SRRK · Scholar Rock Holding Corp

$57.80 -1.46 (-2.46%) At close · Aug 28

Russell 2000 member Rebalance outlook locked

Market Cap

$6.99B

121.79M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Quarter:

Holders: 308

Total Value: $7,658,995,410

Total Shares: 139,254,462

Showing 1–100 of 314 holdings by value · page 1 of 4

Institutional Ownership Trend

26 quarters

Holder Concentration

Across 308 13F filers · Δ vs Mar 31, 2026

Top Holder

12.9%

+0.0 pts

Top 5

36.4%

−2.0 pts

Top 10

55.5%

−1.7 pts

HHI

428

Broadly held−25

Quarterly Activity

vs Mar 31, 2026

New

Increased

128

Decreased

Sold Out

Unchanged

Holder count: +27

Institutional value: +16.4%

Shares held: +4.0%

Top Buyers

vs Mar 31, 2026

Institution	Added	Now Holds
Mega BlackRock, Inc.	+2,024,791	9,494,084
Large Orbis Allan Gray Ltd	+1,753,550	5,933,533
Mega MORGAN STANLEY	+1,319,749	1,538,776
Large ADAGE CAPITAL PARTNERS GP, L.L.C.	+1,031,402	1,731,402
Mega VANGUARD CAPITAL MANAGEMENT LLC	+876,533	4,767,880

Top Sellers

vs Mar 31, 2026

Institution	Sold	Now Holds
Mid Redmile Group, LLC	−4,755,529	4,311,092
Mid Siren, L.L.C.	−1,250,000	9,575,013
Mid Bellevue Group AG	−1,135,026	2,449,547
Mega JENNISON ASSOCIATES LLC	−981,905	2,358,587
Small STEMPOINT CAPITAL LP	−896,350	239,146

Top New Positions

vs Mar 31, 2026

Institution	Acquired	Value (USD)
Mega NORGES BANK	2,769,910	$152,345,050
Mega MASSACHUSETTS FINANCIAL SERVICES CO /MA/	2,607,348	$143,404,140
Mega Jupiter Topco LLC	2,242,106	$123,315,830
Mega Qube Research & Technologies Ltd	463,730	$25,505,150
Mid Senator Investment Group LP	205,000	$11,275,000

Top Sold-Out Positions

vs Mar 31, 2026

Institution	Previously Held	Last Value (USD)
Mega JANUS HENDERSON GROUP PLC	2,236,837	$109,962,906
Mid Vestal Point Capital, LP	1,324,422	$65,108,585
Mid Paradigm Biocapital Advisors LP	1,226,080	$60,274,092
Mid ADAR1 Capital Management, LLC	407,273	$20,021,540
Small SILVERARC CAPITAL MANAGEMENT, LLC	339,613	$16,695,375

Filter: New 57 Increased 128 Reduced 85 Sold out 30 Unchanged 38

Showing 100 of 314

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

FMR LLC

$989,738,255

12.92%

17,995,241

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	11,376,577	$625,711,735	63.2%	Defined
FIDELITY MANAGEMENT TRUST CO	6,273,240	$345,028,200	34.9%	Defined
STRATEGIC ADVISERS LLC	2	$110	0.0%	Defined
Fidelity Institutional Asset Management Trust Co	46,300	$2,546,500	0.3%	Defined
FIAM LLC	299,122	$16,451,710	1.7%	Defined

Siren, L.L.C.

$526,625,715

6.88%

9,575,013

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invus Group, LLC	9,575,013	$526,625,715	100.0%	Defined

BlackRock, Inc.

$522,174,620

6.82%

9,494,084

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	111,940	$6,156,700	1.2%	Sole
BlackRock Investment Management (Australia) Ltd	88	$4,840	0.0%	Sole
BlackRock Asset Management Canada Ltd	9,705	$533,775	0.1%	Sole
BlackRock Investment Management, LLC	322,865	$17,757,575	3.4%	Sole
BLACKROCK ADVISORS LLC	662,627	$36,444,485	7.0%	Sole
BlackRock Fund Advisors	4,886,579	$268,761,845	51.5%	Sole
BlackRock Institutional Trust Company, N.A.	3,178,326	$174,807,930	33.5%	Sole
BlackRock Japan Co. Ltd	6,743	$370,865	0.1%	Sole
BlackRock Fund Managers Ltd.	13,409	$737,495	0.1%	Sole
BlackRock Investment Management (UK) Ltd.	32,212	$1,771,660	0.3%	Sole
BlackRock (Netherlands) B.V.	2,829	$155,595	0.0%	Sole
BlackRock Asset Management Ireland Ltd	173,442	$9,539,310	1.8%	Sole
BlackRock Asset Management Schweiz AG	12,949	$712,195	0.1%	Sole
Aperio Group, LLC	80,367	$4,420,185	0.8%	Sole
SpiderRock Advisors, LLC	3	$165	0.0%	Sole

SAMSARA BIOCAPITAL, LLC

$379,370,860

4.95%

6,897,652

PRICE T ROWE ASSOCIATES INC /MD/

$373,716,035

4.88%

6,794,837

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	158,833	$8,735,815	2.3%	Defined
Held directly	6,636,004	$364,980,220	97.7%	Sole

WELLINGTON MANAGEMENT GROUP LLP

$369,989,675

4.83%

6,727,085

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wellington Management Company LLP	5,829,338	$320,613,590	86.7%	Defined
Wellington Managment Canada LLC	20,301	$1,116,555	0.3%	Defined
Wellington Management Singapore Pte. Ltd.	82,598	$4,542,890	1.2%	Defined
Wellington Management International Ltd	221,158	$12,163,690	3.3%	Defined
Wellington Trust Company, NA joint	555,960	$30,577,800	8.3%	Defined
Wellington Management Australia Pty Ltd	15,627	$859,485	0.2%	Defined
Wellington Management Europe GmbH	2,103	$115,665	0.0%	Defined

Orbis Allan Gray Ltd

$326,344,315

4.26%

5,933,533

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ORBIS INVESTMENT MANAGEMENT LTD	5,926,433	$325,953,815	99.9%	Defined
Held directly	7,100	$390,500	0.1%	Defined

VANGUARD CAPITAL MANAGEMENT LLC

$262,233,400

3.42%

4,767,880

STATE STREET CORP

$260,077,455

3.40%

4,728,681

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	4,728,681	$260,077,455	100.0%	Defined

Redmile Group, LLC

$237,110,060

3.10%

4,311,092

NORGES BANK

$152,345,050

1.99%

2,769,910

GEODE CAPITAL MANAGEMENT, LLC

$151,458,285

1.98%

2,753,787

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	384,976	$21,173,680	14.0%	Defined
Held directly	2,368,811	$130,284,605	86.0%	Defined

MASSACHUSETTS FINANCIAL SERVICES CO /MA/

$143,404,140

1.87%

2,607,348

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MFS Institutional Advisors, Inc.	260,301	$14,316,555	10.0%	Defined
MFS Investment Management CO (LUX) S.A.R.L.	16,130	$887,150	0.6%	Defined
MFS Investment Management K.K.	15,368	$845,240	0.6%	Defined
MFS Heritage Trust CO	416,566	$22,911,130	16.0%	Defined
MFS INVESTMENT MANAGEMENT CANADA LTD	4,366	$240,130	0.2%	Defined
Held directly	1,894,617	$104,203,935	72.7%	Sole

Bellevue Group AG

$134,725,085

1.76%

2,449,547

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BB Biotech AG joint	2,445,032	$134,476,760	99.8%	Defined
Bellevue Asset Management AG	4,515	$248,325	0.2%	Defined

JENNISON ASSOCIATES LLC

$129,722,285

1.69%

2,358,587

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Prudential Financial Inc	1,861,477	$102,381,235	78.9%	Defined
Prudential Trust Company	497,110	$27,341,050	21.1%	Other

Jupiter Topco LLC

$123,315,830

1.61%

2,242,106

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	40,395	$2,221,725	1.8%	Defined
JANUS HENDERSON INVESTORS US LLC	2,201,711	$121,094,105	98.2%	Defined

Hood River Capital Management LLC

$122,363,340

1.60%

2,224,788

Driehaus Capital Management LLC

$121,308,990

1.58%

2,205,618

EVENTIDE ASSET MANAGEMENT, LLC

$116,127,550

1.52%

2,111,410

Polar Capital Holdings Plc

$103,321,900

1.35%

1,878,580

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Polar Capital LLP	1,878,580	$103,321,900	100.0%	Defined

Holocene Advisors, LP

$96,578,790

1.26%

1,755,978

ADAGE CAPITAL PARTNERS GP, L.L.C.

$95,227,110

1.24%

1,731,402

MORGAN STANLEY

$84,632,680

1.11%

1,538,776

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	382	$21,010	0.0%	Defined
MORGAN STANLEY & CO. LLC	42,237	$2,323,035	2.7%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	1,404,585	$77,252,175	91.3%	Defined
MORGAN STANLEY SMITH BARNEY LLC	4,480	$246,400	0.3%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	61,928	$3,406,040	4.0%	Defined
EATON VANCE MANAGEMENT	25,164	$1,384,020	1.6%	Defined

Woodline Partners LP

$82,525,850

1.08%

1,500,470

GOLDMAN SACHS GROUP INC

$77,631,235

1.01%

1,411,477

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	1,197,209	$65,846,495	84.8%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	208,669	$11,476,795	14.8%	Defined
Goldman Sachs Trust Company, N.A.	5,599	$307,945	0.4%	Defined

Capital World Investors

$63,822,495

0.83%

1,160,409

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
CAPITAL GROUP COMPANIES INC joint	1,160,409	$63,822,495	100.0%	Defined

Clearbridge Investments, LLC

$58,287,680

0.76%

1,059,776

VANGUARD PORTFOLIO MANAGEMENT LLC

$56,504,635

0.74%

1,027,357

BNP PARIBAS ASSET MANAGEMENT Holding S.A.

$53,891,475

0.70%

979,845

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNP PARIBAS ASSET MANAGEMENT Europe	1,115	$61,325	0.1%	Defined
BNP Paribas Asset Management USA, Inc.	851,963	$46,857,965	86.9%	Defined
AXA INVESTMENT MANAGERS UK LIMITED	126,767	$6,972,185	12.9%	Defined

AMERICAN CENTURY COMPANIES INC

$53,635,835

0.70%

975,197

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AMERICAN CENTURY INVESTMENT MANAGEMENT INC	975,197	$53,635,835	100.0%	Defined

NORTHERN TRUST CORP

$53,601,680

0.70%

974,576

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	906,162	$49,838,910	93.0%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	51,061	$2,808,355	5.2%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	17,353	$954,415	1.8%	Other

BANK OF AMERICA CORP /DE/

$53,474,135

0.70%

972,257

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	761,408	$41,877,440	78.3%	Defined
BOFA SECURITIES, INC.	87,934	$4,836,370	9.0%	Defined
MANAGED ACCOUNT ADVISORS LLC joint	122,915	$6,760,325	12.6%	Defined

Pictet Asset Management Holding SA

$50,109,125

0.65%

911,075

VOYA INVESTMENT MANAGEMENT LLC

$48,983,385

0.64%

890,607

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	890,607	$48,983,385	100.0%	Defined

FRANKLIN RESOURCES INC

$48,329,930

0.63%

878,726

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FIDUCIARY TRUST INTERNATIONAL LLC	256	$14,080	0.0%	Defined
FRANKLIN ADVISERS INC	33,230	$1,827,650	3.8%	Defined
PUTNAM INVESTMENT MANAGEMENT LLC	553,659	$30,451,245	63.0%	Defined
PUTNAM ADVISORY CO LLC	291,581	$16,036,955	33.2%	Defined

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$47,535,730

0.62%

864,286

Integral Health Asset Management, LLC

$41,250,000

0.54%

750,000

FEDERATED HERMES, INC.

$40,398,215

0.53%

734,513

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FEDERATED GLOBAL INVESTMENT MANAGEMENT CORP	734,513	$40,398,215	100.0%	Defined

VANGUARD FIDUCIARY TRUST CO

$40,154,565

0.52%

730,083

BNP PARIBAS ARBITRAGE, SA

$39,463,160

0.52%

717,512

Rock Springs Capital Management LP

$39,349,860

0.51%

715,452

Casdin Capital, LLC

$36,575,000

0.48%

665,000

Nuveen, LLC

$35,846,195

0.47%

651,749

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	19,472	$1,070,960	3.0%	Defined
TIAA CREF INVESTMENT MANAGEMENT LLC joint	88,690	$4,877,950	13.6%	Defined
TEACHERS ADVISORS, LLC joint	543,587	$29,897,285	83.4%	Defined

LOOMIS SAYLES & CO L P

$32,783,025

0.43%

596,055

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
LOOMIS SAYLES TRUST COMPANY, LLC	100,389	$5,521,395	16.8%	Defined
Held directly	495,666	$27,261,630	83.2%	Defined

Ensign Peak Advisors, Inc

$31,847,860

0.42%

579,052

Eagle Health Investments LP

$30,539,960

0.40%

555,272

UBS Group AG

$28,114,350

0.37%

511,170

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	26,541	$1,459,755	5.2%	Defined
UBS AG	464,846	$25,566,530	90.9%	Defined
UBS Switzerland AG	19,731	$1,085,205	3.9%	Defined
UBS Securities LLC	52	$2,860	0.0%	Defined

Qube Research & Technologies Ltd

$25,505,150

0.33%

463,730

Fisher Asset Management, LLC

$24,169,585

0.32%

439,447

MARSHALL WACE, LLP

$23,677,500

0.31%

430,500

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marshall Wace North America L.P. joint	421,259	$23,169,245	97.9%	Other
MARSHALL WACE ASIA Ltd	9,241	$508,255	2.1%	Other

Bank of New York Mellon Corp

$20,086,990

0.26%

365,218

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNY Mellon Trust of Delaware joint	270	$14,850	0.1%	Defined
Mellon Investments Corporation joint	149,741	$8,235,755	41.0%	Defined
The Bank of New York Mellon	200,020	$11,001,100	54.8%	Defined
BNY Mellon, NA joint	13,262	$729,410	3.6%	Defined
BNY Mellon Advisors, Inc.	1,923	$105,765	0.5%	Defined
Held directly	2	$110	0.0%	Defined

CITADEL ADVISORS LLC

$18,251,805

0.24%

331,851

DEUTSCHE BANK AG\

$16,679,740

0.22%

303,268

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
DWS Investment GmbH	139,925	$7,695,875	46.1%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC	51,181	$2,814,955	16.9%	Defined
DWS Investments UK Ltd	112,162	$6,168,910	37.0%	Defined

Squarepoint Ops LLC

$16,059,780

0.21%

291,996

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Squarepoint Capital LLP	291,996	$16,059,780	100.0%	Defined

NAN FUNG TRINITY (HK) LTD

$15,702,225

0.21%

285,495

ARMISTICE CAPITAL, LLC

$15,290,000

0.20%

278,000

Trexquant Investment LP

$15,164,160

0.20%

275,712

First Turn Management, LLC

$14,917,485

0.19%

271,227

AQR CAPITAL MANAGEMENT LLC

$13,613,655

0.18%

247,521

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RF&L WEALTH MANAGEMENT, LLC	4	$220	0.0%	Other
Prism Planning Partners LLC	92	$5,060	0.0%	Other
Arax Advisory Partners	1,204	$66,220	0.5%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	406	$22,330	0.2%	Other
Stokes Family Office, LLC	303	$16,665	0.1%	Other
Held directly	245,512	$13,503,160	99.2%	Defined

D. E. Shaw & Co., Inc.

$13,476,485

0.18%

245,027

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	153,988	$8,469,340	62.8%	Defined
D. E. Shaw Investment Management, L.L.C. joint	91,039	$5,007,145	37.2%	Defined

RAYMOND JAMES FINANCIAL INC

$13,236,575

0.17%

240,665

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	84,051	$4,622,805	34.9%	Defined
Raymond James Financial Services Advisors, Inc.	44,782	$2,463,010	18.6%	Defined
CHARTWELL INVESTMENT PARTNERS, LLC	111,832	$6,150,760	46.5%	Defined

STEMPOINT CAPITAL LP

$13,153,030

0.17%

239,146

Candriam S.C.A.

$11,888,030

0.16%

216,146

Universal- Beteiligungs- und Servicegesellschaft mbH

$11,549,505

0.15%

209,991

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Universal-Investment-Gesellschaft mbH joint	209,991	$11,549,505	100.0%	Defined

Senator Investment Group LP

$11,275,000

0.15%

205,000

Logos Global Management LP

$11,000,000

0.14%

200,000

Susquehanna Portfolio Strategies, LLC

$10,947,860

0.14%

199,052

SEGALL BRYANT & HAMILL, LLC

$10,372,340

0.14%

188,588

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NEW YORK LIFE INVESTMENT MANAGEMENT LLC	188,588	$10,372,340	100.0%	Sole

JPMORGAN CHASE & CO

$9,773,060

0.13%

177,692

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
J.P. Morgan Trust Co of Delaware	351	$19,305	0.2%	Other
JPMorgan Chase Bank, N.A.	52,258	$2,874,190	29.4%	Defined
J.P. Morgan Investment Management Inc.	116,156	$6,388,580	65.4%	Defined
J.P. Morgan Securities LLC	422	$23,210	0.2%	Defined
J.P. Morgan Securities plc	8,505	$467,775	4.8%	Defined

SEI INVESTMENTS CO

$9,549,265

0.12%

173,623

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI INVESTMENTS MANAGEMENT CORP	3,774	$207,570	2.2%	Sole
SEI TRUST CO	169,849	$9,341,695	97.8%	Sole

Granahan Investment Management, LLC

$9,006,745

0.12%

163,759

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$8,888,000

0.12%

161,600

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	161,600	$8,888,000	100.0%	Other

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$8,624,000

0.11%

156,800

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	156,800	$8,624,000	100.0%	Other

Swiss National Bank

$8,239,000

0.11%

149,800

RHUMBLINE ADVISERS

$8,000,300

0.10%

145,460

ALLIANCEBERNSTEIN L.P.

$7,690,100

0.10%

139,820

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	139,820	$7,690,100	100.0%	Defined

PEREGRINE CAPITAL MANAGEMENT LLC

$7,674,370

0.10%

139,534

AMERIPRISE FINANCIAL INC

$7,341,510

0.10%

133,482

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	124,316	$6,837,380	93.1%	Defined
Columbia Threadneedle Management Ltd	9,166	$504,130	6.9%	Defined

GordonMD Global Investments LP

$7,287,500

0.10%

132,500

Russell Investments Group, Ltd.

$7,178,270

0.09%

130,514

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC	87,554	$4,815,470	67.1%	Defined
Russell Investments Capital, LLC	21,702	$1,193,610	16.6%	Defined
Russell Investments France SAS	7,587	$417,285	5.8%	Defined
Russell Investments Trust Company	13,671	$751,905	10.5%	Defined

MGO ONE SEVEN LLC

$6,879,235

0.09%

125,077

iA Global Asset Management Inc.

$6,754,660

0.09%

122,812

MANUFACTURERS LIFE INSURANCE COMPANY, THE

$6,722,265

0.09%

122,223

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MANULIFE FINANCIAL CORP joint	122,223	$6,722,265	100.0%	Defined

TD Asset Management Inc

$6,620,020

0.09%

120,364

WELLS FARGO & COMPANY/MN

$6,325,605

0.08%

115,011

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	101,523	$5,583,765	88.3%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	5,154	$283,470	4.5%	Defined
WELLS FARGO CLEARING SERVICES, LLC	8,334	$458,370	7.2%	Defined

CITIGROUP INC

$6,216,705

0.08%

113,031

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	113,031	$6,216,705	100.0%	Defined

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$6,115,230

0.08%

111,186

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	1,454	$79,970	1.3%	Defined
UBS Asset Management Switzerland AG	92,625	$5,094,375	83.3%	Sole
UBS Asset Management (UK) Ltd.	7,733	$425,315	7.0%	Defined
Held directly	9,374	$515,570	8.4%	Defined

EFG International AG

$6,084,870

0.08%

110,634

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
EFG Asset Management (North America) Corp.	110,587	$6,082,285	100.0%	Defined
EFG Private Wealth Management (Americas) Corp.	47	$2,585	0.0%	Defined

Man Group plc

$5,739,030

0.07%

104,346

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AHL Partners LLP	104,346	$5,739,030	100.0%	Defined

CITADEL ADVISORS LLC

$5,566,000

0.07%

101,200

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	101,200	$5,566,000	100.0%	Defined

Ghisallo Capital Management LLC

$5,500,000

0.07%

100,000

Artia Global Partners LP

$5,295,675

0.07%

96,285

SPHERA FUNDS MANAGEMENT LTD.

$5,184,575

0.07%

94,265

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Sphera Global Healthcare Management LP joint	94,265	$5,184,575	100.0%	Defined

Invesco Ltd.

$5,178,580

0.07%

94,156

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	16,833	$925,815	17.9%	Defined
Invesco Trust Co joint	44,214	$2,431,770	47.0%	Defined
Invesco Capital Management LLC	33,109	$1,820,995	35.2%	Defined

RENAISSANCE TECHNOLOGIES LLC

$5,032,500

0.07%

91,500

FIDEURAM - INTESA SANPAOLO PRIVATE BANKING S.P.A.

$5,005,990

0.07%

91,018

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Intesa Sanpaolo S.p.A.	91,018	$5,005,990	100.0%	Defined

Nicholas Investment Partners, LP

$4,968,370

0.06%

90,334

BEACONLIGHT CAPITAL, LLC

$4,834,115

0.06%

87,893

CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM

$4,710,420

0.06%

85,644

VANGUARD ASSET MANAGEMENT, Ltd

$4,385,535

0.06%

79,737

Key facts CIK 1727196 CUSIP 80706P103 13F (30d) 252 filings 251 filers Visit website Investor relations