PTCT · Ptc Therapeutics, Inc.

$69.30 -2.10 (-2.94%) At close · Sep 4

S&P 600 member Russell 2000 member Rebalance outlook locked

Market Cap

$5.96B

83.46M

Institutional ownership from SEC Form 13F filings. Investment managers with more than $100 million in U.S. equities must disclose their positions every quarter, within 45 days of quarter-end, so the holdings shown can lag the market by up to a quarter.

Use institutional holdings through the 13F API or MCP →

Quarter:

Holders: 385

Total Value: $8,272,166,374

Total Shares: 101,449,094

Showing 1–100 of 394 holdings by value · page 1 of 4

Institutional Ownership Trend

26 quarters

Holder Concentration

Across 385 13F filers · Δ vs Mar 31, 2026

Top Holder

13.8%

+0.3 pts

Top 5

37.9%

−0.5 pts

Top 10

51.9%

−2.3 pts

HHI

427

Broadly held−9

Quarterly Activity

vs Mar 31, 2026

New

Increased

144

Decreased

113

Sold Out

Unchanged

Holder count: +30

Institutional value: +25.9%

Shares held: +5.2%

Top Buyers

vs Mar 31, 2026

Institution	Added	Now Holds
Mega BlackRock, Inc.	+1,015,837	13,964,554
Mid Blue Sparrow, LLC /DE	+860,100	1,050,700
Mega PRICE T ROWE ASSOCIATES INC /MD/	+639,368	1,114,516
Mega ARROWSTREET CAPITAL, LIMITED PARTNERSHIP	+542,750	654,382
Mega AMERICAN CENTURY COMPANIES INC	+495,992	500,995

Top Sellers

vs Mar 31, 2026

Institution	Sold	Now Holds
Mega UBS Group AG	−1,102,342	262,426
Large TORONTO DOMINION BANK	−860,117	2,165,846
Mid ARMISTICE CAPITAL, LLC	−733,054	1,731,395
Mega BNP PARIBAS ARBITRAGE, SA	−541,756	378,720
Large Balyasny Asset Management L.P.	−439,959	20,605

Top New Positions

vs Mar 31, 2026

Institution	Acquired	Value (USD)
Mega Jupiter Topco LLC	7,236,402	$590,273,310
Large WESTFIELD CAPITAL MANAGEMENT CO LP	1,956,156	$159,563,644
Large WILLIAM BLAIR INVESTMENT MANAGEMENT, LLC	1,722,688	$140,519,660
Large FULLER & THALER ASSET MANAGEMENT, INC.	1,156,819	$94,361,725
Mid Longaeva Partners L.P.	111,803	$9,119,770

Top Sold-Out Positions

vs Mar 31, 2026

Institution	Previously Held	Last Value (USD)
Mega JANUS HENDERSON GROUP PLC	7,193,433	$490,088,588
Large Woodline Partners LP	744,988	$50,756,032
Mega DZ BANK AG Deutsche Zentral Genossenschafts Bank, Frankfurt am Main	213,363	$14,536,421
Large BI Asset Management Fondsmaeglerselskab A/S	81,878	$5,578,348
Large PICTON MAHONEY ASSET MANAGEMENT	53,328	$3,633,236

Filter: New 76 Increased 144 Reduced 113 Sold out 46 Unchanged 52

Showing 100 of 394

All Common Calls Puts

Compact Export CSV

Holder

Value (USD)

% of 13F

BlackRock, Inc.

$1,139,088,662

13.77%

13,964,554

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BLACKROCK FINANCIAL MANAGEMENT INC/DE	166,600	$13,589,562	1.2%	Sole
BlackRock Investment Management (Australia) Ltd	868	$70,802	0.0%	Sole
BlackRock Asset Management Canada Ltd	8,343	$680,538	0.1%	Sole
BlackRock Investment Management, LLC	284,322	$23,192,145	2.0%	Sole
BLACKROCK ADVISORS LLC	470,838	$38,406,255	3.4%	Sole
BlackRock Fund Advisors	9,614,679	$784,269,366	68.9%	Sole
BlackRock Institutional Trust Company, N.A.	2,744,397	$223,860,463	19.7%	Sole
BlackRock Japan Co. Ltd	7,322	$597,255	0.1%	Sole
BlackRock Fund Managers Ltd.	13,653	$1,113,675	0.1%	Sole
BlackRock Investment Management (UK) Ltd.	64,685	$5,276,355	0.5%	Sole
BlackRock (Netherlands) B.V.	2,060	$168,034	0.0%	Sole
BlackRock Asset Management Ireland Ltd	452,792	$36,934,243	3.2%	Sole
BlackRock (Luxembourg) S.A.	9,613	$784,132	0.1%	Sole
BlackRock Life Ltd	343	$27,978	0.0%	Sole
BlackRock Asset Management Schweiz AG	12,324	$1,005,268	0.1%	Sole
Aperio Group, LLC	111,649	$9,107,208	0.8%	Sole
SpiderRock Advisors, LLC	66	$5,383	0.0%	Sole

RTW INVESTMENTS, LP

$631,875,071

7.64%

7,746,415

Jupiter Topco LLC

$590,273,310

7.14%

7,236,402

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
JANUS HENDERSON INVESTORS UK Ltd	149,897	$12,227,098	2.1%	Defined
JANUS HENDERSON INVESTORS US LLC	7,076,322	$577,215,585	97.8%	Defined
Held directly	10,183	$830,627	0.1%	Defined

VANGUARD PORTFOLIO MANAGEMENT LLC

$401,986,014

4.86%

4,928,111

STATE STREET CORP

$375,136,514

4.53%

4,598,952

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
STATE STREET GLOBAL ADVISORS EUROPE LIMITED joint	4,598,952	$375,136,514	100.0%	Defined

WELLINGTON MANAGEMENT GROUP LLP

$336,743,224

4.07%

4,128,273

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Wellington Management Company LLP	3,585,091	$292,435,872	86.8%	Defined
Wellington Managment Canada LLC	27,985	$2,282,736	0.7%	Defined
Wellington Management Singapore Pte. Ltd.	50,969	$4,157,541	1.2%	Defined
Wellington Management International Ltd	148,563	$12,118,283	3.6%	Defined
Wellington Trust Company, NA joint	303,314	$24,741,322	7.3%	Defined
Wellington Management Australia Pty Ltd	10,466	$853,711	0.3%	Defined
Wellington Management Europe GmbH	1,885	$153,759	0.0%	Defined

VANGUARD CAPITAL MANAGEMENT LLC

$305,000,589

3.69%

3,739,127

GEODE CAPITAL MANAGEMENT, LLC

$180,055,659

2.18%

2,207,376

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GEODE CAPITAL MANAGEMENT TRUST COMPANY, LLC	305,959	$24,957,075	13.9%	Defined
Held directly	1,901,417	$155,098,584	86.1%	Defined

TORONTO DOMINION BANK

$176,668,057

2.14%

2,165,846

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
TD SECURITIES INC	1	$81	0.0%	Defined
Held directly	2,165,845	$176,667,976	100.0%	Sole

WESTFIELD CAPITAL MANAGEMENT CO LP

$159,563,644

1.93%

1,956,156

Hood River Capital Management LLC

$141,818,662

1.71%

1,738,613

WILLIAM BLAIR INVESTMENT MANAGEMENT, LLC

$140,519,660

1.70%

1,722,688

D. E. Shaw & Co., Inc.

$126,345,322

1.53%

1,548,919

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
D. E. SHAW & CO, L.P.	949,845	$77,478,856	61.3%	Defined
D. E. Shaw Investment Management, L.L.C. joint	599,074	$48,866,466	38.7%	Defined

TANG CAPITAL MANAGEMENT LLC

$122,355,000

1.48%

1,500,000

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
TANG KEVIN	1,500,000	$122,355,000	100.0%	Other

BANK OF AMERICA CORP /DE/

$118,462,233

1.43%

1,452,277

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BANK OF AMERICA NA	690,828	$56,350,839	47.6%	Defined
MERRILL LYNCH, PIERCE, FENNER & SMITH INC.	136,574	$11,140,341	9.4%	Defined
BOFA SECURITIES, INC.	199,237	$16,251,762	13.7%	Defined
MANAGED ACCOUNT ADVISORS LLC joint	425,638	$34,719,291	29.3%	Defined

Driehaus Capital Management LLC

$117,469,527

1.42%

1,440,107

FMR LLC

$114,056,147

1.38%

1,398,261

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Fidelity Management & Research Co LLC	698,259	$56,956,986	49.9%	Defined
STRATEGIC ADVISERS LLC	8,971	$731,764	0.6%	Defined
Fidelity Institutional Asset Management Trust Co	275,292	$22,455,568	19.7%	Defined
FIAM LLC	388,796	$31,714,089	27.8%	Defined
Fidelity Diversifying Solutions LLC	26,943	$2,197,740	1.9%	Defined

Jefferies Financial Group Inc.

$110,693,915

1.34%

1,357,042

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jefferies Financial Services, Inc.	1,357,042	$110,693,915	100.0%	Defined

FULLER & THALER ASSET MANAGEMENT, INC.

$94,361,725

1.14%

1,156,819

Pictet Asset Management Holding SA

$93,629,635

1.13%

1,147,844

PRICE T ROWE ASSOCIATES INC /MD/

$90,911,069

1.10%

1,114,516

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
T. Rowe Price International Ltd	110,208	$8,989,666	9.9%	Defined
Held directly	1,004,308	$81,921,403	90.1%	Sole

MORGAN STANLEY

$90,026,765

1.09%

1,103,675

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MORGAN STANLEY & CO INTERNATIONAL PLC	413	$33,688	0.0%	Defined
MORGAN STANLEY & CO. LLC	262,490	$21,411,309	23.8%	Defined
MORGAN STANLEY CAPITAL SERVICES, LLC	543,646	$44,345,204	49.3%	Defined
MORGAN STANLEY INVESTMENT MANAGEMENT INC	2,135	$174,151	0.2%	Defined
MORGAN STANLEY SMITH BARNEY LLC	9,390	$765,941	0.9%	Defined
PARAMETRIC PORTFOLIO ASSOCIATES LLC	270,016	$22,025,204	24.5%	Defined
Calvert Research & Management	15,585	$1,271,268	1.4%	Defined

ARMISTICE CAPITAL, LLC

$87,393,690

1.06%

1,071,395

Blue Sparrow, LLC /DE

$85,705,599

1.04%

1,050,700

FRANKLIN RESOURCES INC

$83,799,061

1.01%

1,027,327

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FIDUCIARY TRUST CO INTERNATIONAL	517	$42,171	0.1%	Defined
FIDUCIARY TRUST INTERNATIONAL LLC	2,334	$190,384	0.2%	Defined
FRANKLIN ADVISERS INC	1,024,289	$83,551,253	99.7%	Defined
Franklin Templeton International Services S.a r.l	187	$15,253	0.0%	Defined

Invesco Ltd.

$76,804,760

0.93%

941,581

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Invesco Advisers, Inc. joint	32,583	$2,657,794	3.5%	Defined
Invesco Trust Co joint	33,011	$2,692,707	3.5%	Defined
Invesco Investment Advisers LLC	5,825	$475,145	0.6%	Defined
Invesco Capital Management LLC	870,162	$70,979,114	92.4%	Defined

Palo Alto Investors LP

$74,950,268

0.91%

918,846

NORTHERN TRUST CORP

$71,783,963

0.87%

880,029

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NORTHERN TRUST CO joint	790,678	$64,495,603	89.8%	Defined
NORTHERN TRUST GLOBAL INVESTMENTS LTD	48,761	$3,977,434	5.5%	Defined
MASON STREET ADVISORS, LLC joint	27,723	$2,261,365	3.2%	Other
DWS INVESTMENT MANAGEMENT AMERICAS, INC joint	12,867	$1,049,561	1.5%	Other

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

$64,449,435

0.78%

790,112

CITADEL ADVISORS LLC

$60,999,840

0.74%

747,822

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citadel Securities GP LLC	498,490	$40,661,829	66.7%	Defined
Held directly	249,332	$20,338,011	33.3%	Defined

RENAISSANCE TECHNOLOGIES LLC

$59,015,895

0.71%

723,500

Boston Partners

$58,614,080

0.71%

718,574

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Boston Partners Trust Co	18,703	$1,525,603	2.6%	Defined
Held directly	699,871	$57,088,477	97.4%	Defined

JUPITER ASSET MANAGEMENT LTD

$57,706,695

0.70%

707,450

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Jupiter Investment Management Limited	109,442	$8,927,183	15.5%	Defined
Held directly	598,008	$48,779,512	84.5%	Sole

MARSHALL WACE, LLP

$57,489,474

0.69%

704,787

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Marshall Wace North America L.P. joint	704,787	$57,489,474	100.0%	Other

JPMORGAN CHASE & CO

$55,797,302

0.67%

684,042

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
J.P. Morgan Trust Co of Delaware	1,100	$89,726	0.2%	Other
JPMorgan Chase Bank, N.A.	40,000	$3,262,800	5.8%	Defined
J.P. Morgan Investment Management Inc.	113,492	$9,257,542	16.6%	Defined
J.P. Morgan Securities LLC	126,983	$10,358,002	18.6%	Defined
JPMorgan Asset Management (UK) Ltd	402,460	$32,828,662	58.8%	Defined
55I, LLC	7	$570	0.0%	Other

ARMISTICE CAPITAL, LLC

$53,836,200

0.65%

660,000

ARROWSTREET CAPITAL, LIMITED PARTNERSHIP

$53,377,939

0.65%

654,382

DIMENSIONAL FUND ADVISORS LP

$52,017,514

0.63%

637,704

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Dimensional Ireland Ltd	6,505	$530,612	1.0%	Defined
Held directly	631,199	$51,486,902	99.0%	Sole

GOLDMAN SACHS GROUP INC

$50,430,406

0.61%

618,247

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
GOLDMAN SACHS & CO. LLC	418,702	$34,153,522	67.7%	Defined
GOLDMAN SACHS ASSET MANAGEMENT, L.P.	195,696	$15,962,922	31.7%	Defined
Goldman Sachs Trust Company, N.A.	3,849	$313,962	0.6%	Defined

SG Americas Securities, LLC

$49,397,242

0.60%

605,581

LOOMIS SAYLES & CO L P

$48,156,806

0.58%

590,374

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
LOOMIS SAYLES TRUST COMPANY, LLC	156,539	$12,768,886	26.5%	Defined
Held directly	433,835	$35,387,920	73.5%	Defined

LORD, ABBETT & CO. LLC

$48,132,336

0.58%

590,074

VANGUARD FIDUCIARY TRUST CO

$47,953,942

0.58%

587,887

Nuveen, LLC

$47,622,931

0.58%

583,829

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Nuveen Asset Management, LLC joint	25,083	$2,046,020	4.3%	Defined
TIAA CREF INVESTMENT MANAGEMENT LLC joint	96,109	$7,839,611	16.5%	Defined
TEACHERS ADVISORS, LLC joint	462,637	$37,737,300	79.2%	Defined

Bank of New York Mellon Corp

$45,145,891

0.55%

553,462

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Mellon Investments Corporation joint	322,617	$26,315,867	58.3%	Defined
BNY Mellon Investment Adviser, Inc.	67,360	$5,494,555	12.2%	Defined
The Bank of New York Mellon	155,430	$12,678,425	28.1%	Defined
BNY Mellon, NA joint	3,200	$261,023	0.6%	Defined
BNY Mellon Advisors, Inc. joint	4,286	$349,608	0.8%	Defined
Newton Investment Management North America, LLC	567	$46,250	0.1%	Defined
Held directly	2	$163	0.0%	Defined

PRINCIPAL FINANCIAL GROUP INC

$44,181,166

0.53%

541,635

VOYA INVESTMENT MANAGEMENT LLC

$43,992,249

0.53%

539,319

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Voya Investment Management Co. LLC joint	539,319	$43,992,249	100.0%	Defined

Ensign Peak Advisors, Inc

$41,442,127

0.50%

508,056

AMERICAN CENTURY COMPANIES INC

$40,866,162

0.49%

500,995

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
AMERICAN CENTURY INVESTMENT MANAGEMENT INC	500,995	$40,866,162	100.0%	Defined

FEDERATED HERMES, INC.

$39,432,894

0.48%

483,424

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
FEDERATED EQUITY MANAGEMENT CO OF PENNSYLVANIA	12,177	$993,277	2.5%	Defined
FEDERATED GLOBAL INVESTMENT MANAGEMENT CORP	1,474	$120,234	0.3%	Defined
FEDERATED MDTA LLC	469,773	$38,319,383	97.2%	Defined

MANUFACTURERS LIFE INSURANCE COMPANY, THE

$36,050,432

0.44%

441,957

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
MANULIFE FINANCIAL CORP joint	441,957	$36,050,432	100.0%	Defined

Parkman Healthcare Partners LLC

$35,539,314

0.43%

435,691

Defilade Capital Management, L.P.

$35,376,827

0.43%

433,699

Russell Investments Group, Ltd.

$34,098,295

0.41%

418,025

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Russell Investment Management, LLC joint	183,461	$14,964,913	43.9%	Defined
Russell Investments Canada Limited joint	5,797	$472,860	1.4%	Defined
Russell Investments Capital, LLC	73,795	$6,019,458	17.7%	Defined
Russell Investments France SAS	24,210	$1,974,809	5.8%	Defined
Russell Investments Limited	1,878	$153,188	0.4%	Defined
Russell Investments Trust Company joint	128,884	$10,513,067	30.8%	Defined

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$32,274,149

0.39%

395,662

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	395,662	$32,274,149	100.0%	Other

BNP PARIBAS ARBITRAGE, SA

$30,892,190

0.37%

378,720

Legal & General Group Plc

$28,772,512

0.35%

352,734

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Legal & General Investment Management America Inc	127,044	$10,362,979	36.0%	Defined
Legal & General Investment Management Ltd joint	225,690	$18,409,533	64.0%	Defined

AQR CAPITAL MANAGEMENT LLC

$27,101,712

0.33%

332,251

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Prism Planning Partners LLC	67	$5,465	0.0%	Other
Compound Planning, Inc.	3	$244	0.0%	Other
Arax Advisory Partners	6,590	$537,546	2.0%	Other
HALBERT HARGROVE GLOBAL ADVISORS, LLC	1,534	$125,128	0.5%	Other
Stokes Family Office, LLC	918	$74,881	0.3%	Other
Held directly	323,139	$26,358,448	97.3%	Defined

Qube Research & Technologies Ltd

$25,398,777

0.31%

311,374

BNP PARIBAS ASSET MANAGEMENT Holding S.A.

$24,771,584

0.30%

303,685

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BNP PARIBAS ASSET MANAGEMENT Europe	1,016	$82,875	0.3%	Defined
BNP Paribas Asset Management USA, Inc.	301,361	$24,582,016	99.2%	Defined
AXA INVESTMENT MANAGERS UK LIMITED	1,308	$106,693	0.4%	Defined

Rockefeller Capital Management L.P.

$24,666,195

0.30%

302,393

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
ROCKEFELLER & CO. LLC	719	$58,648	0.2%	Defined
ROCKEFELLER TRUST COMPANY, N.A.	1,646	$134,264	0.5%	Defined
ROCKEFELLER FINANCIAL LLC	300,028	$24,473,283	99.2%	Defined

DEUTSCHE BANK AG\

$24,218,049

0.29%

296,899

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Deutsche Bank Trust Co Americas	1,758	$143,400	0.6%	Defined
DWS Investment GmbH	182,842	$14,914,421	61.6%	Defined
DWS INVESTMENT MANAGEMENT AMERICAS, INC	25,585	$2,086,968	8.6%	Defined
DWS Investments UK Ltd	84,888	$6,924,314	28.6%	Defined
DWS International GmbH	1,826	$148,946	0.6%	Defined

Public Sector Pension Investment Board

$23,249,326

0.28%

285,023

SEGALL BRYANT & HAMILL, LLC

$22,165,423

0.27%

271,735

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
NEW YORK LIFE INVESTMENT MANAGEMENT LLC	271,735	$22,165,423	100.0%	Sole

UBS Group AG

$21,406,088

0.26%

262,426

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Financial Services Inc	11,567	$943,520	4.4%	Defined
UBS AG	231,515	$18,884,678	88.2%	Defined
UBS Switzerland AG	19,344	$1,577,890	7.4%	Defined

Allianz Asset Management GmbH

$18,850,990

0.23%

231,102

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Allianz Global Investors GmbH	219,702	$17,921,092	95.1%	Defined
Allianz Global Invesstors Taiwan Ltd.	11,400	$929,898	4.9%	Defined

Boxer Capital Management, LLC

$16,314,000

0.20%

200,000

RHUMBLINE ADVISERS

$15,719,028

0.19%

192,706

PRUDENTIAL FINANCIAL INC

$15,157,337

0.18%

185,820

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
PGIM Quantitative Solutions LLC	168,520	$13,746,176	90.7%	Defined
Prudential Trust Co	17,300	$1,411,161	9.3%	Defined

AMERIPRISE FINANCIAL INC

$14,019,761

0.17%

171,874

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Columbia Management Investment Advisers, LLC	170,457	$13,904,177	99.2%	Defined
Ameriprise Financial Services, LLC	1,393	$113,627	0.8%	Defined
Ameriprise Bank, FSB	24	$1,957	0.0%	Defined

Rock Springs Capital Management LP

$13,238,647

0.16%

162,298

Hudson Bay Capital Management LP

$13,033,091

0.16%

159,778

Schonfeld Strategic Advisors LLC

$12,025,457

0.15%

147,425

Swiss National Bank

$11,539,300

0.14%

141,465

Universal- Beteiligungs- und Servicegesellschaft mbH

$11,063,583

0.13%

135,633

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Universal-Investment-Gesellschaft mbH joint	135,633	$11,063,583	100.0%	Defined

HSBC HOLDINGS PLC

$10,717,400

0.13%

131,389

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
HSBC GLOBAL ASSET MANAGEMENT (FRANCE)	92,657	$7,558,031	70.5%	Defined
HSBC BANK PLC	38,732	$3,159,369	29.5%	Defined

UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC

$10,278,713

0.12%

126,011

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
UBS Group AG	1,254	$102,288	1.0%	Defined
UBS Asset Management Switzerland AG	93,026	$7,588,130	73.8%	Sole
UBS Asset Management (UK) Ltd.	17,494	$1,426,984	13.9%	Defined
Held directly	14,237	$1,161,311	11.3%	Defined

SEI INVESTMENTS CO

$10,231,650

0.12%

125,434

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SEI INVESTMENTS MANAGEMENT CORP	385	$31,404	0.3%	Sole
SEI TRUST CO	125,049	$10,200,246	99.7%	Sole

CITIGROUP INC

$9,678,769

0.12%

118,656

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Citigroup Financial Products Inc. joint	118,656	$9,678,769	100.0%	Defined

FIRST TRUST ADVISORS LP

$9,246,285

0.11%

113,354

Longaeva Partners L.P.

$9,119,770

0.11%

111,803

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Balyasny Asset Management L.P.	111,803	$9,119,770	100.0%	Other

ALLIANCEBERNSTEIN L.P.

$8,641,281

0.10%

105,937

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Equitable Holdings, Inc.	105,937	$8,641,281	100.0%	Defined

SCHRODER INVESTMENT MANAGEMENT GROUP

$8,273,563

0.10%

101,429

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Schroder Investment Management Limited	100,716	$8,215,404	99.3%	Sole
Schroder Investment Management North America Inc.	713	$58,159	0.7%	Sole

SECTOR GAMMA AS

$8,157,000

0.10%

100,000

WCM INVESTMENT MANAGEMENT, LLC

$7,651,184

0.09%

93,799

PANAGORA ASSET MANAGEMENT INC

$7,256,140

0.09%

88,956

BESSEMER GROUP INC

$6,938,017

0.08%

85,056

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BESSEMER TRUST Co NATIONAL ASSOCIATION	84,963	$6,930,431	99.9%	Defined
Bessemer Trust CO of Delaware, N.A.	93	$7,586	0.1%	Other

Aquatic Capital Management LLC

$6,808,647

0.08%

83,470

WELLS FARGO & COMPANY/MN

$6,776,099

0.08%

83,071

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
WELLS FARGO BANK N A	77,464	$6,318,738	93.3%	Defined
WELLS FARGO ADVISORS FINANCIAL NETWORK, LLC	4,003	$326,524	4.8%	Defined
WELLS FARGO CLEARING SERVICES, LLC	1,575	$128,472	1.9%	Defined
WELLS FARGO INVESTMENT INSTITUTE, INC.	29	$2,365	0.0%	Defined

XTX Topco Ltd

$6,682,459

0.08%

81,923

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
XTX EXECUTION SERVICES LLC joint	81,923	$6,682,459	100.0%	Defined

CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM

$6,666,960

0.08%

81,733

Empowered Funds, LLC

$6,629,275

0.08%

81,271

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
Strive Asset Management, LLC	2,923	$238,429	3.6%	Other
The Burney Company	78,348	$6,390,846	96.4%	Other

MOORE CAPITAL MANAGEMENT, LP

$6,165,713

0.07%

75,588

PINNACLE ASSOCIATES LTD

$5,745,382

0.07%

70,435

RAYMOND JAMES FINANCIAL INC

$5,682,165

0.07%

69,660

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
RAYMOND JAMES & ASSOCIATES	31,915	$2,603,306	45.8%	Defined
Raymond James Financial Services Advisors, Inc.	650	$53,020	0.9%	Defined
ClariVest Asset Management LLC joint	37,095	$3,025,839	53.3%	Defined

VANGUARD ASSET MANAGEMENT, Ltd

$5,614,299

0.07%

68,828

CANADA LIFE ASSURANCE Co

$5,229,615

0.06%

64,112

Capricorn Fund Managers Ltd

$5,212,323

0.06%

63,900

SUSQUEHANNA INTERNATIONAL GROUP, LLP

$5,000,241

0.06%

61,300

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
SUSQUEHANNA SECURITIES, LLC	61,300	$5,000,241	100.0%	Other

BARCLAYS PLC

$4,761,239

0.06%

58,370

This filing is a combination report: the position is filed across the managers below.

Manager	Shares	Value (USD)	% of position	Discretion
BARCLAYS BANK PLC	51,120	$4,169,858	87.6%	Defined
BARCLAYS CAPITAL INC.	6,355	$518,377	10.9%	Defined
BARCLAYS CAPITAL SECURITIES LTD	48	$3,915	0.1%	Defined
Barclays Investment Solutions Ltd	847	$69,089	1.5%	Defined

Key facts CIK 1070081 CUSIP 69366J200 13F (30d) 271 filings 263 filers Visit website Investor relations