SRRK · Scholar Rock Holding Corp · Holdings

$41.02 +0.78 (+1.94%)

Russell 2000 member Rebalance outlook locked

Market Cap

$4.90B

121.79M

Volume · Oct 20 1.61M Avg daily vol (3M) 1.6M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Use institutional holdings through the 13F API or MCP → See every holder of SRRK: 13F, 13D and 13G, insiders and funds →

Institutional holdings are being reconciled with SEC filings. Holder totals and changes may be incomplete until reconciliation finishes.

The Sep 30, 2026 filing window is still open — showing the latest known positions. Funds that have not filed yet carry their prior-quarter positions; a fund that filed without this stock counts as sold out. Each fund's Change is measured against its own previous reported quarter, shown in the Reported column. Pick a completed quarter in the Quarter selector below to put every fund on one quarter and see the stock-wide activity breakdown.

Quarter:

Holders: 309

Total Value: $7,660,319,820

Total Shares: 139,281,459

Showing 1–100 of 315 holdings by value · page 1 of 4

Institutional Ownership Trend

27 quarters

Holder Concentration

Across 310 13F filers

Top Holder

12.9%

of institutional value

Top 5

36.4%

of institutional value

Top 10

55.4%

of institutional value

HHI

428

Broadly held

Who Holds SRRK

Last 8 quarters through Sep 30, 2026 (filing window open)

Each kind of 13F filer's share of the common shares 13F filers report. Holder type comes from regulatory records, never the filer's name; style, turnover and horizon are measured from each filer's own 13F book. The quarter selector does not change this card.

Jun 30, 2026, the newest closed quarter: 306 filers, like-for-like change +2.4%; actively managed books -2.8%; index-like books +27.8%.

Holder type	Share	Filers	Change	New	Exited
Registered adviser	40.5%	154	-3.9%	31	14
Exempt adviser	0.7%	5	-13.2%	3	1
Adviser	2.5%	10	+33.2%	1	0
Bank	5.8%	17	+8.9%	0	0
Insurer	0.0%	3	+171.4%	1	0
Broker-dealer	0.0%	2	+62.9%	0	0
Benefit plan	0.1%	5	+23.1%	0	0
Non-US	5.3%	9	-19.1%	0	0
Holding company	26.8%	7	+12.5%	0	0
Unknown	18.2%	94	+4.0%	17	6

Style	Share	Filers	Change	New	Exited
Index-like	22.6%	34	+27.8%	2	0
Systematic-like	2.8%	26	-7.3%	9	2
Concentrated	9.3%	40	-29.8%	11	4
Diversified active	65.3%	206	+3.1%	31	15
Not measured	0.0%	0	—	0	0
Unknown	0.0%	0	—	0	0

Turnover	Share	Filers	Change	New	Exited
Low turnover	51.3%	149	+9.2%	17	7
Medium turnover	35.1%	81	-2.7%	12	7
High turnover	4.8%	45	-29.5%	17	6
Unknown	8.8%	31	+16.4%	7	1
Not measured	0.0%	0	—	0	0

Holding horizon	Share	Filers	Change	New	Exited
Short horizon	7.4%	50	+6.6%	18	3
Medium horizon	17.0%	51	-7.2%	9	6
Long horizon	58.9%	178	+3.2%	21	9
Unknown	16.7%	27	+9.1%	5	3
Not measured	0.0%	0	—	0	0

13F book size	Share	Filers	Change	New	Exited
Under $1B	1.4%	60	-39.5%	20	4
$1B to $10B	22.0%	86	-22.4%	18	10
$10B to $100B	16.9%	92	+15.5%	13	5
$100B or more	59.8%	68	+14.3%	2	2
Not measured	0.0%	0	—	0	0

Client base	Share	Filers	Change	New	Exited
Individuals and high net worth	0.3%	58	+32.5%	13	5
Pooled investment vehicles	16.7%	39	-25.1%	9	8
Registered funds and BDCs	18.5%	22	+19.6%	5	0
Pension and profit sharing plans	0.0%	2	+573.8%	1	0
Insurance companies	0.7%	3	+23.8%	0	0
Governments and sovereign funds	0.6%	6	+16.0%	1	0
Other investment advisers	0.0%	3	+5.6%	0	0
Other	0.0%	1	0.0%	0	0
Diversified	3.7%	19	+20.7%	2	1
Corporations	0.0%	0	—	0	0
Unknown	59.5%	153	+7.4%	22	7

Private funds	Share	Filers	Change	New	Exited
Hedge funds	14.7%	47	-27.5%	11	10
Private equity funds	1.8%	12	+7.5%	0	0
Venture capital funds	6.6%	4	+1.3%	1	0
Real estate funds	0.0%	3	+278.9%	0	1
Liquidity funds	0.0%	1	new	1	0
Securitized asset funds	0.1%	1	+8.0%	0	0
Other private funds	7.6%	14	-1.7%	2	0
Mixed private funds	0.6%	4	+69.6%	0	0
Private funds, type not stated	1.9%	3	+4981.0%	1	0
No private funds	8.0%	70	+24.5%	18	4
Unknown	58.7%	147	+7.7%	19	6

Location	Share	Filers	Change	New	Exited
United States	86.4%	247	+2.6%	44	18
Outside the US	13.6%	59	+1.0%	9	3

The Sep 30, 2026 filing window is still open, so its bar carries the Jun 30, 2026 positions of filers that have not filed yet.

How the mix is measured

Shares are 13F common shares restated to today's split basis; options and principal amounts are left out. The change compares only filers with a 13F in both quarters and keeps each filer in its bucket of the later quarter on both sides; a filer that continues under a new CIK is compared through its successor. Holder type comes from each filer's current regulatory profile; client base and private funds come from the Form ADV roster current at each quarter end (rosters from December 2019, so an earlier quarter reads Unknown). Style, turnover, horizon and book size are measured from each filer's 13F book in the newest measured quarter, and Not measured means none was. Actively managed books are those measured systematic-like, concentrated or diversified active.

Excluded 3 holders with no observed 2026-03-31 13F and 3 holders with no observed 2026-06-30 13F. Compared across CIK changes: JANUS HENDERSON GROUP PLC continues as Jupiter Topco LLC.

Conviction and Crowding

306 13F filers, quarter ended Jun 30, 2026

Measured across every 13F filer's common-share book once the quarter's filing window closed. Crowding counts filers with high turnover or that advise private funds on Form ADV.

Conviction holders

1.96% of filers hold it in their top 10

Mar 31, 2026: 10

Holders above 5%

median weight 0.005%

Mar 31, 2026: 6

Crowding

—

holders exceed shares outstanding: 13F filers report more shares than the cover-page count, which predates an issuance, is misstated, or misses shares lent and reported twice

Private-fund advisers

—

of shares outstanding

Institutional ownership

—

holders exceed shares outstanding: 13F filers report more shares than the cover-page count, which predates an issuance, is misstated, or misses shares lent and reported twice

Top 10 holders

55.7%

of 13F shares, Herfindahl 0.043

Active holders

-2.8%

like-for-like share change of active filers

5% stakes on record

1 13D, 6 13G (2 institutional, 2 passive, 0 exempt)

How long holders have held it value-weighted 13.9 quarters

Under 1 year 13.2% 1 to 3 years 42.6% 3 years or more 44.2% Since coverage start 0.000%

Managers in the top fifth by current three-year FundScore alpha were net buyers of $184,877 (1 buying, 0 selling).

Estimated Holder Cost Basis

Quarter ended Jun 30, 2026, on that quarter's share basis

Estimated, never filed: each 13F buy is costed at its quarter's average traded price and each sale keeps the average cost, so this is what holders paid on average if they traded evenly through each quarter.

Estimated avg cost

$33.77

estimated per share

Mar 31, 2026: $31.60

Quarter-end price

$55.00

implied by 13F values

Estimated gain or loss

+62.9%

price against the estimated cost

Estimated at a loss

0.000%

of costed 13F value

Estimated cost covers 45.2% of the 13F shares; positions already held when a filer's 13F history starts, or bought in a quarter without a trusted average price, have no estimate.

Shared Holders

Companies SRRK's active holders also own, quarter ended Jun 30, 2026

#	Company	Shared holders	Similarity
1	XENE Xenon Pharmaceuticals Inc.	15	0.313
2	PRAX Praxis Precision Medicines, Inc.	14	0.300
3	CYTK Cytokinetics Inc	17	0.285
4	SION Sionna Therapeutics, Inc.	9	0.276
5	ELVN Enliven Therapeutics, Inc.	12	0.265
6	PTGX Protagonist Therapeutics, Inc	11	0.259
7	SYRE Spyre Therapeutics, Inc.	12	0.257
8	MLYS Mineralys Therapeutics, Inc.	11	0.256
9	BBIO BridgeBio Pharma, Inc.	13	0.252
10	VERA Vera Therapeutics, Inc.	10	0.247

Shareholder Targets

13F filers that own its peers but not SRRK, quarter ended Jun 30, 2026

Found through its holder-overlap peers: XENE, PRAX, CYTK, SION, ELVN, PTGX, SYRE, MLYS, BBIO, VERA, NAMS, IRON, KYMR, INSM, RYTM, CLDX, SEPN, CRNX, ERAS, TRVI. 855 filers in all.

#	Filer	In peers	Of book	Largest peers
1	BAKER BROS. ADVISORS LP Registered adviser, Concentrated	$3,401,234,918 11 held	17.1%	KYMR, INSM, RYTM
2	T. Rowe Price Investment Management, Inc. Adviser, Diversified active	$3,148,399,977 8 held	2.07%	CYTK, KYMR, ERAS
3	RA CAPITAL MANAGEMENT, L.P. Registered adviser, Concentrated	$2,127,771,981 6 held	18.7%	RYTM, SION, NAMS
4	RTW INVESTMENTS, LP Registered adviser, Diversified active	$2,116,552,468 7 held	18.1%	PTGX, INSM, SYRE
5	Avoro Capital Advisors LLC Registered adviser, Concentrated	$1,922,230,680 5 held	17.3%	KYMR, CYTK, XENE
6	FARALLON CAPITAL MANAGEMENT LLC Registered adviser, Diversified active	$1,914,764,276 3 held	11.3%	BBIO, PTGX, CRNX
7	Frazier Life Sciences Management, L.P. Registered adviser, Concentrated	$1,579,850,493 6 held	34.4%	NAMS, ERAS, TRVI
8	PERCEPTIVE ADVISORS LLC Registered adviser, Diversified active	$1,512,438,616 5 held	24.4%	PRAX, SYRE, RYTM
9	Capital International Investors Adviser, Diversified active	$1,275,354,046 5 held	0.260%	INSM, XENE, CYTK
10	VIKING GLOBAL INVESTORS LP Registered adviser, Diversified active	$1,052,840,338 4 held	3.06%	BBIO, NAMS, SION

Fund Ownership of SRRK by Fund Type

Registered funds' Form N-PORT reports for the quarter ended Jun 30, 2026

Index funds and index ETFs hold 13.56% of shares outstanding.

Fund type	Of shares outstanding	Change	Shares	Funds
Index funds	2.70%	+0.59 pp	3,282,945	65
Index ETFs	10.86%	+1.97 pp	13,230,608	43
Active ETFs	0.25%	+0.12 pp	308,891	14
Active funds	27.17%	+6.05 pp	33,084,501	159
Unclassified	0.12%	+0.12 pp	149,988	11
All registered funds	41.10%	+8.84 pp	50,056,933	292

How fund types are measured

Each fund series counts once, with its latest Form N-PORT report dated inside the quarter; only long share positions count, restated to today's split basis. The fund type is the series' latest Form N-CEN answer on or before the quarter end (the index-fund box and the exchange-traded flag), never its name. Unclassified funds have no usable N-CEN; unit investment trusts are among them, so index ownership is a floor. Funds sit inside their advisers' 13F reports, so these shares overlap the 13F figures and are never added to them.

Holder type: Registered adviser 157 Exempt adviser 5 Adviser 10 Bank 17 Insurer 3 Broker-dealer 2 Benefit plan 5 Non-US 9 Holding company 7 Unknown 94

Showing 100 of 315

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

FMR LLC

$989,738,255

12.92%

17,995,241

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	11,376,577	$625,711,735	63.2%	Defined
FIDELITY MANAGEMENT TRUST CO	6,273,240	$345,028,200	34.9%	Defined
STRATEGIC ADVISERS LLC	2	$110	0.0%	Defined
Fidelity Institutional Asset Management Trust Co	46,300	$2,546,500	0.3%	Defined
FIAM LLC	299,122	$16,451,710	1.7%	Defined

Siren, L.L.C.

$526,625,715

6.87%

9,575,013

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invus Group, LLC	9,575,013	$526,625,715	100.0%	Defined

BlackRock, Inc.

$522,174,620

6.82%

9,494,084

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	111,940	$6,156,700	1.2%	Sole
BlackRock Investment Management (Australia) Ltd	88	$4,840	0.0%	Sole
BlackRock Asset Management Canada Ltd	9,705	$533,775	0.1%	Sole
BlackRock Investment Management, LLC	322,865	$17,757,575	3.4%	Sole
BLACKROCK ADVISORS LLC	662,627	$36,444,485	7.0%	Sole
BlackRock Fund Advisors	4,886,579	$268,761,845	51.5%	Sole
BlackRock Institutional Trust Company, N.A.	3,178,326	$174,807,930	33.5%	Sole
BlackRock Japan Co. Ltd	6,743	$370,865	0.1%	Sole
BlackRock Fund Managers Ltd.	13,409	$737,495	0.1%	Sole
BlackRock Investment Management (UK) Ltd.	32,212	$1,771,660	0.3%	Sole
BlackRock (Netherlands) B.V.	2,829	$155,595	0.0%	Sole
BlackRock Asset Management Ireland Ltd	173,442	$9,539,310	1.8%	Sole
BlackRock Asset Management Schweiz AG	12,949	$712,195	0.1%	Sole
Aperio Group, LLC	80,367	$4,420,185	0.8%	Sole
SpiderRock Advisors, LLC	3	$165	0.0%	Sole

SAMSARA BIOCAPITAL, LLC

$379,370,860

4.95%

6,897,652

PRICE T ROWE ASSOCIATES INC /MD/

$373,716,035

4.88%

6,794,837

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	158,833	$8,735,815	2.3%	Defined
Held directly	6,636,004	$364,980,220	97.7%	Sole

WELLINGTON MANAGEMENT GROUP LLP

$369,989,675

4.83%

6,727,085

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wellington Management Company LLP	5,829,338	$320,613,590	86.7%	Defined
Wellington Managment Canada LLC	20,301	$1,116,555	0.3%	Defined
Wellington Management Singapore Pte. Ltd.	82,598	$4,542,890	1.2%	Defined
Wellington Management International Ltd	221,158	$12,163,690	3.3%	Defined
Wellington Trust Company, NA joint	555,960	$30,577,800	8.3%	Defined
Wellington Management Australia Pty Ltd	15,627	$859,485	0.2%	Defined
Wellington Management Europe GmbH	2,103	$115,665	0.0%	Defined

Orbis Allan Gray Ltd

$326,344,315

4.26%

5,933,533

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ORBIS INVESTMENT MANAGEMENT LTD	5,926,433	$325,953,815	99.9%	Defined
Held directly	7,100	$390,500	0.1%	Defined

VANGUARD CAPITAL MANAGEMENT LLC

$262,233,400

3.42%

4,767,880

STATE STREET CORP

$260,077,455

3.40%

4,728,681

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	4,728,681	$260,077,455	100.0%	Defined

Redmile Group, LLC

$237,110,060

3.10%

4,311,092

NORGES BANK

$152,345,050

1.99%

2,769,910

GEODE CAPITAL MANAGEMENT, LLC

$151,458,285

1.98%

2,753,787

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	384,976	$21,173,680	14.0%	Defined
Held directly	2,368,811	$130,284,605	86.0%	Defined

MASSACHUSETTS FINANCIAL SERVICES CO /MA/

$143,404,140

1.87%

2,607,348

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MFS Institutional Advisors, Inc.	260,301	$14,316,555	10.0%	Defined
MFS Investment Management CO (LUX) S.A.R.L.	16,130	$887,150	0.6%	Defined
MFS Investment Management K.K.	15,368	$845,240	0.6%	Defined
MFS Heritage Trust CO	416,566	$22,911,130	16.0%	Defined
MFS INVESTMENT MANAGEMENT CANADA LTD	4,366	$240,130	0.2%	Defined
Held directly	1,894,617	$104,203,935	72.7%	Sole

Bellevue Group AG

$134,725,085

1.76%

2,449,547

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BB Biotech AG joint	2,445,032	$134,476,760	99.8%	Defined
Bellevue Asset Management AG	4,515	$248,325	0.2%	Defined

JENNISON ASSOCIATES LLC

$129,722,285

1.69%

2,358,587

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Prudential Financial Inc	1,861,477	$102,381,235	78.9%	Defined
Prudential Trust Company	497,110	$27,341,050	21.1%	Other

Jupiter Topco LLC

$123,315,830

1.61%

2,242,106

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	40,395	$2,221,725	1.8%	Defined
JANUS HENDERSON INVESTORS US LLC	2,201,711	$121,094,105	98.2%	Defined

Hood River Capital Management LLC

$122,363,340

1.60%

2,224,788

Driehaus Capital Management LLC

$121,308,990

1.58%

2,205,618

EVENTIDE ASSET MANAGEMENT, LLC

$116,127,550

1.52%

2,111,410

Polar Capital Holdings Plc

$103,321,900

1.35%

1,878,580

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Polar Capital LLP	1,878,580	$103,321,900	100.0%	Defined

Holocene Advisors, LP

$96,578,790

1.26%

1,755,978

ADAGE CAPITAL PARTNERS GP, L.L.C.

$95,227,110

1.24%

1,731,402

MORGAN STANLEY

$84,632,680

1.10%

1,538,776

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	382	$21,010	0.0%	Defined
MORGAN STANLEY & CO. LLC	42,237	$2,323,035	2.7%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	1,404,585	$77,252,175	91.3%	Defined
MORGAN STANLEY SMITH BARNEY LLC	4,480	$246,400	0.3%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	61,928	$3,406,040	4.0%	Defined
EATON VANCE MANAGEMENT	25,164	$1,384,020	1.6%	Defined

Woodline Partners LP

$82,525,850

1.08%

1,500,470

GOLDMAN SACHS GROUP INC

$77,631,235

1.01%

1,411,477

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	1,197,209	$65,846,495	84.8%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	208,669	$11,476,795	14.8%	Defined
Goldman Sachs Trust Company, N.A.	5,599	$307,945	0.4%	Defined

Capital World Investors

$63,822,495

0.83%

1,160,409

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CAPITAL GROUP COMPANIES INC joint	1,160,409	$63,822,495	100.0%	Defined

Clearbridge Investments, LLC

$58,287,680

0.76%

1,059,776

VANGUARD PORTFOLIO MANAGEMENT LLC

$56,504,635

0.74%

1,027,357

BNP PARIBAS ASSET MANAGEMENT Holding S.A.

$53,891,475

0.70%

979,845

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNP PARIBAS ASSET MANAGEMENT Europe	1,115	$61,325	0.1%	Defined
BNP Paribas Asset Management USA, Inc.	851,963	$46,857,965	86.9%	Defined
AXA INVESTMENT MANAGERS UK LIMITED	126,767	$6,972,185	12.9%	Defined

AMERICAN CENTURY COMPANIES INC

$53,635,835

0.70%

975,197

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AMERICAN CENTURY INVESTMENT MANAGEMENT INC	975,197	$53,635,835	100.0%	Defined

NORTHERN TRUST CORP

$53,601,680

0.70%

974,576

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	906,162	$49,838,910	93.0%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	51,061	$2,808,355	5.2%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	17,353	$954,415	1.8%	Other

BANK OF AMERICA CORP /DE/

$53,474,135

0.70%

972,257

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	761,408	$41,877,440	78.3%	Defined
BOFA SECURITIES, INC.	87,934	$4,836,370	9.0%	Defined
MANAGED ACCOUNT ADVISORS LLC joint	122,915	$6,760,325	12.6%	Defined

Pictet Asset Management Holding SA

$50,109,125

0.65%

911,075

VOYA INVESTMENT MANAGEMENT LLC

$48,983,385

0.64%

890,607

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	890,607	$48,983,385	100.0%	Defined

FRANKLIN RESOURCES INC

$48,329,930

0.63%

878,726

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FIDUCIARY TRUST INTERNATIONAL LLC	256	$14,080	0.0%	Defined
FRANKLIN ADVISERS INC	33,230	$1,827,650	3.8%	Defined
PUTNAM INVESTMENT MANAGEMENT LLC	553,659	$30,451,245	63.0%	Defined
PUTNAM ADVISORY CO LLC	291,581	$16,036,955	33.2%	Defined

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$47,535,730

0.62%

864,286

Integral Health Asset Management, LLC

$41,250,000

0.54%

750,000

FEDERATED HERMES, INC.

$40,398,215

0.53%

734,513

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FEDERATED GLOBAL INVESTMENT MANAGEMENT CORP	734,513	$40,398,215	100.0%	Defined

VANGUARD FIDUCIARY TRUST CO

$40,154,565

0.52%

730,083

BNP PARIBAS ARBITRAGE, SA

$39,463,160

0.52%

717,512

Rock Springs Capital Management LP

$39,349,860

0.51%

715,452

Casdin Capital, LLC

$36,575,000

0.48%

665,000

Nuveen, LLC

$35,846,195

0.47%

651,749

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	19,472	$1,070,960	3.0%	Defined
TIAA CREF INVESTMENT MANAGEMENT LLC joint	88,690	$4,877,950	13.6%	Defined
TEACHERS ADVISORS, LLC joint	543,587	$29,897,285	83.4%	Defined

LOOMIS SAYLES & CO L P

$32,783,025

0.43%

596,055

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
LOOMIS SAYLES TRUST COMPANY, LLC	100,389	$5,521,395	16.8%	Defined
Held directly	495,666	$27,261,630	83.2%	Defined

Ensign Peak Advisors, Inc

$31,847,860

0.42%

579,052

Eagle Health Investments LP

$30,539,960

0.40%

555,272

UBS Group AG

$28,114,350

0.37%

511,170

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	26,541	$1,459,755	5.2%	Defined
UBS AG	464,846	$25,566,530	90.9%	Defined
UBS Switzerland AG	19,731	$1,085,205	3.9%	Defined
UBS Securities LLC	52	$2,860	0.0%	Defined

Qube Research & Technologies Ltd

$25,505,150

0.33%

463,730

Fisher Asset Management, LLC

$24,169,585

0.32%

439,447

MARSHALL WACE, LLP

$23,677,500

0.31%

430,500

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marshall Wace North America L.P. joint	421,259	$23,169,245	97.9%	Other
MARSHALL WACE ASIA Ltd	9,241	$508,255	2.1%	Other

Bank of New York Mellon Corp

$20,086,990

0.26%

365,218

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNY Mellon Trust of Delaware joint	270	$14,850	0.1%	Defined
Mellon Investments Corporation joint	149,741	$8,235,755	41.0%	Defined
The Bank of New York Mellon	200,020	$11,001,100	54.8%	Defined
BNY Mellon, NA joint	13,262	$729,410	3.6%	Defined
BNY Mellon Advisors, Inc.	1,923	$105,765	0.5%	Defined
Held directly	2	$110	0.0%	Defined

CITADEL ADVISORS LLC

$18,251,805

0.24%

331,851

DEUTSCHE BANK AG\

$16,679,740

0.22%

303,268

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DWS Investment GmbH	139,925	$7,695,875	46.1%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC	51,181	$2,814,955	16.9%	Defined
DWS Investments UK Ltd	112,162	$6,168,910	37.0%	Defined

Squarepoint Ops LLC

$16,059,780

0.21%

291,996

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Squarepoint Capital LLP	291,996	$16,059,780	100.0%	Defined

NAN FUNG TRINITY (HK) LTD

$15,702,225

0.20%

285,495

ARMISTICE CAPITAL, LLC

$15,290,000

0.20%

278,000

Trexquant Investment LP

$15,164,160

0.20%

275,712

First Turn Management, LLC

$14,917,485

0.19%

271,227

AQR CAPITAL MANAGEMENT LLC

$13,613,655

0.18%

247,521

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RF&L WEALTH MANAGEMENT, LLC	4	$220	0.0%	Other
Prism Planning Partners LLC	92	$5,060	0.0%	Other
Arax Advisory Partners	1,204	$66,220	0.5%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	406	$22,330	0.2%	Other
Stokes Family Office, LLC	303	$16,665	0.1%	Other
Held directly	245,512	$13,503,160	99.2%	Defined

D. E. Shaw & Co., Inc.

$13,476,485

0.18%

245,027

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	153,988	$8,469,340	62.8%	Defined
D. E. Shaw Investment Management, L.L.C. joint	91,039	$5,007,145	37.2%	Defined

RAYMOND JAMES FINANCIAL INC

$13,236,575

0.17%

240,665

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	84,051	$4,622,805	34.9%	Defined
Raymond James Financial Services Advisors, Inc.	44,782	$2,463,010	18.6%	Defined
CHARTWELL INVESTMENT PARTNERS, LLC	111,832	$6,150,760	46.5%	Defined

STEMPOINT CAPITAL LP

$13,153,030

0.17%

239,146

Candriam S.C.A.

$11,888,030

0.16%

216,146

Universal- Beteiligungs- und Servicegesellschaft mbH

$11,549,505

0.15%

209,991

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Universal-Investment-Gesellschaft mbH joint	209,991	$11,549,505	100.0%	Defined

Senator Investment Group LP

$11,275,000

0.15%

205,000

Logos Global Management LP

$11,000,000

0.14%

200,000

Susquehanna Portfolio Strategies, LLC

$10,947,860

0.14%

199,052

SEGALL BRYANT & HAMILL, LLC

$10,372,340

0.14%

188,588

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NEW YORK LIFE INVESTMENT MANAGEMENT LLC	188,588	$10,372,340	100.0%	Sole

JPMORGAN CHASE & CO

$9,773,060

0.13%

177,692

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
J.P. Morgan Trust Co of Delaware	351	$19,305	0.2%	Other
JPMorgan Chase Bank, N.A.	52,258	$2,874,190	29.4%	Defined
J.P. Morgan Investment Management Inc.	116,156	$6,388,580	65.4%	Defined
J.P. Morgan Securities LLC	422	$23,210	0.2%	Defined
J.P. Morgan Securities plc	8,505	$467,775	4.8%	Defined

SEI INVESTMENTS CO

$9,549,265

0.12%

173,623

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI INVESTMENTS MANAGEMENT CORP	3,774	$207,570	2.2%	Sole
SEI TRUST CO	169,849	$9,341,695	97.8%	Sole

Granahan Investment Management, LLC

$9,006,745

0.12%

163,759

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$8,888,000

0.12%

161,600

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	161,600	$8,888,000	100.0%	Other

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$8,624,000

0.11%

156,800

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	156,800	$8,624,000	100.0%	Other

Swiss National Bank

$8,239,000

0.11%

149,800

RHUMBLINE ADVISERS

$8,000,300

0.10%

145,460

ALLIANCEBERNSTEIN L.P.

$7,690,100

0.10%

139,820

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	139,820	$7,690,100	100.0%	Defined

PEREGRINE CAPITAL MANAGEMENT LLC

$7,674,370

0.10%

139,534

AMERIPRISE FINANCIAL INC

$7,341,510

0.10%

133,482

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	124,316	$6,837,380	93.1%	Defined
Columbia Threadneedle Management Ltd	9,166	$504,130	6.9%	Defined

GordonMD Global Investments LP

$7,287,500

0.10%

132,500

Russell Investments Group, Ltd.

$7,178,270

0.09%

130,514

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC	87,554	$4,815,470	67.1%	Defined
Russell Investments Capital, LLC	21,702	$1,193,610	16.6%	Defined
Russell Investments France SAS	7,587	$417,285	5.8%	Defined
Russell Investments Trust Company	13,671	$751,905	10.5%	Defined

MGO ONE SEVEN LLC

$6,879,235

0.09%

125,077

iA Global Asset Management Inc.

$6,754,660

0.09%

122,812

MANUFACTURERS LIFE INSURANCE COMPANY, THE

$6,722,265

0.09%

122,223

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MANULIFE FINANCIAL CORP joint	122,223	$6,722,265	100.0%	Defined

TD Asset Management Inc

$6,620,020

0.09%

120,364

WELLS FARGO & COMPANY/MN

$6,325,605

0.08%

115,011

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	101,523	$5,583,765	88.3%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	5,154	$283,470	4.5%	Defined
WELLS FARGO CLEARING SERVICES, LLC	8,334	$458,370	7.2%	Defined

CITIGROUP INC

$6,216,705

0.08%

113,031

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	113,031	$6,216,705	100.0%	Defined

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$6,115,230

0.08%

111,186

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	1,454	$79,970	1.3%	Defined
UBS Asset Management Switzerland AG	92,625	$5,094,375	83.3%	Sole
UBS Asset Management (UK) Ltd.	7,733	$425,315	7.0%	Defined
Held directly	9,374	$515,570	8.4%	Defined

EFG International AG

$6,084,870

0.08%

110,634

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
EFG Asset Management (North America) Corp.	110,587	$6,082,285	100.0%	Defined
EFG Private Wealth Management (Americas) Corp.	47	$2,585	0.0%	Defined

Man Group plc

$5,739,030

0.07%

104,346

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AHL Partners LLP	104,346	$5,739,030	100.0%	Defined

CITADEL ADVISORS LLC

$5,566,000

0.07%

101,200

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	101,200	$5,566,000	100.0%	Defined

Ghisallo Capital Management LLC

$5,500,000

0.07%

100,000

Artia Global Partners LP

$5,295,675

0.07%

96,285

SPHERA FUNDS MANAGEMENT LTD.

$5,184,575

0.07%

94,265

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Sphera Global Healthcare Management LP joint	94,265	$5,184,575	100.0%	Defined

Invesco Ltd.

$5,178,580

0.07%

94,156

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	16,833	$925,815	17.9%	Defined
Invesco Trust Co joint	44,214	$2,431,770	47.0%	Defined
Invesco Capital Management LLC	33,109	$1,820,995	35.2%	Defined

RENAISSANCE TECHNOLOGIES LLC

$5,032,500

0.07%

91,500

FIDEURAM - INTESA SANPAOLO PRIVATE BANKING S.P.A.

$5,005,990

0.07%

91,018

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Intesa Sanpaolo S.p.A.	91,018	$5,005,990	100.0%	Defined

Nicholas Investment Partners, LP

$4,968,370

0.06%

90,334

BEACONLIGHT CAPITAL, LLC

$4,834,115

0.06%

87,893

CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM

$4,710,420

0.06%

85,644

VANGUARD ASSET MANAGEMENT, Ltd

$4,385,535

0.06%

79,737

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Key facts CIK 1727196 CUSIP 80706P103 13F (30d) 16 filings 16 filers Visit website Investor relations