PTGX · Protagonist Therapeutics, Inc · Holdings

$143.73 +3.82 (+2.73%) At close · Oct 9

S&P 600 member Russell 2000 member Rebalance outlook locked

Market Cap

$9.30B

64.71M

Volume · Oct 9 763.1K Avg daily vol (3M) 625.26K

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Use institutional holdings through the 13F API or MCP →

Some SEC filings for this period have not finished importing. Holder totals and changes may be incomplete; these filings remain queued for recovery.

The Sep 30, 2026 filing window is still open — showing the latest known positions. Funds that have not filed yet carry their prior-quarter positions; a fund that filed without this stock counts as sold out. Each fund's Change is measured against its own previous reported quarter, shown in the Reported column. Pick a completed quarter in the Quarter selector below to put every fund on one quarter and see the stock-wide activity breakdown.

Quarter:

Holders: 347

Total Value: $9,367,115,066

Total Shares: 76,427,015

Showing 1–100 of 358 holdings by value · page 1 of 4

Institutional Ownership Trend

27 quarters

Superinvestors Holding PTGX

All superinvestors →

Stanley Druckenmiller $25,606,962 Joel Greenblatt $2,287,710 Ray Dalio $1,136,194

Holder Concentration

Across 346 13F filers

Top Holder

13.5%

of institutional value

Top 5

37.6%

of institutional value

Top 10

56.4%

of institutional value

HHI

458

Broadly held

Showing 100 of 358

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

BlackRock, Inc.

$1,263,534,651

13.49%

10,307,837

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	66,011	$8,091,628	0.6%	Sole
BlackRock Investment Management (Australia) Ltd	228	$27,948	0.0%	Sole
BlackRock Asset Management Canada Ltd	8,436	$1,034,084	0.1%	Sole
BlackRock Investment Management, LLC	214,378	$26,278,455	2.1%	Sole
BLACKROCK ADVISORS LLC	664,115	$81,407,216	6.4%	Sole
BlackRock Fund Advisors	7,002,179	$858,327,101	67.9%	Sole
BlackRock Institutional Trust Company, N.A.	1,872,422	$229,521,488	18.2%	Sole
BlackRock Japan Co. Ltd	1,363	$167,076	0.0%	Sole
BlackRock Fund Managers Ltd.	9,511	$1,165,858	0.1%	Sole
BlackRock Investment Management (UK) Ltd.	51,604	$6,325,618	0.5%	Sole
BlackRock (Netherlands) B.V.	2,183	$267,592	0.0%	Sole
BlackRock Asset Management Ireland Ltd	269,307	$33,011,652	2.6%	Sole
BlackRock (Luxembourg) S.A.	8,542	$1,047,078	0.1%	Sole
BlackRock Life Ltd	79	$9,683	0.0%	Sole
BlackRock Asset Management Schweiz AG	8,703	$1,066,813	0.1%	Sole
Aperio Group, LLC	128,743	$15,781,316	1.2%	Sole
SpiderRock Advisors, LLC	33	$4,045	0.0%	Sole

FARALLON CAPITAL MANAGEMENT LLC

$778,248,897

8.31%

6,348,906

RTW INVESTMENTS, LP

$620,398,831

6.62%

5,061,175

VANGUARD PORTFOLIO MANAGEMENT LLC

$444,879,348

4.75%

3,629,298

WELLINGTON MANAGEMENT GROUP LLP

$418,829,010

4.47%

3,416,781

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wellington Management Company LLP	2,849,468	$349,287,787	83.4%	Defined
Wellington Managment Canada LLC	52,198	$6,398,430	1.5%	Defined
Wellington Management Singapore Pte. Ltd.	40,283	$4,937,890	1.2%	Defined
Wellington Management International Ltd	118,572	$14,534,555	3.5%	Defined
Wellington Trust Company, NA joint	326,043	$39,966,350	9.5%	Defined
Wellington Management Australia Pty Ltd	8,312	$1,018,884	0.2%	Defined
Wellington Management Europe GmbH	21,905	$2,685,114	0.6%	Defined

UBS Group AG

$416,676,508

4.45%

3,399,221

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	867	$106,276	0.0%	Defined
UBS AG	3,332,313	$408,474,927	98.0%	Defined
UBS Switzerland AG	66,041	$8,095,305	1.9%	Defined

STATE STREET CORP

$407,156,089

4.35%

3,321,554

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	3,321,554	$407,156,089	100.0%	Defined

VANGUARD CAPITAL MANAGEMENT LLC

$349,769,036

3.73%

2,853,394

JOHNSON & JOHNSON

$300,220,852

3.21%

2,449,183

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Johnson & Johnson Innovation - JJDC, Inc.	2,449,183	$300,220,852	100.0%	Defined

Invesco Ltd.

$286,918,957

3.06%

2,340,667

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	1,624,571	$199,139,911	69.4%	Defined
Invesco Trust Co joint	24,505	$3,003,822	1.0%	Defined
Invesco Investment Advisers LLC	2,920	$357,933	0.1%	Defined
Invesco Capital Management LLC	688,671	$84,417,291	29.4%	Defined

PRICE T ROWE ASSOCIATES INC /MD/

$263,505,567

2.81%

2,149,662

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	83,246	$10,204,294	3.9%	Defined
Held directly	2,066,416	$253,301,273	96.1%	Sole

Jupiter Topco LLC

$261,068,431

2.79%

2,129,780

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	119,259	$14,618,768	5.6%	Defined
JANUS HENDERSON INVESTORS US LLC	2,001,105	$245,295,450	94.0%	Defined
Held directly	9,416	$1,154,213	0.4%	Defined

GEODE CAPITAL MANAGEMENT, LLC

$213,437,399

2.28%

1,741,209

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	224,233	$27,486,481	12.9%	Defined
Held directly	1,516,976	$185,950,918	87.1%	Defined

Woodline Partners LP

$173,349,326

1.85%

1,414,173

BANK OF AMERICA CORP /DE/

$162,356,963

1.73%

1,324,498

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	1,146,579	$140,547,653	86.6%	Defined
BOFA SECURITIES, INC.	42,490	$5,208,424	3.2%	Defined
MANAGED ACCOUNT ADVISORS LLC joint	135,429	$16,600,886	10.2%	Defined

JENNISON ASSOCIATES LLC

$141,173,056

1.51%

1,151,681

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Prudential Financial Inc	920,855	$112,878,405	80.0%	Defined
Prudential Trust Company	230,826	$28,294,651	20.0%	Other

PERCEPTIVE ADVISORS LLC

$120,358,727

1.28%

981,879

ADAR1 Capital Management, LLC

$117,782,586

1.26%

960,863

MORGAN STANLEY

$111,056,494

1.19%

905,992

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	310	$37,999	0.0%	Defined
MORGAN STANLEY & CO. LLC	155,165	$19,020,125	17.1%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	495,117	$60,691,441	54.6%	Defined
MORGAN STANLEY INVESTMENT MANAGEMENT INC	508	$62,270	0.1%	Defined
MORGAN STANLEY SMITH BARNEY LLC	31,468	$3,857,347	3.5%	Defined
Morgan Stanley Strategic Investments, Inc	1	$122	0.0%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	217,136	$26,616,530	24.0%	Defined
EATON VANCE MANAGEMENT	6,287	$770,660	0.7%	Defined

Alyeska Investment Group, L.P.

$109,010,148

1.16%

889,298

Hood River Capital Management LLC

$108,206,024

1.16%

882,738

GOLDMAN SACHS GROUP INC

$99,639,397

1.06%

812,852

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	565,028	$69,261,132	69.5%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	244,405	$29,959,164	30.1%	Defined
Goldman Sachs Trust Company, N.A.	3,419	$419,101	0.4%	Defined

Driehaus Capital Management LLC

$89,612,476

0.96%

731,053

Clearbridge Investments, LLC

$88,911,564

0.95%

725,335

FMR LLC

$82,396,559

0.88%

672,186

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	670,967	$82,247,134	99.8%	Defined
STRATEGIC ADVISERS LLC	1,219	$149,425	0.2%	Defined

NORTHERN TRUST CORP

$81,651,147

0.87%

666,105

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	598,391	$73,350,767	89.8%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	37,948	$4,651,665	5.7%	Defined
MASON STREET ADVISORS, LLC joint	20,213	$2,477,709	3.0%	Other
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	9,553	$1,171,006	1.4%	Other

AMERICAN CENTURY COMPANIES INC

$73,376,388

0.78%

598,600

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AMERICAN CENTURY INVESTMENT MANAGEMENT INC	598,600	$73,376,388	100.0%	Defined

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$71,559,016

0.76%

583,774

VOYA INVESTMENT MANAGEMENT LLC

$70,068,932

0.75%

571,618

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	571,618	$70,068,932	100.0%	Defined

Pictet Asset Management Holding SA

$69,624,214

0.74%

567,990

FRANKLIN RESOURCES INC

$63,745,519

0.68%

520,032

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Brandywine Global Investment Management, LLC	17	$2,083	0.0%	Defined
FIDUCIARY TRUST CO INTERNATIONAL	289	$35,425	0.1%	Defined
FIDUCIARY TRUST INTERNATIONAL LLC	180	$22,064	0.0%	Defined
FRANKLIN ADVISERS INC	71,483	$8,762,386	13.7%	Defined
FRANKLIN ADVISORY SERVICES LLC	3,444	$422,165	0.7%	Defined
Franklin Templeton International Services S.a r.l	178	$21,819	0.0%	Defined
PUTNAM INVESTMENT MANAGEMENT LLC	305,251	$37,417,667	58.7%	Defined
PUTNAM ADVISORY CO LLC	139,190	$17,061,910	26.8%	Defined

VANGUARD FIDUCIARY TRUST CO

$53,659,027

0.57%

437,747

LORD, ABBETT & CO. LLC

$48,908,684

0.52%

398,994

PRINCIPAL FINANCIAL GROUP INC

$47,225,415

0.50%

385,262

Bank of New York Mellon Corp

$43,920,411

0.47%

358,300

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNY Mellon Trust of Delaware joint	535	$65,580	0.1%	Defined
Mellon Investments Corporation joint	183,917	$22,544,545	51.3%	Defined
BNY Mellon Investment Adviser, Inc.	49,069	$6,014,878	13.7%	Defined
The Bank of New York Mellon	114,706	$14,060,661	32.0%	Defined
BNY Mellon, NA joint	7,803	$956,491	2.2%	Defined
BNY Mellon Advisors, Inc.	2,269	$278,134	0.6%	Defined
Held directly	1	$122	0.0%	Defined

NEUBERGER BERMAN GROUP LLC

$37,071,501

0.40%

302,427

BNP PARIBAS ARBITRAGE, SA

$36,954,437

0.39%

301,472

683 Capital Management, LLC

$35,548,200

0.38%

290,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
683 Capital Partners, LP	290,000	$35,548,200	100.0%	Defined

Nuveen, LLC

$34,823,137

0.37%

284,085

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	72,462	$8,882,391	25.5%	Defined
TIAA CREF INVESTMENT MANAGEMENT LLC joint	81,993	$10,050,701	28.9%	Defined
TEACHERS ADVISORS, LLC joint	129,630	$15,890,045	45.6%	Defined

Stephens Investment Management Group LLC

$34,546,966

0.37%

281,832

BNP PARIBAS ASSET MANAGEMENT Holding S.A.

$31,867,366

0.34%

259,972

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNP PARIBAS ASSET MANAGEMENT Europe	855	$104,805	0.3%	Defined
BNP Paribas Asset Management USA, Inc.	259,117	$31,762,561	99.7%	Defined

Eagle Health Investments LP

$30,439,310

0.32%

248,322

NEXTBio Capital Management LP

$26,885,839

0.29%

219,333

SEGALL BRYANT & HAMILL, LLC

$26,320,867

0.28%

214,724

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NEW YORK LIFE INVESTMENT MANAGEMENT LLC	214,724	$26,320,867	100.0%	Sole

SEB Asset Management AB

$25,737,264

0.27%

209,963

Duquesne Family Office LLC

$25,606,962

0.27%

208,900

Public Sector Pension Investment Board

$25,458,395

0.27%

207,688

CITIGROUP INC

$24,352,722

0.26%

198,668

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citibank, N.A. joint	4	$490	0.0%	Defined
Citigroup Financial Products Inc. joint	198,664	$24,352,232	100.0%	Defined

JPMORGAN CHASE & CO

$22,444,150

0.24%

183,098

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
J.P. Morgan Trust Co of Delaware	182	$22,309	0.1%	Other
JPMorgan Chase Bank, N.A.	26,802	$3,285,389	14.6%	Defined
J.P. Morgan Investment Management Inc.	155,854	$19,104,583	85.1%	Defined
J.P. Morgan Securities LLC	27	$3,309	0.0%	Defined
J.P. Morgan Securities plc	232	$28,438	0.1%	Defined
55I, LLC	1	$122	0.0%	Defined

DIMENSIONAL FUND ADVISORS LP

$21,837,870

0.23%

178,152

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Fund Advisors Ltd.	2,464	$302,037	1.4%	Defined
DFA Australia Ltd.	8,707	$1,067,304	4.9%	Defined
Dimensional Fund Advisors Pte. Ltd.	259	$31,748	0.1%	Defined
Dimensional Ireland Ltd	15,048	$1,844,583	8.4%	Defined
Held directly	151,674	$18,592,198	85.1%	Sole

Affinity Asset Advisors, LLC

$21,099,082

0.23%

172,125

Fisher Asset Management, LLC

$20,902,831

0.22%

170,524

Boxer Capital Management, LLC

$20,838,600

0.22%

170,000

RAYMOND JAMES FINANCIAL INC

$20,626,533

0.22%

168,270

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	19,836	$2,431,496	11.8%	Defined
Raymond James Financial Services Advisors, Inc.	124	$15,199	0.1%	Defined
EAGLE ASSET MANAGEMENT INC joint	148,310	$18,179,838	88.1%	Defined

Caption Management, LLC

$19,833,444

0.21%

161,800

NAN FUNG TRINITY (HK) LTD

$19,694,928

0.21%

160,670

Caption Management, LLC

$18,387,000

0.20%

150,000

ORBIMED ADVISORS LLC

$18,387,000

0.20%

150,000

AMERIPRISE FINANCIAL INC

$17,628,229

0.19%

143,810

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	141,913	$17,395,695	98.7%	Defined
Columbia Threadneedle Management Ltd	1,897	$232,534	1.3%	Defined

DEUTSCHE BANK AG\

$17,386,500

0.19%

141,838

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DWS Investment GmbH	45,953	$5,632,918	32.4%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC	28,256	$3,463,620	19.9%	Defined
DWS Investments UK Ltd	67,629	$8,289,962	47.7%	Defined

RHUMBLINE ADVISERS

$17,317,979

0.18%

141,279

NOMURA ASSET MANAGEMENT INTERNATIONAL INC.

$17,293,708

0.18%

141,081

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NOMURA HOLDINGS INC joint	141,081	$17,293,708	100.0%	Defined

TD Asset Management Inc

$16,542,171

0.18%

134,950

KENNEDY CAPITAL MANAGEMENT LLC

$16,428,049

0.18%

134,019

Legal & General Group Plc

$16,223,585

0.17%

132,351

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	92,699	$11,363,043	70.0%	Defined
Legal & General Investment Management Ltd joint	39,652	$4,860,542	30.0%	Defined

Russell Investments Group, Ltd.

$14,148,794

0.15%

115,425

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC	54,006	$6,620,055	46.8%	Defined
Russell Investments Canada Limited	180	$22,064	0.2%	Defined
Russell Investments Capital, LLC	34,678	$4,250,829	30.0%	Defined
Russell Investments France SAS	8,803	$1,079,071	7.6%	Defined
Russell Investments Limited	408	$50,012	0.4%	Defined
Russell Investments Trust Company	17,350	$2,126,763	15.0%	Defined

MARSHALL WACE, LLP

$13,363,303

0.14%

109,017

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marshall Wace North America L.P. joint	109,017	$13,363,303	100.0%	Other

MANUFACTURERS LIFE INSURANCE COMPANY, THE

$13,306,671

0.14%

108,555

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MANULIFE FINANCIAL CORP joint	108,555	$13,306,671	100.0%	Defined

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$13,127,579

0.14%

107,094

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	2,016	$247,121	1.9%	Defined
UBS Asset Management Switzerland AG	82,078	$10,061,121	76.6%	Defined
UBS Asset Management (UK) Ltd.	16,165	$1,981,505	15.1%	Defined
Held directly	6,835	$837,832	6.4%	Defined

Catalio Capital Management, LP

$13,091,421

0.14%

106,799

Swiss National Bank

$12,552,192

0.13%

102,400

Universal- Beteiligungs- und Servicegesellschaft mbH

$12,503,405

0.13%

102,002

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Universal-Investment-Gesellschaft mbH joint	102,002	$12,503,405	100.0%	Defined

Opaleye Management Inc.

$12,258,000

0.13%

100,000

ASSETMARK, INC

$12,097,052

0.13%

98,687

SEI INVESTMENTS CO

$11,369,172

0.12%

92,749

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI INVESTMENTS MANAGEMENT CORP	11,580	$1,419,476	12.5%	Sole
SEI TRUST CO	81,169	$9,949,696	87.5%	Sole

ALGERT GLOBAL LLC

$10,750,020

0.11%

87,698

PRUDENTIAL FINANCIAL INC

$10,441,364

0.11%

85,180

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PGIM Quantitative Solutions LLC	79,080	$9,693,626	92.8%	Defined
Prudential Trust Co	6,100	$747,738	7.2%	Defined

VICTORY CAPITAL MANAGEMENT INC

$10,282,378

0.11%

83,883

AXIOM INVESTORS LLC /DE

$10,012,702

0.11%

81,683

ALLIANCEBERNSTEIN L.P.

$9,752,832

0.10%

79,563

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	79,563	$9,752,832	100.0%	Defined

WELLS FARGO & COMPANY/MN

$8,818,893

0.09%

71,944

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	58,274	$7,143,226	81.0%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	4,846	$594,022	6.7%	Defined
WELLS FARGO CLEARING SERVICES, LLC	8,769	$1,074,904	12.2%	Defined
WELLS FARGO INVESTMENT INSTITUTE, INC.	55	$6,741	0.1%	Defined

NORDEA INVESTMENT MANAGEMENT AB

$7,963,286

0.09%

64,964

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nordea Funds Ltd	13,642	$1,672,236	21.0%	Defined
Nordea Investment Funds S.A.	51,322	$6,291,050	79.0%	Defined

Knott David M Jr

$7,722,540

0.08%

63,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dorset Partners LLC	63,000	$7,722,540	100.0%	Defined

Atika Capital Management LLC

$7,612,217

0.08%

62,100

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Atika Capital Partners LP	36,256	$4,444,260	58.4%	Defined
Optika Master Portfolio Ltd	15,671	$1,920,951	25.2%	Other
Held directly	10,173	$1,247,006	16.4%	Sole

Moody Aldrich Partners LLC

$7,305,890

0.08%

59,601

CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM

$7,227,439

0.08%

58,961

Eversept Partners, LP

$7,171,420

0.08%

58,504

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$6,994,659

0.07%

57,062

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	57,062	$6,994,659	100.0%	Other

GREAT LAKES ADVISORS, LLC

$6,975,292

0.07%

56,904

Creative Planning

$6,887,156

0.07%

56,185

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CREATIVE PLANNING TRUST Co LLC	1,499	$183,747	2.7%	Defined
Held directly	54,686	$6,703,409	97.3%	Sole

Bellevue Group AG

$6,825,499

0.07%

55,682

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Bellevue Asset Management AG	55,682	$6,825,499	100.0%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$6,790,932

0.07%

55,400

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	55,400	$6,790,932	100.0%	Other

FIRST TRUST ADVISORS LP

$6,725,351

0.07%

54,865

KVP Capital Advisors, LP

$6,496,740

0.07%

53,000

BESSEMER GROUP INC

$6,383,228

0.07%

52,074

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BESSEMER TRUST CO	1,055	$129,321	2.0%	Other
BESSEMER TRUST Co NATIONAL ASSOCIATION	50,765	$6,222,773	97.5%	Defined
Bessemer Trust CO of Delaware, N.A.	65	$7,967	0.1%	Other
BESSEMER TRUST Co OF FLORIDA	189	$23,167	0.4%	Defined

VANGUARD ASSET MANAGEMENT, Ltd

$6,329,908

0.07%

51,639

CASTLEARK MANAGEMENT LLC

$6,198,870

0.07%

50,570

Granahan Investment Management, LLC

$6,105,587

0.07%

49,809

NEW YORK STATE TEACHERS RETIREMENT SYSTEM

$6,073,839

0.06%

49,550

NORTHWESTERN MUTUAL WEALTH MANAGEMENT CO

$5,993,426

0.06%

48,894

Key facts CIK 1377121 CUSIP 74366E102 13F (30d) 12 filings 12 filers Visit website Investor relations