one quarter and see the stock-wide activity breakdown.

Quarter:

Holders: 213

Total Value: $2,507,399,856

Total Shares: 136,463,352

Showing 1–100 of 220 holdings by value · page 1 of 3

Institutional Ownership Trend

24 quarters

Holder Concentration

Across 217 13F filers

Top Holder

12.2%

of institutional value

Top 5

33.0%

of institutional value

Top 10

51.4%

of institutional value

HHI

383

Broadly held

Who Holds VRDN

Last 8 quarters through Sep 30, 2026 (filing window open)

Each kind of 13F filer's share of the common shares 13F filers report. Holder type comes from regulatory records, never the filer's name; style, turnover and horizon are measured from each filer's own 13F book. The quarter selector does not change this card.

Jun 30, 2026, the newest closed quarter: 214 filers, like-for-like change +21.1%; actively managed books +24.1%; index-like books +10.6%.

Holder type	Share	Filers	Change	New	Exited
Registered adviser	51.7%	100	+37.0%	26	34
Exempt adviser	0.1%	4	-70.7%	2	2
Adviser	4.3%	10	+22.3%	2	1
Bank	7.8%	14	+27.2%	0	1
Insurer	0.0%	2	-66.2%	1	1
Benefit plan	0.1%	5	-3.2%	1	0
Non-US	6.2%	7	+3.0%	1	0
Holding company	22.6%	6	+6.8%	0	0
Broker-dealer	0.0%	0	-100.0%	0	1
Unknown	7.2%	66	-4.3%	11	15

Style	Share	Filers	Change	New	Exited
Index-like	20.6%	29	+10.6%	1	1
Systematic-like	6.2%	22	+2.0%	8	11
Concentrated	25.7%	31	+52.4%	6	11
Diversified active	47.5%	132	+15.8%	29	32
Not measured	0.0%	0	—	0	0
Unknown	0.0%	0	—	0	0

Turnover	Share	Filers	Change	New	Exited
Low turnover	53.9%	96	+13.8%	14	14
Medium turnover	32.2%	58	+41.8%	11	19
High turnover	7.0%	38	+3.4%	17	17
Unknown	7.0%	22	+19.6%	2	5
Not measured	0.0%	0	—	0	0

Holding horizon	Share	Filers	Change	New	Exited
Short horizon	8.0%	39	+19.1%	13	14
Medium horizon	24.5%	40	+32.4%	9	11
Long horizon	57.0%	116	+16.4%	14	15
Unknown	10.5%	19	+25.0%	8	15
Not measured	0.0%	0	—	0	0

13F book size	Share	Filers	Change	New	Exited
Under $1B	4.6%	36	+44.2%	13	15
$1B to $10B	33.1%	67	+41.9%	16	18
$10B to $100B	10.6%	58	+7.0%	15	14
$100B or more	51.7%	53	+11.9%	0	8
Not measured	0.0%	0	—	0	0

Client base	Share	Filers	Change	New	Exited
Individuals and high net worth	0.1%	24	+54.4%	10	7
Pooled investment vehicles	32.0%	40	+46.0%	11	17
Registered funds and BDCs	13.9%	14	+41.5%	2	2
Pension and profit sharing plans	0.0%	1	-95.3%	0	1
Insurance companies	0.3%	4	-63.2%	0	0
Governments and sovereign funds	2.3%	5	+41.5%	1	2
Other investment advisers	0.0%	2	+0.7%	0	0
Diversified	3.0%	9	-12.2%	2	5
Other	0.0%	0	—	0	0
Unknown	48.4%	115	+7.6%	18	21

Private funds	Share	Filers	Change	New	Exited
Hedge funds	28.5%	46	+49.7%	14	18
Private equity funds	2.5%	8	-31.7%	1	1
Venture capital funds	0.0%	1	-99.9%	0	1
Real estate funds	0.0%	2	+195.5%	1	2
Securitized asset funds	0.7%	1	+12.6%	0	0
Other private funds	7.2%	8	+51.2%	0	5
Mixed private funds	2.9%	3	+36.3%	1	0
Private funds, type not stated	0.0%	1	-82.6%	0	2
No private funds	10.0%	34	+30.5%	11	7
Unknown	48.2%	110	+8.3%	16	19

Location	Share	Filers	Change	New	Exited
United States	89.1%	171	+22.6%	35	47
Outside the US	10.9%	43	+9.5%	9	8

The Sep 30, 2026 filing window is still open, so its bar carries the Jun 30, 2026 positions of filers that have not filed yet.

How the mix is measured

Shares are 13F common shares restated to today's split basis; options and principal amounts are left out. The change compares only filers with a 13F in both quarters and keeps each filer in its bucket of the later quarter on both sides; a filer that continues under a new CIK is compared through its successor. Holder type comes from each filer's current regulatory profile; client base and private funds come from the Form ADV roster current at each quarter end (rosters from December 2019, so an earlier quarter reads Unknown). Style, turnover, horizon and book size are measured from each filer's 13F book in the newest measured quarter, and Not measured means none was. Actively managed books are those measured systematic-like, concentrated or diversified active.

Excluded 1 holder with no observed 2026-03-31 13F and 1 holder with no observed 2026-06-30 13F. Compared across CIK changes: JANUS HENDERSON GROUP PLC continues as Jupiter Topco LLC.

Conviction and Crowding

214 13F filers, quarter ended Jun 30, 2026

Measured across every 13F filer's common-share book once the quarter's filing window closed. Crowding counts filers with high turnover or that advise private funds on Form ADV.

Conviction holders

3.27% of filers hold it in their top 10

Mar 31, 2026: 6

Holders above 5%

median weight 0.003%

Mar 31, 2026: 1

Crowding

—

holders exceed shares outstanding: 13F filers report more shares than the cover-page count, which predates an issuance, is misstated, or misses shares lent and reported twice

Private-fund advisers

—

of shares outstanding

Institutional ownership

—

holders exceed shares outstanding: 13F filers report more shares than the cover-page count, which predates an issuance, is misstated, or misses shares lent and reported twice

Top 10 holders

51.6%

of 13F shares, Herfindahl 0.039

Active holders

+24.1%

like-for-like share change of active filers

5% stakes on record

1 13D, 6 13G (5 institutional, 1 passive, 0 exempt)

How long holders have held it value-weighted 12.3 quarters

Under 1 year 25.6% 1 to 3 years 18.9% 3 years or more 55.5% Since coverage start 0.000%

Managers in the top fifth by current three-year FundScore alpha were net buyers of $19,760,607 (2 buying, 0 selling).

Estimated Holder Cost Basis

Quarter ended Jun 30, 2026, on that quarter's share basis

Estimated, never filed: each 13F buy is costed at its quarter's average traded price and each sale keeps the average cost, so this is what holders paid on average if they traded evenly through each quarter.

Estimated avg cost

$20.45

estimated per share

Mar 31, 2026: $22.49

Quarter-end price

$18.37

implied by 13F values

Estimated gain or loss

-10.2%

price against the estimated cost

Estimated at a loss

68.6%

of costed 13F value

Estimated cost covers 78.3% of the 13F shares; positions already held when a filer's 13F history starts, or bought in a quarter without a trusted average price, have no estimate.

Shared Holders

Companies VRDN's active holders also own, quarter ended Jun 30, 2026

#	Company	Shared holders	Similarity
1	VERA Vera Therapeutics, Inc.	13	0.353
2	ELVN Enliven Therapeutics, Inc.	14	0.340
3	STRO Sutro Biopharma, Inc.	7	0.333
4	COGT Cogent Biosciences, Inc.	15	0.319
5	PCVX Vaxcyte, Inc.	12	0.307
6	AVTX Avalo Therapeutics, Inc.	11	0.302
7	JBIO Jade Biosciences, Inc.	8	0.286
8	COAG Hemab Therapeutics Holdings, Inc.	6	0.285
9	EWTX Edgewise Therapeutics, Inc.	10	0.282
10	NRIX Nurix Therapeutics, Inc.	10	0.282

Shareholder Targets

13F filers that own its peers but not VRDN, quarter ended Jun 30, 2026

Found through its holder-overlap peers: VERA, ELVN, STRO, COGT, PCVX, AVTX, JBIO, COAG, EWTX, NRIX, CGEM, CLDX, PRQR, IMCR, TYRA, CTMX, TVTX, ORKA, RLAY, DMRA. 589 filers in all.

#	Filer	In peers	Of book	Largest peers
1	RA CAPITAL MANAGEMENT, L.P. Registered adviser, Concentrated	$1,994,868,718 8 held	17.6%	PCVX, TYRA, EWTX
2	ORBIMED ADVISORS LLC Registered adviser, Diversified active	$1,097,685,827 7 held	21.7%	EWTX, ELVN, COGT
3	T. Rowe Price Investment Management, Inc. Adviser, Diversified active	$898,799,680 5 held	0.590%	PCVX, ORKA, RLAY
4	Polar Capital Holdings Plc Registered adviser, Diversified active	$711,846,961 6 held	2.01%	ELVN, COGT, ORKA
5	Commodore Capital LP Registered adviser, Concentrated	$688,134,175 7 held	41.3%	RLAY, ELVN, COGT
6	RTW INVESTMENTS, LP Registered adviser, Diversified active	$589,005,441 5 held	5.03%	ORKA, IMCR, COGT
7	Siren, L.L.C. Unknown, Diversified active	$554,141,255 10 held	11.3%	EWTX, ELVN, COAG
8	SB INVESTMENT ADVISERS (UK) LTD Registered adviser, Concentrated	$522,101,857 1 held	6.81%	RLAY
9	Vestal Point Capital, LP Registered adviser, Diversified active	$484,720,750 5 held	11.3%	ELVN, NRIX, IMCR
10	ADAGE CAPITAL PARTNERS GP, L.L.C. Registered adviser, Diversified active	$441,092,293 13 held	0.620%	TVTX, ELVN, PCVX

Fund Ownership of VRDN by Fund Type

Registered funds' Form N-PORT reports for the quarter ended Jun 30, 2026

Index funds and index ETFs hold 15.15% of shares outstanding.

Fund type	Of shares outstanding	Change	Shares	Funds
Index funds	2.90%	+0.68 pp	3,280,904	62
Index ETFs	12.25%	+2.05 pp	13,878,947	43
Active ETFs	0.81%	+0.39 pp	921,194	16
Active funds	21.78%	+7.38 pp	24,678,894	126
Unclassified	0.52%	+0.52 pp	584,718	6
All registered funds	38.26%	+11.02 pp	43,344,657	253

How fund types are measured

Each fund series counts once, with its latest Form N-PORT report dated inside the quarter; only long share positions count, restated to today's split basis. The fund type is the series' latest Form N-CEN answer on or before the quarter end (the index-fund box and the exchange-traded flag), never its name. Unclassified funds have no usable N-CEN; unit investment trusts are among them, so index ownership is a floor. Funds sit inside their advisers' 13F reports, so these shares overlap the 13F figures and are never added to them.

Holder type: Registered adviser 100 Exempt adviser 3 Adviser 10 Bank 14 Insurer 2 Benefit plan 5 Non-US 7 Holding company 6 Unknown 66

Showing 100 of 220

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

FMR LLC

$305,029,072

12.17%

16,604,740

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC joint	13,715,330	$251,950,612	82.6%	Defined
Fidelity Institutional Asset Management Trust Co	1,250,683	$22,975,046	7.5%	Defined
FIAM LLC	1,638,727	$30,103,414	9.9%	Defined

BlackRock, Inc.

$157,090,828

6.27%

8,551,488

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	83,322	$1,530,625	1.0%	Sole
BlackRock Investment Management (Australia) Ltd	44	$808	0.0%	Sole
BlackRock Asset Management Canada Ltd	27,560	$506,277	0.3%	Sole
BlackRock Investment Management, LLC	306,925	$5,638,212	3.6%	Sole
BLACKROCK ADVISORS LLC	934,950	$17,175,031	10.9%	Sole
BlackRock Fund Advisors	3,839,978	$70,540,395	44.9%	Sole
BlackRock Institutional Trust Company, N.A.	2,944,415	$54,088,903	34.4%	Sole
BlackRock Fund Managers Ltd.	16,010	$294,103	0.2%	Sole
BlackRock Investment Management (UK) Ltd.	92,098	$1,691,840	1.1%	Sole
BlackRock (Netherlands) B.V.	3,986	$73,222	0.0%	Sole
BlackRock Asset Management Ireland Ltd	245,581	$4,511,322	2.9%	Sole
BlackRock Asset Management Schweiz AG	11,397	$209,362	0.1%	Sole
Aperio Group, LLC	45,222	$830,728	0.5%	Sole

Bellevue Group AG

$123,665,167

4.93%

6,731,909

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BB Biotech AG joint	6,717,515	$123,400,750	99.8%	Defined
Bellevue Asset Management AG	14,394	$264,417	0.2%	Defined

Paradigm Biocapital Advisors LP

$123,515,985

4.93%

6,723,788

Kynam Capital Management, LP

$118,557,720

4.73%

6,453,877

STATE STREET CORP

$112,515,809

4.49%

6,124,976

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	6,124,976	$112,515,809	100.0%	Defined

PRICE T ROWE ASSOCIATES INC /MD/

$105,598,291

4.21%

5,748,410

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	133,257	$2,447,931	2.3%	Defined
Held directly	5,615,153	$103,150,360	97.7%	Sole

Fairmount Funds Management LLC

$93,520,347

3.73%

5,090,928

VANGUARD CAPITAL MANAGEMENT LLC

$84,076,918

3.35%

4,576,860

Deep Track Capital, LP

$65,998,633

2.63%

3,592,740

Venrock Adviser, LLC

$59,847,935

2.39%

3,257,917

WELLINGTON MANAGEMENT GROUP LLP

$57,468,007

2.29%

3,128,362

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wellington Management Company LLP	2,140,618	$39,323,152	68.4%	Defined
Wellington Managment Canada LLC	48,358	$888,336	1.5%	Defined
Wellington Management Singapore Pte. Ltd.	67,192	$1,234,317	2.1%	Defined
Wellington Management International Ltd	56,505	$1,037,996	1.8%	Defined
Wellington Trust Company, NA joint	802,593	$14,743,633	25.7%	Defined
Wellington Management Australia Pty Ltd	11,147	$204,770	0.4%	Defined
Wellington Management Europe GmbH	1,949	$35,803	0.1%	Defined

VANGUARD PORTFOLIO MANAGEMENT LLC

$50,569,431

2.02%

2,752,827

GEODE CAPITAL MANAGEMENT, LLC

$45,959,333

1.83%

2,501,869

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	359,698	$6,607,652	14.4%	Defined
Held directly	2,142,171	$39,351,681	85.6%	Defined

Avoro Capital Advisors LLC

$45,925,000

1.83%

2,500,000

BAKER BROS. ADVISORS LP

$43,776,169

1.75%

2,383,025

Point72 Asset Management, L.P.

$43,044,730

1.72%

2,343,208

Caligan Partners LP

$40,303,430

1.61%

2,193,981

NOVO HOLDINGS A/S

$39,219,950

1.56%

2,135,000

TANG CAPITAL MANAGEMENT LLC

$32,930,539

1.31%

1,792,626

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
TANG KEVIN	1,792,626	$32,930,539	100.0%	Other

Fisher Asset Management, LLC

$32,163,114

1.28%

1,750,850

Candriam S.C.A.

$31,370,412

1.25%

1,707,698

GOLDMAN SACHS GROUP INC

$31,069,253

1.24%

1,691,304

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	1,110,431	$20,398,617	65.7%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	566,327	$10,403,426	33.5%	Defined
Goldman Sachs Trust Company, N.A. joint	14,546	$267,210	0.9%	Other

First Turn Management, LLC

$30,677,900

1.22%

1,670,000

DIMENSIONAL FUND ADVISORS LP

$29,603,639

1.18%

1,611,521

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	9,917	$182,175	0.6%	Defined
DFA Australia Ltd.	27,340	$502,235	1.7%	Defined
Dimensional Fund Advisors Pte. Ltd.	671	$12,326	0.0%	Defined
Dimensional Ireland Ltd	67,600	$1,241,812	4.2%	Defined
Held directly	1,505,993	$27,665,091	93.5%	Sole

UBS Group AG

$26,247,844

1.05%

1,428,843

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	904	$16,606	0.1%	Defined
UBS AG	939,101	$17,251,285	65.7%	Defined
UBS Switzerland AG	485,851	$8,925,082	34.0%	Defined
UBS Securities LLC	2,987	$54,871	0.2%	Defined

ALLIANCEBERNSTEIN L.P.

$25,564,408

1.02%

1,391,639

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	1,391,639	$25,564,408	100.0%	Defined

Parkman Healthcare Partners LLC

$25,306,071

1.01%

1,377,576

Balyasny Asset Management L.P.

$23,511,009

0.94%

1,279,859

D. E. Shaw & Co., Inc.

$21,717,418

0.87%

1,182,222

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	400,438	$7,356,046	33.9%	Defined
D. E. Shaw Investment Management, L.L.C. joint	781,784	$14,361,372	66.1%	Defined

RICE HALL JAMES & ASSOCIATES, LLC

$21,601,393

0.86%

1,175,906

ARROWSTREET CAPITAL, LIMITED PARTNERSHIP

$20,435,026

0.81%

1,112,413

BNP PARIBAS ARBITRAGE, SA

$18,592,975

0.74%

1,012,138

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$18,368,435

0.73%

999,915

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	6,066	$111,432	0.6%	Defined
UBS Asset Management Switzerland AG	514,115	$9,444,291	51.4%	Defined
UBS Asset Management (UK) Ltd.	29,360	$539,343	2.9%	Defined
Held directly	450,374	$8,273,369	45.0%	Defined

Elmind Capital, LP

$18,093,182

0.72%

984,931

BANK OF AMERICA CORP /DE/

$17,347,617

0.69%

944,345

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	872,368	$16,025,400	92.4%	Defined
BOFA SECURITIES, INC.	71,977	$1,322,217	7.6%	Defined

NORTHERN TRUST CORP

$15,862,327

0.63%

863,491

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	813,646	$14,946,676	94.2%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	35,649	$654,871	4.1%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	14,196	$260,780	1.6%	Other

Eversept Partners, LP

$15,268,078

0.61%

831,142

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$14,972,045

0.60%

815,027

Schonfeld Strategic Advisors LLC

$14,576,741

0.58%

793,508

RENAISSANCE TECHNOLOGIES LLC

$14,119,182

0.56%

768,600

MORGAN STANLEY

$13,388,365

0.53%

728,817

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	1,580	$29,024	0.2%	Defined
MORGAN STANLEY & CO. LLC	45,170	$829,772	6.2%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	480,799	$8,832,277	66.0%	Defined
MORGAN STANLEY SMITH BARNEY LLC	4,311	$79,192	0.6%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	196,957	$3,618,100	27.0%	Defined

Woodline Partners LP

$12,876,065

0.51%

700,929

VANGUARD FIDUCIARY TRUST CO

$12,079,891

0.48%

657,588

JPMORGAN CHASE & CO

$11,755,493

0.47%

639,929

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JPMorgan Chase Bank, N.A.	37,448	$687,919	5.9%	Defined
J.P. Morgan Investment Management Inc.	31,197	$573,088	4.9%	Defined
J.P. Morgan Securities LLC	26	$477	0.0%	Defined
J.P. Morgan Securities plc	636	$11,683	0.1%	Defined
JPMorgan Asset Management (UK) Ltd	570,622	$10,482,326	89.2%	Defined

PERCEPTIVE ADVISORS LLC

$10,690,972

0.43%

581,980

BOOTHBAY FUND MANAGEMENT, LLC

$9,939,235

0.40%

541,058

EMERALD ADVISERS, LLC

$9,870,715

0.39%

537,328

DEUTSCHE BANK AG\

$8,870,412

0.35%

482,875

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DWS Investment GmbH	140,883	$2,588,020	29.2%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC	185	$3,398	0.0%	Defined
DWS Investments UK Ltd	341,807	$6,278,994	70.8%	Defined

ALGERT GLOBAL LLC

$8,858,693

0.35%

482,237

Ikarian Capital, LLC

$7,807,250

0.31%

425,000

Yiheng Capital Management, L.P.

$7,356,523

0.29%

400,464

Jump Financial, LLC

$7,315,870

0.29%

398,251

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jump Trading Holdings, LLC joint	2,751	$50,535	0.7%	Defined
Leap GP LLC joint	395,500	$7,265,335	99.3%	Defined

EMERALD MUTUAL FUND ADVISERS TRUST

$7,266,180

0.29%

395,546

PANAGORA ASSET MANAGEMENT INC

$5,963,967

0.24%

324,658

Hennion & Walsh Asset Management, Inc.

$5,245,611

0.21%

254,765

XTX Topco Ltd

$5,054,781

0.20%

275,165

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	275,165	$5,054,781	100.0%	Defined

Bank of New York Mellon Corp

$4,924,260

0.20%

268,060

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Investments Corporation	97,281	$1,787,051	36.3%	Defined
The Bank of New York Mellon	170,048	$3,123,781	63.4%	Defined
BNY Mellon, NA joint	730	$13,410	0.3%	Defined
Held directly	1	$18	0.0%	Defined

Blue Owl Capital Holdings LP

$4,776,200

0.19%

260,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Blue Owl Healthcare Opportunities Advisors LLC	260,000	$4,776,200	100.0%	Defined

Affinity Asset Advisors, LLC

$4,592,500

0.18%

250,000

VOYA INVESTMENT MANAGEMENT LLC

$4,588,145

0.18%

249,763

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	249,763	$4,588,145	100.0%	Defined

TD Asset Management Inc

$4,328,082

0.17%

235,606

Nuveen, LLC

$3,659,854

0.15%

199,230

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	7,988	$146,739	4.0%	Defined
TEACHERS ADVISORS, LLC joint	191,242	$3,513,115	96.0%	Defined

CAPITAL FUND MANAGEMENT S.A.

$3,634,467

0.14%

197,848

FIRST TRUST ADVISORS LP

$3,329,268

0.13%

181,234

Knott David M Jr

$3,324,970

0.13%

181,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dorset Partners LLC	181,000	$3,324,970	100.0%	Defined

MANUFACTURERS LIFE INSURANCE COMPANY, THE

$3,312,992

0.13%

180,348

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MANULIFE FINANCIAL CORP joint	180,348	$3,312,992	100.0%	Defined

Squarepoint Ops LLC

$3,250,663

0.13%

176,955

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Squarepoint Capital LLP	176,955	$3,250,663	100.0%	Defined

Privium Fund Management B.V.

$3,187,488

0.13%

173,516

HighVista Strategies LLC

$2,937,087

0.12%

159,885

Swiss National Bank

$2,849,187

0.11%

155,100

CITIGROUP INC

$2,579,422

0.10%

140,415

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	140,415	$2,579,422	100.0%	Defined

CITADEL ADVISORS LLC

$2,555,267

0.10%

139,100

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	139,100	$2,555,267	100.0%	Defined

F/m Investments LLC

$2,475,559

0.10%

134,761

BNP PARIBAS ASSET MANAGEMENT Holding S.A.

$2,469,937

0.10%

134,455

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNP PARIBAS ASSET MANAGEMENT Europe	1,253	$23,017	0.9%	Defined
AXA INVESTMENT MANAGERS UK LIMITED	133,202	$2,446,920	99.1%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$2,393,317

0.10%

130,284

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	130,284	$2,393,317	100.0%	Other

AQR CAPITAL MANAGEMENT LLC

$2,338,517

0.09%

127,301

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RF&L WEALTH MANAGEMENT, LLC	3	$55	0.0%	Other
Arax Advisory Partners	218	$4,004	0.2%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	156	$2,865	0.1%	Other
Stokes Family Office, LLC	1	$18	0.0%	Other
Held directly	126,923	$2,331,575	99.7%	Defined

KVP Capital Advisors, LP

$2,296,250

0.09%

125,000

Lighthouse Investment Partners, LLC

$2,204,400

0.09%

120,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
North Rock Capital Management, LLC	120,000	$2,204,400	100.0%	Defined

RHUMBLINE ADVISERS

$2,163,012

0.09%

117,747

WELLS FARGO & COMPANY/MN

$2,122,889

0.08%

115,563

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	76,359	$1,402,714	66.1%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	8,215	$150,909	7.1%	Defined
WELLS FARGO CLEARING SERVICES, LLC	30,929	$568,165	26.8%	Defined
Wells Fargo Securities, LLC	1	$18	0.0%	Defined
WELLS FARGO INVESTMENT INSTITUTE, INC.	59	$1,083	0.1%	Defined

Russell Investments Group, Ltd.

$2,117,985

0.08%

115,296

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC joint	45,947	$844,045	39.9%	Defined
Russell Investments Capital, LLC joint	54,134	$994,441	47.0%	Defined
Russell Investments Trust Company joint	15,215	$279,499	13.2%	Other

ALPS ADVISORS INC

$2,049,577

0.08%

111,572

SG Americas Securities, LLC

$2,003,671

0.08%

109,073

Rafferty Asset Management, LLC

$1,997,719

0.08%

108,749

J. Safra Sarasin Holding AG

$1,981,002

0.08%

107,839

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
J. Safra Sarasin Fund Management (Luxembourg) S.A.	107,839	$1,981,002	100.0%	Defined

JACOBS LEVY EQUITY MANAGEMENT, INC

$1,862,975

0.07%

101,414

Invesco Ltd.

$1,853,439

0.07%

100,895

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	12,084	$221,982	12.0%	Defined
Invesco Trust Co joint	35,693	$655,680	35.4%	Defined
Invesco Capital Management LLC	53,118	$975,777	52.6%	Defined

NOMURA HOLDINGS INC

$1,840,674

0.07%

100,200

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NOMURA SECURITIES INTERNATIONAL INC	100,200	$1,840,674	100.0%	Defined

DIADEMA PARTNERS LP

$1,837,000

0.07%

100,000

Voleon Capital Management LP

$1,792,875

0.07%

97,598

Merck & Co., Inc.

$1,716,015

0.07%

93,414

FIDEURAM - INTESA SANPAOLO PRIVATE BANKING S.P.A.

$1,692,758

0.07%

92,148

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Intesa Sanpaolo S.p.A.	92,148	$1,692,758	100.0%	Defined

CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM

$1,679,054

0.07%

91,402

Caption Management, LLC

$1,580,554

0.06%

86,040

VICTORY CAPITAL MANAGEMENT INC

$1,499,524

0.06%

81,629

BARCLAYS PLC

$1,495,849

0.06%

81,429

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	73,480	$1,349,827	90.2%	Defined
BARCLAYS CAPITAL INC.	7,822	$143,690	9.6%	Defined
BARCLAYS CAPITAL SECURITIES LTD	127	$2,332	0.2%	Defined

Aquatic Capital Management LLC

$1,466,752

0.06%

79,845

Knights of Columbus Asset Advisors LLC

$1,296,058

0.05%

70,553

J. Goldman & Co LP

$1,239,975

0.05%

67,500

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Key facts CIK 1590750 CUSIP 92790C104 13F (30d) 7 filings 7 filers Visit website Investor relations